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''Eucalyptus'' () is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
of over seven hundred species of
flowering A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechanism ...
trees, shrubs or mallees in the myrtle family, Myrtaceae. Along with several other genera in the tribe
Eucalypteae Eucalypteae is a large tribe of flowering plants in the family Myrtaceae; members of this tribe are known as eucalypts. In Australia the genera ''Angophora'', ''Corymbia'', and ''Eucalyptus'' are commonly known as gum trees, for the sticky substa ...
, including '' Corymbia'', they are commonly known as
eucalypt Eucalypt is a descriptive name for woody plants with capsule fruiting bodies belonging to seven closely related genera (of the tribe Eucalypteae) found across Australasia: ''Eucalyptus'', ''Corymbia'', ''Angophora'', '' Stockwellia'', ''Allosyn ...
s. Plants in the genus ''Eucalyptus'' have bark that is either smooth, fibrous, hard or stringy, leaves with oil glands, and
sepal A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 The term ''sepalum'' was coine ...
s and petals that are fused to form a "cap" or operculum over the stamens. The fruit is a woody capsule commonly referred to as a "gumnut". Most species of ''Eucalyptus'' are native to Australia, and every state and territory has representative species. About three-quarters of Australian forests are eucalypt forests. Wildfire is a feature of the Australian landscape and many eucalypt species are adapted to fire, and resprout after fire or have seeds which survive fire. A few species are native to islands north of Australia and a smaller number are only found outside the continent. Eucalypts have been grown in plantations in many other countries because they are fast growing and have valuable timber, or can be used for pulpwood, for honey production or essential oils. In some countries, however, they have been removed because of the danger of forest fires due to their high inflammability.


Description


Size and habit

Eucalypts vary in size and habit from shrubs to tall trees. Trees usually have a single main stem or trunk but many eucalypts are mallees that are multistemmed from ground level and rarely taller than . There is no clear distinction between a mallee and a shrub but in eucalypts, a shrub is a mature plant less than tall and growing in an extreme environment. '' E. vernicosa'' in the
Tasmania ) , nickname = , image_map = Tasmania in Australia.svg , map_caption = Location of Tasmania in AustraliaCoordinates: , subdivision_type = Country , subdi ...
n highlands, '' E. yalatensis'' on the
Nullarbor The Nullarbor Plain ( ; Latin: feminine of , 'no', and , 'tree') is part of the area of flat, almost treeless, arid or semi-arid country of southern Australia, located on the Great Australian Bight coast with the Great Victoria Desert to its ...
and '' E. surgens'' growing on coastal cliffs in
Western Australia Western Australia (commonly abbreviated as WA) is a state of Australia occupying the western percent of the land area of Australia excluding external territories. It is bounded by the Indian Ocean to the north and west, the Southern Ocean to th ...
are examples of eucalypt shrubs. The terms " mallet" and " marlock" are only applied to
Western Australia Western Australia (commonly abbreviated as WA) is a state of Australia occupying the western percent of the land area of Australia excluding external territories. It is bounded by the Indian Ocean to the north and west, the Southern Ocean to th ...
n eucalypts. A mallet is a tree with a single thin trunk with a steeply branching habit but lacks both a
lignotuber A lignotuber is a woody swelling of the root crown possessed by some plants as a protection against destruction of the plant stem, such as by fire. Other woody plants may develop basal burls as a similar survival strategy, often as a response t ...
and epicormic buds. '' E. astringens'' is an example of a mallet. A marlock is a shrub or small tree with a single, short trunk, that lacks a lignotuber and has spreading, densely leafy branches that often reach almost to the ground. '' E. platypus'' is an example of a marlock. ''Eucalyptus'' trees, including mallets and marlocks, are single-stemmed and include ''
Eucalyptus regnans ''Eucalyptus'' () is a genus of over seven hundred species of flowering trees, shrubs or mallees in the myrtle family, Myrtaceae. Along with several other genera in the tribe Eucalypteae, including ''Corymbia'', they are commonly known as euc ...
'', the tallest known flowering plant on Earth. Tree sizes follow the convention of: * Small: to in height * Medium-sized: * Tall: * Very tall: over Tasmania logging 08 Mighty tree.jpg, ''
Eucalyptus regnans ''Eucalyptus'' () is a genus of over seven hundred species of flowering trees, shrubs or mallees in the myrtle family, Myrtaceae. Along with several other genera in the tribe Eucalypteae, including ''Corymbia'', they are commonly known as euc ...
'', a forest tree, showing crown dimension,
Tasmania ) , nickname = , image_map = Tasmania in Australia.svg , map_caption = Location of Tasmania in AustraliaCoordinates: , subdivision_type = Country , subdi ...
TocumwalTownBeach.jpg, '' E. camaldulensis'', immature woodland trees, showing collective crown habit,
Murray River The Murray River (in South Australia: River Murray) (Ngarrindjeri: ''Millewa'', Yorta Yorta: ''Tongala'') is a river in Southeastern Australia. It is Australia's longest river at extent. Its tributaries include five of the next six longest ...
,
Tocumwal Tocumwal ( ) is a town in the southern Riverina region of New South Wales, Australia, in the Berrigan Shire local government area, near the Victorian border. The town is situated on the banks of the Murray River, north of the city of Melbourne ...
,
New South Wales ) , nickname = , image_map = New South Wales in Australia.svg , map_caption = Location of New South Wales in AustraliaCoordinates: , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , es ...
Eucalyptus cretata.jpg, '' E. cretata'', juvenile, showing low branching 'mallee' form, Melbourne,
Victoria Victoria most commonly refers to: * Victoria (Australia), a state of the Commonwealth of Australia * Victoria, British Columbia, provincial capital of British Columbia, Canada * Victoria (mythology), Roman goddess of Victory * Victoria, Seychelle ...
Eucalyptus angustissima1.jpg, '' E. angustissima'', showing shrub form, Melbourne Eucalyptus platypus.jpg, '' E. platypus'', showing ‘marlock’ form, Melbourne


Bark

All eucalypts add a layer of bark every year and the outermost layer dies. In about half of the species, the dead bark is shed exposing a new layer of fresh, living bark. The dead bark may be shed in large slabs, in ribbons or in small flakes. These species are known as "smooth barks" and include '' E. sheathiana'', '' E. diversicolor'', '' E. cosmophylla'' and '' E. cladocalyx''. The remaining species retain the dead bark which dries out and accumulates. In some of these species, the fibres in the bark are loosely intertwined (in stringybarks such as '' E. macrorhyncha'' or peppermints such as '' E. radiata'') or more tightly adherent (as in the "boxes" such as '' E. leptophleba''). In some species (the "ironbarks" such as '' E. crebra'' and '' E. jensenii'') the rough bark is infused with gum resin. Many species are ‘half-barks’ or ‘blackbutts’ in which the dead bark is retained in the lower half of the trunks or stems—for example, '' E. brachycalyx'', '' E. ochrophloia'', and '' E. occidentalis''—or only in a thick, black accumulation at the base, as in '' E. clelandii''. In some species in this category, for example '' E. youngiana'' and '' E. viminalis'', the rough basal bark is very ribbony at the top, where it gives way to the smooth upper stems. The smooth upper bark of the half-barks and that of the completely smooth-barked trees and mallees can produce remarkable colour and interest, for example '' E. deglupta''. '' E. globulus'' bark cells are able to photosynthesize in the absence of foliage, conferring an "increased capacity to re-fix internal CO2 following partial defoliation". This allows the tree to grow in less-than-ideal climates, in addition to providing a better chance of recovery from damage sustained to its leaves in an event such as a fire. Different commonly recognised types of bark include: * Stringybark—consists of long fibres and can be pulled off in long pieces. It is usually thick with a spongy texture. *
Ironbark Ironbark is a common name of a number of species in three taxonomic groups within the genus ''Eucalyptus'' that have dark, deeply furrowed bark. Instead of being shed annually as in many of the other species of ''Eucalyptus'', the dead bark accum ...
—is hard, rough, and deeply furrowed. It is impregnated with dried kino (a
sap Sap is a fluid transported in xylem cells (vessel elements or tracheids) or phloem sieve tube elements of a plant. These cells transport water and nutrients throughout the plant. Sap is distinct from latex, resin, or cell sap; it is a separ ...
exuded by the tree) which gives a dark red or even black colour. * Tessellated—bark is broken up into many distinct flakes. They are corkish and can flake off. * Box—has short fibres. Some also show tessellation. * Ribbon—has the bark coming off in long, thin pieces, but is still loosely attached in some places. They can be long ribbons, firmer strips, or twisted curls. Apple box bark.jpg, Bark detail of '' E. angophoroides'', the apple-topped box Eucalyptus deglupta-trees.jpg, Coloured bark of '' E. deglupta'' native to Southeast Asia Eucalyptus bark.jpg, 'Box' bark of '' E. quadrangulata'', the white-topped box Eucalyptus sideroxylon - bark.jpg, Dark, fissured '
ironbark Ironbark is a common name of a number of species in three taxonomic groups within the genus ''Eucalyptus'' that have dark, deeply furrowed bark. Instead of being shed annually as in many of the other species of ''Eucalyptus'', the dead bark accum ...
' of '' E. sideroxylon''


Leaves

Nearly all ''Eucalyptus'' are evergreen, but some tropical species lose their leaves at the end of the dry season. As in other members of the myrtle family, ''Eucalyptus'' leaves are covered with oil glands. The copious oils produced are an important feature of the genus. Although mature ''Eucalyptus'' trees may be towering and fully leafed, their shade is characteristically patchy because the leaves usually hang downwards. The leaves on a mature ''Eucalyptus'' plant are commonly lanceolate, petiolate, apparently alternate and waxy or glossy green. In contrast, the leaves of seedlings are often opposite,
sessile Sessility, or sessile, may refer to: * Sessility (motility), organisms which are not able to move about * Sessility (botany), flowers or leaves that grow directly from the stem or peduncle of a plant * Sessility (medicine), tumors and polyps that ...
and glaucous, but many exceptions to this pattern exist. Many species such as '' E. melanophloia'' and ''E. setosa'' retain the juvenile leaf form even when the plant is reproductively mature. Some species, such as '' E. macrocarpa'', '' E. rhodantha'', and '' E. crucis'', are sought-after ornamentals due to this lifelong juvenile leaf form. A few species, such as '' E. petraea'', '' E. dundasii'', and '' E. lansdowneana'', have shiny green leaves throughout their life cycle. '' E. caesia'' exhibits the opposite pattern of leaf development to most ''Eucalyptus'', with shiny green leaves in the seedling stage and dull, glaucous leaves in mature crowns. The contrast between juvenile and adult leaf phases is valuable in field identification. Four leaf phases are recognised in the development of a ''Eucalyptus'' plant: the ‘seedling’, ‘juvenile’, ‘intermediate’, and ‘adult’ phases. However, no definite transitional point occurs between the phases. The intermediate phase, when the largest leaves are often formed, links the juvenile and adult phases.Brooker & Kleinig (2001) In all except a few species, the leaves form in pairs on opposite sides of a square stem, consecutive pairs being at right angles to each other (decussate). In some narrow-leaved species, for example '' E. oleosa'', the seedling leaves after the second leaf pair are often clustered in a detectable spiral arrangement about a five-sided stem. After the spiral phase, which may last from several to many nodes, the arrangement reverts to decussate by the absorption of some of the leaf-bearing faces of the stem. In those species with opposite adult foliage the leaf pairs, which have been formed opposite at the stem apex, become separated at their bases by unequal elongation of the stem to produce the apparently alternate adult leaves.


Flowers and fruits

The most readily recognisable characteristics of ''Eucalyptus'' species are the distinctive flowers and fruit (capsules or "gumnuts"). Flowers have numerous fluffy stamens which may be white, cream, yellow, pink, or red; in bud, the stamens are enclosed in a cap known as an operculum which is composed of the fused sepals or petals, or both. Thus, flowers have no petals, but instead decorate themselves with the many showy stamens. As the stamens expand, the operculum is forced off, splitting away from the cup-like base of the flower; this is one of the features that unites the genus. The woody fruits or capsules are roughly cone-shaped and have valves at the end which open to release the seeds, which are waxy, rod-shaped, about 1 mm in length, and yellow-brown in colour. Most species do not flower until adult foliage starts to appear; ''E. cinerea'' and ''E. perriniana'' are notable exceptions.


Taxonomy

The genus ''Eucalyptus'' was first formally described in 1789 by
Charles Louis L'Héritier de Brutelle Charles Louis L'Héritier de Brutelle (; 15 June 1746 – 18 August 1800) was an 18th-century French botanist and civil servant. Born into an affluent upper-class Parisian family, connections with the French Royal Court secured him the position of ...
who published the description in his book ''Sertum Anglicum, seu, Plantae rariores quae in hortis juxta Londinum'' along with a description of the type species, ''
Eucalyptus obliqua ''Eucalyptus obliqua'', commonly known as messmate stringybark or messmate, but also known as brown top, brown top stringbark, stringybark or Tasmanian oak, is a species of tree that is endemic to south-eastern Australia. It has rough, stringy or ...
''. The name ''Eucalyptus'' is derived from the
Ancient Greek Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic p ...
words meaning 'good, well, true, beautiful, very' and meaning '(I) cover, conceal, hide' referring to the operculum covering the flower buds. The type specimen was collected in 1777 by David Nelson, the gardener-botanist on Cook's third voyage. He collected the specimen on
Bruny Island Bruny Island ( Nuenonne: Lunawanna-alonnah) is a island located off the south-eastern coast of Tasmania, Australia. The island is separated from the Tasmanian mainland by the D'Entrecasteaux Channel, and its east coast lies within the Tasman ...
and sent it to de Brutelle who was working in
London London is the capital and List of urban areas in the United Kingdom, largest city of England and the United Kingdom, with a population of just under 9 million. It stands on the River Thames in south-east England at the head of a estuary dow ...
at that time.


History

Although eucalypts must have been seen by the very early European explorers and collectors, no botanical collections of them are known to have been made until 1770 when Joseph Banks and
Daniel Solander Daniel Carlsson Solander or Daniel Charles Solander (19 February 1733 – 13 May 1782) was a Swedish naturalist and an apostle of Carl Linnaeus. Solander was the first university-educated scientist to set foot on Australian soil. Biography ...
arrived at Botany Bay with
Captain James Cook James Cook (7 November 1728 Old Style date: 27 October – 14 February 1779) was a British explorer, navigator, cartographer, and captain in the British Royal Navy, famous for his three voyages between 1768 and 1779 in the Pacific Ocean and ...
. There they collected specimens of ''E. gummifera'' and later, near the
Endeavour River The Endeavour River ( Guugu Yimithirr: ''Wabalumbaal''), inclusive of the Endeavour River Right Branch, the Endeavour River South Branch, and the Endeavour River North Branch, is a river system located on Cape York Peninsula in Far North Queens ...
in northern ''Queensland'', ''E. platyphylla''; neither of these species was named as such at the time. In 1777, on Cook's third expedition, David Nelson collected a eucalypt on
Bruny Island Bruny Island ( Nuenonne: Lunawanna-alonnah) is a island located off the south-eastern coast of Tasmania, Australia. The island is separated from the Tasmanian mainland by the D'Entrecasteaux Channel, and its east coast lies within the Tasman ...
in southern
Tasmania ) , nickname = , image_map = Tasmania in Australia.svg , map_caption = Location of Tasmania in AustraliaCoordinates: , subdivision_type = Country , subdi ...
. This specimen was taken to the British Museum in
London London is the capital and List of urban areas in the United Kingdom, largest city of England and the United Kingdom, with a population of just under 9 million. It stands on the River Thames in south-east England at the head of a estuary dow ...
, and was named ''Eucalyptus obliqua'' by the French botanist L'Héritier, who was working in London at the time. He coined the generic name from the Greek roots ''eu'' and ''calyptos'', meaning "well" and "covered" in reference to the operculum of the flower bud which protects the developing flower parts as the flower develops and is shed by the pressure of the emerging
stamens The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filame ...
at flowering. It was most likely an accident that L'Héritier chose a feature common to all eucalypts. The name ''obliqua'' was derived from the Latin ''obliquus'', meaning "oblique", which is the botanical term describing a leaf base where the two sides of the leaf blade are of unequal length and do not meet the petiole at the same place. ''E. obliqua'' was published in 1788–89, which coincided with the first official European settlement of Australia. Between then and the turn of the 19th century, several more species of ''Eucalyptus'' were named and published. Most of these were by the English botanist James Edward Smith and most were, as might be expected, trees of the Sydney region. These include the economically valuable ''E. pilularis'', ''E. saligna'' and ''E. tereticornis''. The first endemic Western Australian ''Eucalyptus'' to be collected and subsequently named was the Yate ('' E. cornuta'') by the French botanist
Jacques Labillardière Jacques-Julien Houtou de Labillardière (28 October 1755 – 8 January 1834) was a French biologist noted for his descriptions of the flora of Australia. Labillardière was a member of a voyage in search of the La Pérouse expedition. He pub ...
, who collected in what is now the Esperance area in 1792. Several Australian botanists were active during the 19th century, particularly Ferdinand von Mueller, whose work on ''eucalypts'' contributed greatly to the first comprehensive account of the genus in
George Bentham George Bentham (22 September 1800 – 10 September 1884) was an English botanist, described by the weed botanist Duane Isely as "the premier systematic botanist of the nineteenth century". Born into a distinguished family, he initially studie ...
's ''Flora Australiensis'' in 1867. The account is the most important early systematic treatment of the genus. Bentham divided it into five series whose distinctions were based on characteristics of the stamens, particularly the anthers (Mueller, 1879–84), work elaborated by Joseph Henry Maiden (1903–33) and still further by
William Faris Blakely William Faris Blakely (November 1875 – 1 September 1941) was an Australian botanist and collector. From 1913 to 1940 he worked in the National Herbarium of New South Wales, working with Joseph Maiden on ''Eucalyptus'', Maiden named a ''red g ...
(1934). The anther system became too complex to be workable and more recent systematic work has concentrated on the characteristics of buds, fruits, leaves and bark.


Species and hybrids

Over 700 species of ''Eucalyptus'' are known. Some have diverged from the mainstream of the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
to the extent that they are quite isolated
genetically Genetics is the study of genes, genetic variation, and heredity in organisms.Hartl D, Jones E (2005) It is an important branch in biology because heredity is vital to organisms' evolution. Gregor Mendel, a Moravian Augustinian friar working i ...
and are able to be recognised by only a few relatively invariant characteristics. Most, however, may be regarded as belonging to large or small groups of related species, which are often in geographical contact with each other and between which gene exchange still occurs. In these situations, many species appear to grade into one another, and intermediate forms are common. In other words, some species are relatively fixed genetically, as expressed in their
morphology Morphology, from the Greek and meaning "study of shape", may refer to: Disciplines * Morphology (archaeology), study of the shapes or forms of artifacts * Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies ...
, while others have not diverged completely from their nearest relatives. Hybrid individuals have not always been recognised as such on first collection and some have been named as new species, such as ''E. chrysantha'' (''E. preissiana'' × ''E. sepulcralis'') and ''E.'' "rivalis" (''E. marginata'' × ''E. megacarpa''). Hybrid combinations are not particularly common in the field, but some other published species frequently seen in Australia have been suggested to be hybrid combinations. For example, '' Eucalyptus × erythrandra'' is believed to be ''E. angulosa'' × ''E. teraptera'' and due to its wide distribution is often referred to in texts. Renantherin, a phenolic compound present in the leaves of some ''Eucalyptus'' species, allows chemotaxonomic discrimination in the sections renantheroideae and renantherae and the ratio of the amount of leucoanthocyanins varies considerably in certain species.


Related genera

''Eucalyptus'' is one of three similar genera that are commonly referred to as "
eucalypt Eucalypt is a descriptive name for woody plants with capsule fruiting bodies belonging to seven closely related genera (of the tribe Eucalypteae) found across Australasia: ''Eucalyptus'', ''Corymbia'', ''Angophora'', '' Stockwellia'', ''Allosyn ...
s", the others being '' Corymbia'' and '' Angophora''. Many species, though by no means all, are known as gum trees because they exude copious kino from any break in the bark (e.g., scribbly gum). The generic name is derived from the
Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...
words ευ (''eu'') "well" and καλύπτω (''kalýpto'') "to cover", referring to the operculum on the calyx that initially conceals the flower.


Distribution

There are more than 700 species of ''Eucalyptus'' and most are native to Australia; a very small number are found in adjacent areas of New Guinea and Indonesia. One species, ''
Eucalyptus deglupta ''Eucalyptus deglupta'' is a species of tall tree, commonly known as the rainbow eucalyptus, Mindanao gum, or rainbow gum that is native to the Philippines, Indonesia, and Papua New Guinea. It is the only ''Eucalyptus'' species that usually liv ...
,'' ranges as far north as the Philippines. Of the 15 species found outside Australia, just nine are exclusively non-Australian. Species of ''Eucalyptus'' are cultivated widely in the tropical and temperate world, including the Americas,
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a subcontinent of Eurasia and it is located entirel ...
,
Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...
, the Mediterranean Basin, the Middle East, China, and the
Indian subcontinent The Indian subcontinent is a list of the physiographic regions of the world, physiographical region in United Nations geoscheme for Asia#Southern Asia, Southern Asia. It is situated on the Indian Plate, projecting southwards into the Indian O ...
. However, the range over which many eucalypts can be planted in the temperate zone is constrained by their limited cold tolerance. Australia is covered by of eucalypt forest, comprising three quarters of the area covered by native forest. The Blue Mountains of southeastern Australia have been a centre of eucalypt diversification; their name is in reference to the blue haze prevalent in the area, believed derived from the volatile
terpenoid The terpenoids, also known as isoprenoids, are a class of naturally occurring organic chemicals derived from the 5-carbon compound isoprene and its derivatives called terpenes, diterpenes, etc. While sometimes used interchangeably with "terpenes" ...
s emitted by these trees.


Fossil record

The oldest definitive ''Eucalyptus'' fossils are from
Patagonia Patagonia () refers to a geographical region that encompasses the southern end of South America, governed by Argentina and Chile. The region comprises the southern section of the Andes Mountains with lakes, fjords, temperate rainforests, and g ...
in South America, where eucalypts are no longer native, though they have been introduced from Australia. The fossils are from the early Eocene (51.9 Mya), and were found in the Laguna del Hunco Formation in Chubut Province in Argentina. This shows that the genus had a Gondwanan distribution. Fossil leaves also occur in the Miocene of New Zealand, where the genus is not native today, but again have been introduced from Australia. Despite the prominence of ''Eucalyptus'' in modern Australia, estimated to contribute some 75% of the modern vegetation, the fossil record is very scarce throughout much of the Cenozoic, and suggests that this rise to dominance is a geologically more recent phenomenon. The oldest reliably dated macrofossil of ''Eucalyptus'' is a 21-million-year-old tree-stump encased in basalt in the upper Lachlan Valley in
New South Wales ) , nickname = , image_map = New South Wales in Australia.svg , map_caption = Location of New South Wales in AustraliaCoordinates: , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , es ...
. Other fossils have been found, but many are either unreliably dated or else unreliably identified.Hill, R. S. "The history of selected Australian taxa." History of the Australian vegetation: Cretaceous to Recent (1994): 390. It is useful to consider where ''Eucalyptus'' fossils have not been found. Extensive research has gone into the fossil floras of the Paleocene to Oligocene of South-Eastern Australia, and has failed to uncover a single ''Eucalyptus'' specimen. Although the evidence is sparse, the best hypothesis is that in the mid-Tertiary, the continental margins of Australia only supported more mesic noneucalypt vegetation, and that eucalypts probably contributed to the drier vegetation of the arid continental interior. With the progressive drying out of the continent since the
Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...
, eucalypts were displaced to the continental margins, and much of the mesic and rainforest vegetation that was once there was eliminated entirely. The current superdominance of ''Eucalyptus'' in Australia may be an artefact of human influence on its ecology. In more recent sediments, numerous findings of a dramatic increase in the abundance of ''Eucalyptus'' pollen are associated with increased charcoal levels. Though this occurs at different rates throughout Australia, it is compelling evidence for a relationship between the artificial increase of fire frequency with the arrival of Aboriginals and increased prevalence of this exceptionally fire-tolerant genus.


Tall timber

Several eucalypt species are among the tallest trees in the world. ''
Eucalyptus regnans ''Eucalyptus'' () is a genus of over seven hundred species of flowering trees, shrubs or mallees in the myrtle family, Myrtaceae. Along with several other genera in the tribe Eucalypteae, including ''Corymbia'', they are commonly known as euc ...
'', the Australian 'mountain ash', is the tallest of all flowering plants ( angiosperms); today, the tallest measured specimen named Centurion is tall.
Coast Douglas-fir ''Pseudotsuga menziesii'' var. ''menziesii'', commonly known as Coast Douglas-fir, Pacific Douglas-fir, Oregon pine, or Douglas spruce, is an evergreen conifer native to western North America from west-central British Columbia, Canada southward t ...
is about the same height; only
coast redwood ''Sequoia sempervirens'' ()''Sunset Western Garden Book,'' 1995:606–607 is the sole living species of the genus '' Sequoia'' in the cypress family Cupressaceae (formerly treated in Taxodiaceae). Common names include coast redwood, coastal ...
is taller, and they are conifers ( gymnosperms). Six other eucalypt species exceed 80 metres in height: ''
Eucalyptus obliqua ''Eucalyptus obliqua'', commonly known as messmate stringybark or messmate, but also known as brown top, brown top stringbark, stringybark or Tasmanian oak, is a species of tree that is endemic to south-eastern Australia. It has rough, stringy or ...
'', '' Eucalyptus delegatensis'', ''
Eucalyptus diversicolor ''Eucalyptus diversicolor'', commonly known as karri, is a species of flowering plant in the family Myrtaceae and is endemic to the south-west of Western Australia. It is a tall tree with smooth light grey to cream-coloured, often mottled bark ...
'', '' Eucalyptus nitens'', ''
Eucalyptus globulus ''Eucalyptus globulus'', commonly known as southern blue gum or blue gum, is a species of tall, evergreen tree endemic to southeastern Australia. This ''Eucalyptus'' species has mostly smooth bark, juvenile leaves that are whitish and waxy on ...
'' and ''
Eucalyptus viminalis ''Eucalyptus viminalis'', commonly known as the manna gum, white gum or ribbon gum, is a species of small to very tall tree that is endemic to south-eastern Australia. It has smooth bark, sometimes with rough bark near the base, lance-shaped to c ...
''.


Frost intolerance

Most eucalypts are not tolerant of severe cold. Eucalypts do well in a range of climates but are usually damaged by anything beyond a light frost of ; the hardiest are the snow gums, such as '' Eucalyptus pauciflora'', which is capable of withstanding cold and frost down to about . Two subspecies, ''E. pauciflora subsp. niphophila'' and ''E. pauciflora subsp. debeuzevillei'' in particular are even hardier and can tolerate even quite severe winters. Several other species, especially from the high plateau and mountains of central
Tasmania ) , nickname = , image_map = Tasmania in Australia.svg , map_caption = Location of Tasmania in AustraliaCoordinates: , subdivision_type = Country , subdi ...
such as '' Eucalyptus coccifera'', '' Eucalyptus subcrenulata'' and '' Eucalyptus gunnii'', have also produced extreme cold-hardy forms and it is seed procured from these
genetically Genetics is the study of genes, genetic variation, and heredity in organisms.Hartl D, Jones E (2005) It is an important branch in biology because heredity is vital to organisms' evolution. Gregor Mendel, a Moravian Augustinian friar working i ...
hardy strains that are planted for ornament in colder parts of the world.


Animal relationships

An essential oil extracted from ''Eucalyptus'' leaves contains compounds that are powerful natural disinfectants and can be toxic in large quantities. Several marsupial herbivores, notably
koala The koala or, inaccurately, koala bear (''Phascolarctos cinereus''), is an arboreal herbivorous marsupial native to Australia. It is the only extant representative of the family Phascolarctidae and its closest living relatives are the w ...
s and some possums, are relatively tolerant of it. The close correlation of these oils with other more potent toxins called formylated
phloroglucinol Phloroglucinol is an organic compound with the formula C6H3(OH)3. It is a colorless solid. It is used in the synthesis of pharmaceuticals and explosives. Phloroglucinol is one of three isomeric benzenetriols. The other two isomers are hydroxyqu ...
compounds ( euglobals, macrocarpals and sideroxylonals) allows koalas and other marsupial species to make food choices based on the smell of the leaves. For koalas, these compounds are the most important factor in leaf choice. ''Eucalyptus'' flowers produce a great abundance of nectar, providing food for many pollinators including
insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three ...
s, birds, bats and possums. Many lizard species in Australia feed on ''Eucalyptus'' sap as well, famously in the case of the Dubious dtella. Although ''Eucalyptus'' trees are seemingly well-defended from herbivores by the oils and phenolic compounds, they have insect pests. These include the eucalyptus longhorn borer '' Phoracantha semipunctata'' and the aphid-like
psyllid Psyllidae, the jumping plant lice or psyllids, are a family of small plant-feeding insects that tend to be very host-specific, i.e. each plant-louse species only feeds on one plant species (monophagous) or feeds on a few closely related plants ...
s that create "bell lerps", both of which have become established as pests throughout the world wherever eucalypts are cultivated. The
eusocial Eusociality (from Greek εὖ ''eu'' "good" and social), the highest level of organization of sociality, is defined by the following characteristics: cooperative brood care (including care of offspring from other individuals), overlapping gen ...
beetle ''
Austroplatypus incompertus ''Austroplatypus incompertus'' is a species of ambrosia beetle belonging to the true weevil family, native to Australia, with a verified distribution in New South Wales and Victoria. It forms colonies in the heartwood of ''Eucalyptus'' trees and ...
'' makes and defends its galleries exclusively inside ''Eucalyptus'' plants. The trunks and branches of the ''Eucalyptus'' tree allow the largest known moth, ''Zelotypia stacyi'' (the bentwing ghost moth, having a wingspan up to 250 mm) to feed and protect their
larva A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. ...
and pupa, respectively.


Adaptation to fire

Eucalypts originated between 35 and 50 million years ago, not long after Australia-New Guinea separated from Gondwana, their rise coinciding with an increase in fossil charcoal deposits (suggesting that fire was a factor even then), but they remained a minor component of the Tertiary rainforest until about 20 million years ago, when the gradual drying of the continent and depletion of soil nutrients led to the development of a more open forest type, predominantly '' Casuarina'' and '' Acacia'' species. The two valuable timber trees, alpine ash '' E. delegatensis'' and Australian mountain ash '' E. regnans'', are killed by fire and only regenerate from seed. The same 2003 bushfire that had little impact on forests around Canberra resulted in thousands of hectares of dead ash forests. However, a small amount of ash survived and put out new ash trees as well.


Fire hazard

''Eucalyptus'' oil is highly flammable; ignited trees have been known to explode.
Bushfires A wildfire, forest fire, bushfire, wildland fire or rural fire is an unplanned, uncontrolled and unpredictable fire in an area of combustible vegetation. Depending on the type of vegetation present, a wildfire may be more specifically identif ...
can travel easily through the oil-rich air of the tree crowns. Eucalypts obtain long-term fire survivability from their ability to regenerate from epicormic buds situated deep within their thick bark, or from
lignotuber A lignotuber is a woody swelling of the root crown possessed by some plants as a protection against destruction of the plant stem, such as by fire. Other woody plants may develop basal burls as a similar survival strategy, often as a response t ...
s, or by producing
serotinous Serotiny in botany simply means 'following' or 'later'. In the case of serotinous flowers, it means flowers which grow following the growth of leaves, or even more simply, flowering later in the season than is customary with allied species. Havi ...
fruits. In seasonally dry climates oaks are often fire-resistant, particularly in open grasslands, as a grass fire is insufficient to ignite the scattered trees. In contrast, a ''Eucalyptus'' forest tends to promote fire because of the volatile and highly combustible oils produced by the leaves, as well as the production of large amounts of
litter Litter consists of waste products that have been discarded incorrectly, without consent, at an unsuitable location. Litter can also be used as a verb; to litter means to drop and leave objects, often man-made, such as aluminum cans, paper cups ...
high in phenolics, preventing its breakdown by fungi and thus accumulating as large amounts of dry, combustible fuel.Reid, J.B. & Potts, B.M. (2005). Eucalypt Biology. In: Reid ''et al.'' (eds.) Vegetation of Tasmania., pp. 198-223. Australian Government. Consequently, dense eucalypt plantings may be subject to catastrophic firestorms. In fact, almost thirty years before the
Oakland firestorm of 1991 : The Oakland firestorm of 1991 was a large suburban wildland–urban interface conflagration that occurred on the hillsides of northern Oakland, California, and southeastern Berkeley over the weekend of October 19–20, 1991, before being b ...
, a study of ''Eucalyptus'' in the area warned that the litter beneath the trees builds up very rapidly and should be regularly monitored and removed. It has been estimated that 70% of the energy released through the combustion of vegetation in the Oakland fire was due to ''Eucalyptus''. In a National Park Service study, it was found that the fuel load (in tons per acre) of non-native ''Eucalyptus'' woods is almost three times as great as native oak woodland. During
World War II World War II or the Second World War, often abbreviated as WWII or WW2, was a world war that lasted from 1939 to 1945. It involved the vast majority of the world's countries—including all of the great powers—forming two opposing ...
, one California town cut down their ''Eucalyptus'' trees to “about a third of their height in the vicinity of
anti-aircraft guns Anti-aircraft warfare, counter-air or air defence forces is the battlespace response to aerial warfare, defined by NATO as "all measures designed to nullify or reduce the effectiveness of hostile air action".AAP-6 It includes surface based, ...
” because of the known fire-fueling qualities of the trees, with the mayor telling a newspaper reporter, “If a
shell Shell may refer to: Architecture and design * Shell (structure), a thin structure ** Concrete shell, a thin shell of concrete, usually with no interior columns or exterior buttresses ** Thin-shell structure Science Biology * Seashell, a hard o ...
so much as hits a leaf, it’s supposed to explode.”


Falling branches

Some species of ''Eucalyptus'' drop branches unexpectedly. In Australia, Parks Victoria warns campers not to camp under river red gums. Some councils in Australia such as
Gosnells, Western Australia Gosnells is a suburb located within the City of Gosnells. Gosnells is approximately south-east of the Perth central business district. It contains the Gosnells town centre which includes the Council offices, library and the Gosnells Railway ...
, have removed eucalypts after reports of damage from dropped branches, even in the face of lengthy, well publicised protests to protect particular trees. A former Australian National Botanic Gardens director and consulting arborist, Robert Boden, has been quoted referring to "summer branch drop". Dropping of branches is recognised in Australia literature through the fictional death of Judy in
Seven Little Australians ''Seven Little Australians'' is a classic Australian children's literature novel by Ethel Turner, published in 1894. Set mainly in Sydney in the 1880s, it relates the adventures of the seven mischievous Woolcot children, their stern army father ...
. Although all large trees can drop branches, the density of ''Eucalyptus'' wood is high due to its high resin content, increasing the hazard.


Cultivation and uses

Eucalypts were introduced from Australia to the rest of the world following the Cook expedition in 1770. Collected by
Sir Joseph Banks Sir Joseph Banks, 1st Baronet, (19 June 1820) was an English naturalist, botanist, and patron of the natural sciences. Banks made his name on the 1766 natural-history expedition to Newfoundland and Labrador. He took part in Captain James C ...
, botanist on the expedition, they were subsequently introduced to many parts of the world, notably California, southern Europe, Africa, the Middle East, South Asia and South America. About 250 species are under cultivation in California. In
Portugal Portugal, officially the Portuguese Republic ( pt, República Portuguesa, links=yes ), is a country whose mainland is located on the Iberian Peninsula of Southwestern Europe, and whose territory also includes the Atlantic archipelagos of ...
and also
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = ''Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , i ...
, ''eucalypts'' have been grown in plantations for the production of pulpwood. ''Eucalyptus'' are the basis for several industries, such as sawmilling, pulp, charcoal and others. Several species have become invasive and are causing major problems for local ecosystems, mainly due to the absence of
wildlife corridor A wildlife corridor, habitat corridor, or green corridor is an area of habitat connecting wildlife populations separated by human activities or structures (such as roads, development, or logging). This allows an exchange of individuals between ...
s and rotations management. Eucalypts have many uses which have made them economically important trees, and they have become a cash crop in poor areas such as Timbuktu, Mali WorldWatch Institute. (2007) '' State of the World: Our Urban Future''. and the
Peru , image_flag = Flag of Peru.svg , image_coat = Escudo nacional del Perú.svg , other_symbol = Great Seal of the State , other_symbol_type = National seal , national_motto = "Firm and Happy f ...
vian
Andes The Andes, Andes Mountains or Andean Mountains (; ) are the longest continental mountain range in the world, forming a continuous highland along the western edge of South America. The range is long, wide (widest between 18°S – 20°S ...
,Luzar J. (2007). The Political Ecology of a "Forest Transition": Eucalyptus forestry in the Southern Peruvian. ''Ethnobotany Research & Applications''. despite concerns that the trees are invasive in some environments like those of
South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. It is bounded to the south by of coastline that stretch along the Atlantic Ocean, South Atlantic and Indian Oceans; to the ...
. Best-known are perhaps the varieties karri and
yellow box ''Eucalyptus melliodora'', commonly known as yellow box, honey box or yellow ironbark, is a species of medium-sized to occasionally tall tree that is Endemism, endemic to south-eastern, continental Australia. It has rough, flaky or fibrous bark ...
. Due to their fast growth, the foremost benefit of these trees is their wood. They can be chopped off at the root and grow back again. They provide many desirable characteristics for use as ornament, timber, firewood and pulpwood. Eucaplytus wood is also used in a number of industries, from fence posts (where the oil-rich wood's high resistance to decay is valued) and charcoal to
cellulose Cellulose is an organic compound with the formula , a polysaccharide consisting of a linear chain of several hundred to many thousands of β(1→4) linked D-glucose units. Cellulose is an important structural component of the primary cell w ...
extraction for biofuels. Fast growth also makes eucalypts suitable as windbreaks and to reduce
erosion Erosion is the action of surface processes (such as water flow or wind) that removes soil, rock, or dissolved material from one location on the Earth's crust, and then transports it to another location where it is deposited. Erosion is dis ...
. Some ''Eucalyptus'' species have attracted attention from horticulturists,
global development International development or global development is a broad concept denoting the idea that societies and countries have differing levels of economic or human development on an international scale. It is the basis for international classifications ...
researchers, and environmentalists because of desirable traits such as being fast-growing sources of wood, producing oil that can be used for cleaning and as a natural insecticide, or an ability to be used to drain swamps and thereby reduce the risk of malaria.
Eucalyptus oil Eucalyptus oil is the generic name for distilled oil from the leaf of ''Eucalyptus'', a genus of the plant family Myrtaceae native to Australia and cultivated worldwide. Eucalyptus oil has a history of wide application, as a pharmaceutical, anti ...
finds many uses like in fuels, fragrances, insect repellence and antimicrobial activity. ''Eucalyptus'' trees show allelopathic effects; they release compounds which inhibit other plant species from growing nearby. Outside their natural ranges, eucalypts are both lauded for their beneficial economic impact on poor populations and criticised for being "water-guzzling" aliens, leading to controversy over their total impact. Eucalypts draw a tremendous amount of water from the soil through the process of transpiration. They have been planted (or re-planted) in some places to lower the water table and reduce soil salination. Eucalypts have also been used as a way of reducing malaria by draining the soil in Algeria, Lebanon, Sicily, elsewhere in
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a subcontinent of Eurasia and it is located entirel ...
, in the Caucasus (Western
Georgia Georgia most commonly refers to: * Georgia (country), a country in the Caucasus region of Eurasia * Georgia (U.S. state), a state in the Southeast United States Georgia may also refer to: Places Historical states and entities * Related to the ...
), and California. Drainage removes swamps which provide a habitat for
mosquito Mosquitoes (or mosquitos) are members of a group of almost 3,600 species of small flies within the family Culicidae (from the Latin ''culex'' meaning " gnat"). The word "mosquito" (formed by ''mosca'' and diminutive ''-ito'') is Spanish for "li ...
larva A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. ...
e, but can also destroy ecologically productive areas. This drainage is not limited to the soil surface, because the ''Eucalyptus'' roots are up to in length and can, depending on the location, even reach the
phreatic zone The phreatic zone, saturated zone, or zone of saturation, is the part of an aquifer, below the water table, in which relatively all pores and fractures are saturated with water. Above the water table is the unsaturated or vadose zone The v ...
.


Pulpwood

''Eucalyptus'' is the most common short fibre source for pulpwood to make
pulp Pulp may refer to: * Pulp (fruit), the inner flesh of fruit Engineering * Dissolving pulp, highly purified cellulose used in fibre and film manufacture * Pulp (paper), the fibrous material used to make paper * Molded pulp, a packaging material ...
. The types most often used in papermaking are ''
Eucalyptus globulus ''Eucalyptus globulus'', commonly known as southern blue gum or blue gum, is a species of tall, evergreen tree endemic to southeastern Australia. This ''Eucalyptus'' species has mostly smooth bark, juvenile leaves that are whitish and waxy on ...
'' (in temperate areas) and the '' Eucalyptus urophylla'' x '' Eucalyptus grandis'' hybrid (in the tropics). The fibre length of ''Eucalyptus'' is relatively short and uniform with low coarseness compared with other hardwoods commonly used as pulpwood. The fibres are slender, yet relatively thick walled. This gives uniform paper formation and high opacity that are important for all types of fine papers. The low coarseness is important for high quality coated papers. ''Eucalyptus'' is suitable for many tissue papers as the short and slender fibres gives a high number of fibres per gram and low coarseness contributes to softness.


''Eucalyptus'' oil

Eucalyptus oil Eucalyptus oil is the generic name for distilled oil from the leaf of ''Eucalyptus'', a genus of the plant family Myrtaceae native to Australia and cultivated worldwide. Eucalyptus oil has a history of wide application, as a pharmaceutical, anti ...
is readily steam distilled from the leaves and can be used for cleaning and as an industrial solvent, as an antiseptic, for deodorising, and in very small quantities in food supplements, especially
sweets Candy, also called sweets (British English) or lollies (Australian English, New Zealand English), is a confection that features sugar as a principal ingredient. The category, called ''sugar confectionery'', encompasses any sweet confection, i ...
,
cough drops A throat lozenge (also known as a cough drop, sore throat sweet, troche, cachou, pastille or cough sweet) is a small, typically medicated tablet intended to be dissolved slowly in the mouth to temporarily stop coughs, lubricate, and soothe irri ...
, toothpaste and decongestants. It has insect-repellent properties, and serves as an active ingredient in some commercial mosquito-repellents. Aromatherapists have adopted ''Eucalyptus'' oils for a wide range of purposes. ''
Eucalyptus globulus ''Eucalyptus globulus'', commonly known as southern blue gum or blue gum, is a species of tall, evergreen tree endemic to southeastern Australia. This ''Eucalyptus'' species has mostly smooth bark, juvenile leaves that are whitish and waxy on ...
'' is the principal source of ''Eucalyptus'' oil worldwide.


Musical instruments

Eucalypt wood is also commonly used to make didgeridoos, a traditional
Australian Aboriginal Aboriginal Australians are the various Indigenous peoples of the Australian mainland and many of its islands, such as Tasmania, Fraser Island, Hinchinbrook Island, the Tiwi Islands, and Groote Eylandt, but excluding the Torres Strait I ...
wind instrument. The trunk of the tree is hollowed out by termites, and then cut down if the bore is of the correct size and shape.


Dyes

All parts of ''Eucalyptus'' may be used to make dyes that are substantive on
protein Proteins are large biomolecules and macromolecules that comprise one or more long chains of amino acid residues. Proteins perform a vast array of functions within organisms, including catalysing metabolic reactions, DNA replication, res ...
fibres (such as silk and wool), simply by processing the plant part with
water Water (chemical formula ) is an Inorganic compound, inorganic, transparent, tasteless, odorless, and Color of water, nearly colorless chemical substance, which is the main constituent of Earth's hydrosphere and the fluids of all known living ...
. Colours to be achieved range from yellow and orange through green, tan, chocolate and deep rust red. The material remaining after processing can be safely used as
mulch A mulch is a layer of material applied to the surface of soil. Reasons for applying mulch include conservation of soil moisture, improving fertility and health of the soil, reducing weed growth and enhancing the visual appeal of the area. A mu ...
or
fertiliser A fertilizer (American English) or fertiliser (British English; see spelling differences) is any material of natural or synthetic origin that is applied to soil or to plant tissues to supply plant nutrients. Fertilizers may be distinct from ...
.


Prospecting

''Eucalyptus'' trees in the Australian outback draw up gold from tens of metres underground through their root system and deposit it as particles in their leaves and branches. A Maia detector for x-ray elemental imaging at the Australian Synchrotron clearly showed deposits of gold and other metals in the structure of ''Eucalyptus'' leaves from the Kalgoorlie region of Western Australia that would have been untraceable using other methods. The microscopic leaf-bound "nuggets", about 8
micrometre The micrometre ( international spelling as used by the International Bureau of Weights and Measures; SI symbol: μm) or micrometer (American spelling), also commonly known as a micron, is a unit of length in the International System of Unit ...
s wide on average, are not worth collecting themselves, but may provide an environmentally benign way of locating subsurface mineral deposits.


''Eucalyptus'' as plantation species

In the 20th century, scientists around the world experimented with ''Eucalyptus'' species. They hoped to grow them in the tropics, but most experimental results failed until breakthroughs in the 1960s-1980s in species selection, silviculture, and breeding programs "unlocked" the potential of eucalypts in the tropics. Prior to then, as Brett Bennett noted in a 2010 article, eucalypts were something of the " El Dorado" of forestry. Today, ''Eucalyptus'' is the most widely planted type of tree in plantations around the world, in South America (mainly in
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
, Argentina, Paraguay and Uruguay),
South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. It is bounded to the south by of coastline that stretch along the Atlantic Ocean, South Atlantic and Indian Oceans; to the ...
, Australia,
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
, Galicia,
Portugal Portugal, officially the Portuguese Republic ( pt, República Portuguesa, links=yes ), is a country whose mainland is located on the Iberian Peninsula of Southwestern Europe, and whose territory also includes the Atlantic archipelagos of ...
and many more.


North America

;California In the 1850s, ''Eucalyptus'' trees were introduced to California by Australians during the California Gold Rush. Much of California is similar in climate to parts of Australia. By the early 1900s, thousands of acres of eucalypts were planted with the encouragement of the state government. It was hoped that they would provide a renewable source of timber for construction, furniture making and
railway sleeper A railroad tie, crosstie (American English), railway tie (Canadian English) or railway sleeper ( Australian and British English) is a rectangular support for the rails in railroad tracks. Generally laid perpendicular to the rails, ties trans ...
s. It was soon found that for the latter purpose ''Eucalyptus'' was particularly unsuitable, as the ties made from ''Eucalyptus'' had a tendency to twist while drying, and the dried ties were so tough that it was nearly impossible to hammer rail spikes into them.
They went on to note that the promise of ''Eucalyptus'' in California was based on the old virgin forests of Australia. This was a mistake, as the young trees being harvested in California could not compare in quality to the centuries-old ''Eucalyptus'' timber of Australia. It reacted differently to harvest. The older trees didn't split or warp as the infant California crop did. There was a vast difference between the two, and this would doom the California ''Eucalyptus'' industry.
The species '' Eucalyptus rostrata, E. tereticornas,'' and '' E. cladocalyx'' are all present in California, but the blue gum '' E. globulus'' makes up by far the largest population in the state. One way in which the ''Eucalyptus'', mainly the blue gum '' E. globulus'', proved valuable in California was in providing windbreaks for highways, orange groves, and farms in the mostly treeless central part of the state. They are also admired as shade and ornamental trees in many cities and gardens. ''Eucalyptus'' plantations in California have been criticised, because they compete with native plants and typically do not support native animals. ''Eucalyptus'' has historically been planted to replace California's
coast live oak ''Quercus agrifolia'', the California live oak, or coast live oak, is a highly variable, often evergreen oak tree, a type of live oak, native to the California Floristic Province. It may be shrubby, depending on age and growing location, but is ...
population, and the new ''Eucalyptus'' is not as hospitable to native flora and fauna as the oaks. In appropriately foggy conditions on the California Coast, ''Eucalyptus'' can spread at a rapid rate. The absence of natural inhibitors such as the
koala The koala or, inaccurately, koala bear (''Phascolarctos cinereus''), is an arboreal herbivorous marsupial native to Australia. It is the only extant representative of the family Phascolarctidae and its closest living relatives are the w ...
or pathogens native to Australia have aided in the spread of California ''Eucalyptus'' trees. This is not as big of an issue further inland, but on the coast invasive eucalypts can disrupt native ecosystems. ''Eucalyptus'' may have adverse effects on local streams due to their chemical composition, and their dominance threatens species that rely on native trees. Nevertheless, some native species have been known to adapt to the ''Eucalyptus'' trees. Notable examples are herons,
great horned owl The great horned owl (''Bubo virginianus''), also known as the tiger owl (originally derived from early naturalists' description as the "winged tiger" or "tiger of the air"), or the hoot owl, is a large owl native to the Americas. It is an extre ...
, and the monarch butterfly using ''Eucalyptus'' groves as habitat. Despite these successes, eucalypts generally has a net negative impact on the overall balance of the native ecosystem. Fire is also a problem. Eucalypts has been noted for their flammable properties and the large fuel loads in the understory of eucalypt forests. ''Eucalyptus'' trees were a catalyst for the spread of the 1923 fire in Berkeley, which destroyed 568 homes. The 1991 Oakland Hills firestorm, which caused US$1.5 billion in damage, destroyed almost 3,000 homes, and killed 25 people, was partly fueled by large numbers of eucalypts close to the houses. Despite these issues, there are calls to preserve the ''Eucalyptus'' plants in California. Advocates for the tree claim its fire risk has been overstated. Some even claim that the ''Eucalyptus's'' absorption of moisture makes it a barrier against fire. These experts believe that the herbicides used to remove the ''Eucalyptus'' would negatively impact the ecosystem, and the loss of the ''Eucalypti'' would release carbon into the atmosphere unnecessarily. There is also an aesthetic argument for keeping the ''Eucalyptus''; the trees are viewed by many as an attractive and iconic part of the California landscape. Many say that although the tree is not native, it has been in California long enough to become an essential part of the ecosystem and therefore should not be attacked as invasive. These arguments have caused experts and citizens in California and the Bay Area to debate the merits of ''Eucalyptus'' removal versus preservation. However, the general consensus remains that some areas urgently require ''Eucalyptus'' management to stave off potential fire hazards. Efforts to remove some of California's ''Eucalyptus'' trees have been met with a mixed reaction from the public, and there have been protests against removal. Removing ''Eucalyptus'' trees can be expensive and often requires machinery or the use of herbicides. The trees struggle to reproduce on their own outside of the foggy regions of Coastal California, and therefore some inland ''Eucalyptus'' forests are predicted to die out naturally. In some parts of California, eucalypt plantations are being removed and native trees and plants restored. Individuals have also illegally destroyed some trees and are suspected of introducing insect pests from Australia which attack the trees. Certain ''Eucalyptus'' species may also be grown for ornament in warmer parts of the Pacific Northwest—western Washington, western
Oregon Oregon () is a state in the Pacific Northwest region of the Western United States. The Columbia River delineates much of Oregon's northern boundary with Washington, while the Snake River delineates much of its eastern boundary with Idaho. T ...
and southwestern
British Columbia British Columbia (commonly abbreviated as BC) is the westernmost province of Canada, situated between the Pacific Ocean and the Rocky Mountains. It has a diverse geography, with rugged landscapes that include rocky coastlines, sandy beaches, ...
.


South America

;Argentina It was introduced in Argentina around 1870 by President Domingo F. Sarmiento, who had brought the seeds from Australia and it quickly became very popular. The most widely planted species were '' E. globulus'', '' E. viminalis'' and '' E. rostrata''. Currently, the Humid Pampas region has small forests and ''Eucalyptus'' barriers, some up to 80 years old, about 50 meters high and a maximum of one meter in diameter. ;Uruguay Antonio Lussich introduced ''Eucalyptus'' into Uruguay in approximately 1896, throughout what is now Maldonado Department, and it has spread all over the south-eastern and eastern coast. There had been no trees in the area because it consisted of dry sand dunes and stones. Lussich also introduced many other trees, particularly '' Acacia'' and pines, but they have not expanded so extensively. Uruguayan forestry crops using ''Eucalyptus'' species have been promoted since 1989, when the new National Forestry Law established that 20% of the national territory would be dedicated to forestry. As the main landscape of Uruguay is grassland (140,000 km2, 87% of the national territory), most of the forestry plantations would be established in prairie regions.Perez-Arrarte, C., 1993. Desarrollo forestal y medio ambiente (compilation). CIEDUR Montevideo, Uruguay The planting of ''Eucalyptus'' sp. has been criticised because of concerns that soil would be degraded by nutrient depletion and other biological changes.Caffera, R.M., Cespedes, C., Gonzalez, A., Gutierrez, M.O., Panario, D.H., 1991. Hacia una evaluacion de efectos ambientales de la forestacion en Uruguay con especies introducidas. CIEDUR, Montevideo, Uruguay. During the last ten years, in the northwestern regions of Uruguay the ''Eucalyptus'' sp. plantations have reached annual forestation rates of 300%. That zone has a potential forested area of 1 million hectares, approximately 29% of the national territory dedicated to forestry, of which approximately 800,000 hectares are currently forested by monoculture of ''Eucalyptus'' spp. It is expected that the radical and durable substitution of vegetation cover leads to changes in the quantity and quality of soil organic matter. Such changes may also influence soil fertility and soil physical and chemical properties. The soil quality effects associated with ''Eucalyptus'' sp. plantations could have adverse effects on soil chemistry; for example: soil acidification, iron leaching, allelopathic activities and a high C:N ratio of litter. Additionally, as most scientific understanding of land cover change effects is related to ecosystems where forests were replaced by grasslands or crops, or grassland was replaced by crops, the environmental effects of the current Uruguayan land cover changes are not well understood. The first scientific publication on soil studies in western zone tree plantations (focused on pulp production) appeared in 2004 and described soil acidification and soil carbon changes, similar to a podzolisation process, and destruction of clay (illite-like minerals), which is the main reservoir of potassium in the soil. Although these studies were carried out in an important zone for forest cultivation, they cannot define the current situation in the rest of the land area under eucalyptus cultivation. Moreover, recently Jackson and Jobbagy have proposed another adverse environmental impact that may result from ''Eucalyptus'' culture on prairie soils—stream acidification. The ''Eucalyptus'' species most planted are ''E. grandis'', ''E. globulus'' and ''E. dunnii''; they are used mainly for pulp mills. Approximately 80,000 ha of ''E. grandis'' situated in the departments of Rivera, Tacuarembó and Paysandú is primarily earmarked for the solid wood market, although a portion of it is used for sawlogs and plywood. The current area under commercial forest plantation is 6% of the total. The main uses of the wood produced are elemental chlorine free pulp mill production (for
cellulose Cellulose is an organic compound with the formula , a polysaccharide consisting of a linear chain of several hundred to many thousands of β(1→4) linked D-glucose units. Cellulose is an important structural component of the primary cell w ...
and
paper Paper is a thin sheet material produced by mechanically or chemically processing cellulose fibres derived from wood, rags, grasses or other vegetable sources in water, draining the water through fine mesh leaving the fibre evenly distrib ...
), sawlogs, plywood and bioenergy ( thermoelectric generation). Most of the products obtained from sawmills and pulp mills, as well as plywood and logs, are exported. This has raised the income of this sector with respect to traditional products from other sectors. Uruguayan forestry plantations have rates of growth of 30 cubic metres per hectare per year and commercial harvesting occurs after nine years. ;Brazil Eucalypts were introduced to
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
in 1910, for timber substitution and the charcoal industry. It has thrived in the local environment, and today there are around 7 million hectares planted. The wood is highly valued by the charcoal and pulp and paper industries. The short rotation allows a larger wood production and supplies wood for several other activities, helping to preserve the native forests from logging. When well managed, the plantation soils can sustain endless replanting. ''Eucalyptus'' plantings are also used as wind breaks. Brazil's plantations have world-record rates of growth, typically over 40 cubic metres per hectare per year, and commercial harvesting occurs after years 5. Due to continual development and governmental funding, year-on-year growth is consistently being improved. ''Eucalyptus'' can produce up to 100 cubic metres per hectare per year. Brazil has become the top exporter and producer of ''Eucalyptus'' round wood and pulp, and has played an important role in developing the Australian market through the country's committed research in this area. The local iron producers in Brazil rely heavily on sustainably grown ''Eucalyptus'' for charcoal; this has greatly pushed up the price of charcoal in recent years. The plantations are generally owned and operated for national and international industry by timber asset companies such as Thomson Forestry, Greenwood Management or cellulose producers such as Aracruz Cellulose and
Stora Enso Stora Enso Oyj (from sv, Stora and fi, Enso ) is a manufacturer of pulp, paper and other forest products, headquartered in Helsinki, Finland. The majority of sales takes place in Europe, but there are also significant operations in Asia and S ...
. Overall, South America was expected to produce 55% of the world's ''Eucalyptus'' round-wood by 2010. Many environmental NGOs have criticised the use of exotic tree species for forestry in Latin America.


Africa

;Ethiopia Eucalypts were introduced to Ethiopia in either 1894 or 1895, either by Emperor Menelik II's French advisor Mondon-Vidailhet or by the Englishman Captain O'Brian. Menelik II endorsed its planting around his new capital city of
Addis Ababa Addis Ababa (; am, አዲስ አበባ, , new flower ; also known as , lit. "natural spring" in Oromo), is the capital and largest city of Ethiopia. It is also served as major administrative center of the Oromia Region. In the 2007 census, t ...
because of the massive deforestation around the city for firewood. According to Richard R.K. Pankhurst, "The great advantage of the eucalypts was that they were fast growing, required little attention and when cut down grew up again from the roots; it could be harvested every ten years. The tree proved successful from the onset". Plantations of eucalypts spread from the capital to other growing urban centres such as Debre Marqos. Pankhurst reports that the most common species found in Addis Ababa in the mid-1960s was '' E. globulus'', although he also found '' E. melliodora'' and '' E. rostrata'' in significant numbers. David Buxton, writing of central Ethiopia in the mid-1940s, observed that eucalyptus trees "have become an integral -- and a pleasing -- element in the Shoan landscape and has largely displaced the slow-growing native 'cedar' (''
Juniperus procera ''Juniperus procera'' (known by the common English names African juniper, African pencil-cedar, East African juniper, East African-cedar, and Kenya-cedar) is a coniferous tree native to mountainous areas in Africa and the Arabian Peninsula. It is ...
'')." It was commonly believed that the thirst of the ''Eucalyptus'' "tended to dry up rivers and wells", creating such opposition to the species that in 1913 a proclamation was issued ordering a partial destruction of all standing trees, and their replacement with mulberry trees. Pankhurst reports, "The proclamation however remained a dead letter; there is no evidence of eucalypts being uprooted, still less of mulberry trees being planted." Eucalypts remain a defining feature of Addis Ababa. ;Madagascar Much of Madagascar's original native forest has been replaced with ''Eucalyptus'', threatening biodiversity by isolating remaining natural areas such as
Andasibe-Mantadia National Park Andasibe-Mantadia National Park is a 155 square kilometre protected area, located about 150 km east of Antananarivo, consisting principally of primary growth forest in Alaotra-Mangoro Region in eastern Madagascar. The park's elevation range ...
. ;South Africa Numerous ''Eucalyptus'' species have been introduced into
South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. It is bounded to the south by of coastline that stretch along the Atlantic Ocean, South Atlantic and Indian Oceans; to the ...
, mainly for timber and firewood but also for ornamental purposes. They are popular with beekeepers for the honey they provide.Palgrave, K. C. 2002: ''Trees of Southern Africa''. Struik Publishers, Cape Town. However, in South Africa they are considered invasive, with their water-sucking capabilities threatening water supplies. They also release a chemical into the surrounding soil which kills native competitors. ''Eucalyptus'' seedlings are usually unable to compete with the indigenous grasses, but after a fire when the grass cover has been removed, a seed-bed may be created. The following ''Eucalyptus'' species have been able to become naturalised in South Africa: '' E. camaldulensis'', '' E. cladocalyx'', '' E. diversicolor'', '' E. grandis'' and '' E. lehmannii''. ;Zimbabwe As in South Africa, many ''Eucalyptus'' species have been introduced into Zimbabwe, mainly for timber and firewood, and '' E. robusta'' and '' E. tereticornis'' have been recorded as having become naturalised there.


Europe


Portugal

Eucalypts have been grown in Portugal since the mid 19th century, the first thought to be a specimen of '' E. obliqua'' introduced to
Vila Nova de Gaia Vila Nova de Gaia (; cel-x-proto, Cale), or simply Gaia, is a city and a municipality in Porto District in Norte Region, Portugal. It is located south of the city of Porto on the other side of the Douro River. The city proper had a population ...
in 1829. First as an ornamental but soon after in plantations, these eucalypts are prized due to their long and upright trunks, rapid growth and the ability to regrow after cutting. These plantations now occupy around 800,000 hectares, 10% of the country's total land area, 90% of the trees being '' E. globulus''. As of the late 20th century, there were an estimated 120 species of ''Eucalyptus'' in Portugal. The genus has also been subject to various controversies. Despite representing a large part of the agricultural economy, eucalypt plantations have a negative impact on soil destruction, inducing resistance to water infiltration and increasing the risks of erosion and soil loss, they are highly inflamable, aggravating the risk for wildfires. Various Portuguese laws on eucalypt plantations have been formed and reformed to better suit both sides. There are various ''Eucalyptus'' species of public interest in Portugal, namely a Karri in Coimbra's Mata Nacional de Vale de Canas, considered to be Europe's tallest tree at high.


Italy

In
Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical ...
, the ''Eucalyptus'' only arrived at the turn of the 19th century and large scale plantations were started at the beginning of the 20th century with the aim of drying up swampy ground to defeat malaria. During the 1930s, Benito Mussolini had thousands of ''Eucalyptus'' planted in the marshes around Rome. This, their rapid growth in the Italian climate and excellent function as windbreaks, has made them a common sight in the south of the country, including the islands of Sardinia and
Sicily (man) it, Siciliana (woman) , population_note = , population_blank1_title = , population_blank1 = , demographics_type1 = Ethnicity , demographics1_footnotes = , demographi ...
. They are also valued for the characteristic smelling and tasting honey that is produced from them. The variety of ''Eucalyptus'' most commonly found in Italy is '' E. camaldulensis''.


Greece

In
Greece Greece,, or , romanized: ', officially the Hellenic Republic, is a country in Southeast Europe. It is situated on the southern tip of the Balkans, and is located at the crossroads of Europe, Asia, and Africa. Greece shares land borders ...
, eucalypts are widely found, especially in southern Greece and Crete. They are cultivated and used for various purposes, including as an ingredient in pharmaceutical products (e.g., creams, elixirs and sprays) and for leather production. They were imported in 1862 by botanist Theodoros Georgios Orphanides. The principal species is ''Eucalyptus globulus''.


Ireland

''Eucalyptus'' has been grown in Ireland since trials in the 1930s and now grows wild in South Western Ireland in the mild climate.


Asia

''Eucalyptus'' seeds of the species ''E. globulus'' were imported into Palestine in the 1860s, but did not acclimatise well. Later, ''E. camaldulensis'' was introduced more successfully and it is still a very common tree in
Israel Israel (; he, יִשְׂרָאֵל, ; ar, إِسْرَائِيل, ), officially the State of Israel ( he, מְדִינַת יִשְׂרָאֵל, label=none, translit=Medīnat Yīsrāʾēl; ), is a country in Western Asia. It is situated ...
. The use of ''Eucalyptus'' trees to drain swampy land was a common practice in the late nineteenth and early twentieth centuries. The German Templer colony of Sarona had begun planting ''Eucalyptus'' for this purpose by 1874, though it is not known where the seeds came from. Many Zionist colonies also adopted the practice in the following years under the guidance of the Mikveh Israel Agricultural School. ''Eucalyptus'' trees are now considered an invasive species in the region. In India, the Institute of Forest Genetics and Tree Breeding, Coimbatore started a ''Eucalyptus'' breeding program in the 1990s. The organisation released four varieties of conventionally bred, high yielding and genetically improved clones for commercial and research interests in 2010. ''Eucalyptus'' trees were introduced to Sri Lanka in the late 19th century by
tea Tea is an aromatic beverage prepared by pouring hot or boiling water over cured or fresh leaves of ''Camellia sinensis'', an evergreen shrub native to East Asia which probably originated in the borderlands of southwestern China and north ...
and
coffee Coffee is a drink prepared from roasted coffee beans. Darkly colored, bitter, and slightly acidic, coffee has a stimulating effect on humans, primarily due to its caffeine content. It is the most popular hot drink in the world. Seeds of ...
planters, for wind protection, shade and fuel. Forestry replanting of ''Eucalyptus'' began in the 1930s in deforested mountain areas, and currently there are about 10 species present in the island. They account for 20% of major reforestation plantings. They provide
railway sleeper A railroad tie, crosstie (American English), railway tie (Canadian English) or railway sleeper ( Australian and British English) is a rectangular support for the rails in railroad tracks. Generally laid perpendicular to the rails, ties trans ...
s, utility poles, sawn timber and
fuelwood Firewood is any wooden material that is gathered and used for fuel. Generally, firewood is not highly processed and is in some sort of recognizable log or branch form, compared to other forms of wood fuel like pellets or chips. Firewood can ...
, but are controversial because of their adverse effect on biodiversity, hydrology and soil fertility. They are associated with another invasive species, the eucalyptus gall wasp, '' Leptocybe invasa''.


Pacific Islands

In Hawaii, some 90 species of ''Eucalyptus'' have been introduced to the islands, where they have displaced some native species due to their higher maximum height, fast growth and lower water needs. Particularly noticeable is the rainbow eucalyptus (''
Eucalyptus deglupta ''Eucalyptus deglupta'' is a species of tall tree, commonly known as the rainbow eucalyptus, Mindanao gum, or rainbow gum that is native to the Philippines, Indonesia, and Papua New Guinea. It is the only ''Eucalyptus'' species that usually liv ...
''), native to Indonesia and the Philippines, whose bark falls off to reveal a trunk that can be green, red, orange, yellow, pink and purple.


Non-native ''Eucalyptus'' and biodiversity

Due to similar favourable climatic conditions, ''Eucalyptus'' plantations have often replaced oak woodlands, for example in California, Spain and Portugal. The resulting monocultures have raised concerns about loss of biological diversity, through loss of acorns that mammals and birds feed on, absence of hollows that in oak trees provide shelter and nesting sites for birds and small mammals and for bee colonies, as well as lack of downed trees in managed plantations. A study of the relationship between birds and ''Eucalyptus'' in the San Francisco Bay Area found that bird diversity was similar in native forest versus ''Eucalyptus'' forest, but the species were different. One way in which the avifauna (local assortment of bird species) changes is that cavity-nesting birds including woodpeckers, owls, chickadees, wood ducks, etc. are
depauperate A depauperate ecosystem is an ecosystem which is lacking in numbers or variety of species, often because it lacks enough stored chemical elements and resources required for life. Thus, depauperate ecosystems often cannot support rapid growth of f ...
in ''Eucalyptus'' groves because the decay-resistant wood of these trees prevents cavity formation by decay or excavation. Also, those bird species that glean insects from foliage, such as warblers and vireos, experience population declines when ''Eucalyptus'' groves replace oak forest. Birds that thrive in ''Eucalyptus'' groves in California tend to prefer tall vertical habitat. These avian species include herons and egrets, which also nest in redwoods. The Point Reyes Bird Observatory observes that sometimes short-billed birds like the ruby-crowned kinglet are found dead beneath ''Eucalyptus'' trees with their nostrils clogged with pitch. Monarch butterflies use ''Eucalyptus'' in California for over-wintering, but in some locations have a preference for
Monterey pine ''Pinus radiata'' ( syn. ''Pinus insignis''), the Monterey pine, insignis pine or radiata pine, is a species of pine native to the Central Coast of California and Mexico ( Guadalupe Island and Cedros island). It is an evergreen conifer in the f ...
s.


Photo album

File:Eucalyptus macarthurii UC.jpg, Mature '' Eucalyptus macarthurii'', at the University of Canterbury File:E.sideroxylon, branchlets, stems, leaves, capsules & buds.jpg, '' Eucalyptus sideroxylon'', showing fruit (capsules) and buds with operculum present. File:Bach dan.jpg, ''Eucalyptus'' forest in
East Gippsland East Gippsland is the eastern region of Gippsland, Victoria, Australia covering 31,740 square kilometres (14%) of Victoria. It has a population of 80,114. Australian Bureau of Statistics2006 Census Community Profile Series: East Gippsland (St ...
,
Victoria Victoria most commonly refers to: * Victoria (Australia), a state of the Commonwealth of Australia * Victoria, British Columbia, provincial capital of British Columbia, Canada * Victoria (mythology), Roman goddess of Victory * Victoria, Seychelle ...
. Mostly ''E. albens'' (white box). File:Applebox.JPG, '' E. bridgesiana'' (apple box) on
Red Hill, Australian Capital Territory Red Hill (postcode: 2603) is a suburb of Canberra, Australian Capital Territory, Australia. The suburb is named after the northernmost hill of the ridge to the west of the suburb. The ridge is a reserve and managed as part of the Canberra Nature ...
. File:euc.uk.600pix.jpg, '' E. gunnii'' planted in southern
England England is a country that is part of the United Kingdom. It shares land borders with Wales to its west and Scotland to its north. The Irish Sea lies northwest and the Celtic Sea to the southwest. It is separated from continental Europe b ...
. The lower part of the trunk is covered in ivy. File:Eucalyptus cinera x pulverulenta.jpg, '' Eucalyptus cinerea'' x '' pulverulenta'' - National Botanical Gardens Canberra File:Eucalyptus gall.jpg, ''Eucalyptus'' gall File:CPonte Eucalyptus.jpg, '' E. grandis''. Province of Buenos Aires, Argentina. File:Eucalipto Galicia.JPG, ''Eucalyptus'' plantation near
Viveiro Viveiro () is a town and municipality in the province of Lugo, in the northwestern Galician autonomous community of Spain. It borders on the Cantabric Sea, to the west of Xove and to the east of O Vicedo. It has a residential population of o ...
, in Galicia in Northwest
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = ''Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , i ...
. Mostly ''E. globulus'' File:Snow Gum1.JPG, A
snow gum ''Eucalyptus pauciflora'', commonly known as snow gum, cabbage gum or white sally, is a species of tree or mallee that is native to eastern Australia. It has smooth bark, lance-shaped to elliptical leaves, flower buds in clusters of between sev ...
(''E. pauciflora''), in winter in the
Australian Alps The Australian Alps is a mountain range in southeast Australia. It comprises an interim Australian bioregion,0042-5184 However, the moth has also been a biovector of arsenic, transporting it from lowland feeding sites over long distances int ...
File:Eucalyptus rubida.jpg, '' E. rubida'' ( candlebark gum) in
Burra, New South Wales Burra is an Australian locality of rural smallholdings lying 20 kilometres to the south of Queanbeyan, New South Wales in the Queanbeyan-Palerang Region. At the , Burra had a population of 790 people. History The Ngarigo people originally inha ...
. File:Sydney Blue Gums Kippara Forest via Wauchope.JPG, Sydney blue gums west of Port Macquarie, New South Wales File:Eucalyptus Chapmaniana 5198.JPG, '' Eucalyptus chapmaniana'' (bogong gum) in Kew Gardens,
London London is the capital and List of urban areas in the United Kingdom, largest city of England and the United Kingdom, with a population of just under 9 million. It stands on the River Thames in south-east England at the head of a estuary dow ...
File:Sherbrooke forest Victoria 220rs.jpg, '' E. regnans'' trees in Sherbrooke Forest,
Victoria Victoria most commonly refers to: * Victoria (Australia), a state of the Commonwealth of Australia * Victoria, British Columbia, provincial capital of British Columbia, Canada * Victoria (mythology), Roman goddess of Victory * Victoria, Seychelle ...
File:Dean-Nicolle-Deanei.JPG, '' E. deanei'', Blue Mountains National Park, Australia File:Prospectcreek.jpg, Eucalypt woodland area near Prospect Creek in western Sydney. Mostly '' E. amplifolia'' and '' E. tereticornis''.


References


General source

* * Blakely, W.F., ''A Key to the Eucalypts: with descriptions of 522 species and 150 varieties''. Third Edition, 1965, Forest and Timber Bureau, Canberra. * 5th edition. * Third edition. vol. 1. ''South-eastern Australia''. * Kelly, Stan, text by G. M. Chippendale and R. D. Johnston, ''Eucalypts: Volume I''. Nelson, Melbourne 1969, 1982, etc. * * *


External links


Lucid Online Player - EUCLID Eucalypts of Australia
( Multi-access key) to 917 species/subspecies taxonomy as of December 2009.
ANPSA Plant Guide: ''Eucalyptus'', ''Corymbia'' and ''Angophora''

Currency Creek Arboretum - Eucalypt Research


b

(2005-current)

* ttp://www.hort.purdue.edu/newcrop/duke_energy/Eucalyptus.html Handbook of Energy CropsDuke, James A. 1983.
''The Eucalyptus of California: Seeds of Good or Seeds of Evil?''
Santos, Robert. 1997 Denair, CA : Alley-Cass Publications
"The Rise and Fall of the Gum Tree: How California Came to Love—then Disown—Eucalyptus"
Farmer, Jared. 2014.
EUCALYPTOLOGICS: Information Resources on ''Eucalyptus'' cultivation around the World
Iglesias Trabado, Gustavo (2007-current) {{Authority control * Taxa described in 1789 Taxa named by Charles Louis L'Héritier de Brutelle Australasian realm flora