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Ernst Heinrich Philipp August Haeckel (; 16 February 1834 – 9 August 1919) was a German zoologist, naturalist,
eugenicist Eugenics ( ; ) is a fringe set of beliefs and practices that aim to improve the genetic quality of a human population. Historically, eugenicists have attempted to alter human gene pools by excluding people and groups judged to be inferior or ...
, philosopher, physician, professor, marine biologist and artist. He discovered, described and named thousands of new
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriat ...
, mapped a
genealogical Genealogy () is the study of families, family history, and the tracing of their lineages. Genealogists use oral interviews, historical records, genetic analysis, and other records to obtain information about a family and to demonstrate kin ...
tree relating all life forms and coined many terms in
biology Biology is the scientific study of life. It is a natural science with a broad scope but has several unifying themes that tie it together as a single, coherent field. For instance, all organisms are made up of cells that process hereditary ...
, including ''
ecology Ecology () is the study of the relationships between living organisms, including humans, and their physical environment. Ecology considers organisms at the individual, population, community, ecosystem, and biosphere level. Ecology overl ...
'', '' phylum'', ''
phylogeny A phylogenetic tree (also phylogeny or evolutionary tree Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, MA.) is a branching diagram or a tree showing the evolutionary relationships among various biological s ...
'', and ''
Protista A protist () is any eukaryotic organism (that is, an organism whose cells contain a cell nucleus) that is not an animal, plant, or fungus. While it is likely that protists share a common ancestor (the last eukaryotic common ancestor), the e ...
.'' Haeckel promoted and popularised
Charles Darwin Charles Robert Darwin ( ; 12 February 1809 – 19 April 1882) was an English naturalist, geologist, and biologist, widely known for his contributions to evolutionary biology. His proposition that all species of life have descended ...
's work in Germany and developed the influential but no longer widely held recapitulation theory ("ontogeny recapitulates phylogeny") claiming that an individual organism's biological development, or
ontogeny Ontogeny (also ontogenesis) is the origination and development of an organism (both physical and psychological, e.g., moral development), usually from the time of fertilization of the egg to adult. The term can also be used to refer to the s ...
, parallels and summarises its species' evolutionary development, or
phylogeny A phylogenetic tree (also phylogeny or evolutionary tree Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, MA.) is a branching diagram or a tree showing the evolutionary relationships among various biological s ...
. The published artwork of Haeckel includes over 100 detailed, multi-colour illustrations of animals and sea creatures, collected in his ''
Kunstformen der Natur (known in English as ''Art Forms in Nature'') is a book of lithographic and halftone prints by German biologist Ernst Haeckel. ...
'' ("Art Forms of Nature"), a book which would go on to influence the Art Nouveau artistic movement. As a philosopher, Ernst Haeckel wrote ''Die Welträthsel'' (1895–1899; in English: ''The Riddle of the Universe'', 1901), the genesis for the term "
world riddle The term "world riddle" or "world-riddle" has been associated, for over 100 years, with Friedrich Nietzsche (who mentioned ''Welträthsel'' in several of his writings) and with the biologist- philosopher Ernst Haeckel, who, as a professor of ...
" (''Welträtsel''); and ''Freedom in Science and Teaching'' to support teaching evolution. Haeckel was also a promoter of scientific racism and embraced the idea of Social Darwinism.


Life

Ernst Haeckel was born on 16 February 1834, in Potsdam (then part of the
Kingdom of Prussia The Kingdom of Prussia (german: Königreich Preußen, ) was a German kingdom that constituted the state of Prussia between 1701 and 1918.Marriott, J. A. R., and Charles Grant Robertson. ''The Evolution of Prussia, the Making of an Empire''. ...
). In 1852 Haeckel completed studies at the ''Domgymnasium'', the cathedral high-school of
Merseburg Merseburg () is a town in central Germany in southern Saxony-Anhalt, situated on the river Saale, and approximately 14 km south of Halle (Saale) and 30 km west of Leipzig. It is the capital of the Saalekreis district. It had a dioces ...
."Ernst Haeckel" (article),''German Wikipedia'', 26 October 2006, webpage: DE-Wiki-Ernst-Haeckel: last paragraph of "Leben" (Life) section. He then studied medicine in
Berlin Berlin ( , ) is the capital and largest city of Germany by both area and population. Its 3.7 million inhabitants make it the European Union's most populous city, according to population within city limits. One of Germany's sixteen constitu ...
and Würzburg, particularly with
Albert von Kölliker Albert von Kölliker (born Rudolf Albert Kölliker'';'' 6 July 18172 November 1905) was a Swiss anatomist, physiologist, and histologist. Biography Albert Kölliker was born in Zurich, Switzerland. His early education was carried on in Zurich, ...
,
Franz Leydig Franz von Leydig, also Franz Leydig (; 21 May 1821 – 13 April 1908), was a German zoologist and comparative anatomist."Franz von Leydig" (biography), Ole Daniel Enerson, 2006, WhoNamedIt.coWNI-675-Leydig Life Franz Leydig was born on 21 May ...
, Rudolf Virchow (with whom he later worked briefly as assistant), and with the anatomist-physiologist Johannes Peter Müller (1801–1858). Together with Hermann Steudner he attended
botany Botany, also called , plant biology or phytology, is the science of plant life and a branch of biology. A botanist, plant scientist or phytologist is a scientist who specialises in this field. The term "botany" comes from the Ancient Greek w ...
lectures in Würzburg. In 1857 Haeckel attained a doctorate in medicine, and afterwards he received the license to practice medicine. The occupation of physician appeared less worthwhile to Haeckel after contact with suffering patients. He studied under Karl Gegenbaur at the
University of Jena The University of Jena, officially the Friedrich Schiller University Jena (german: Friedrich-Schiller-Universität Jena, abbreviated FSU, shortened form ''Uni Jena''), is a public research university located in Jena, Thuringia, Germany. The ...
for three years, earning a habilitation in comparative anatomy in 1861, before becoming a professor of
zoology Zoology ()The pronunciation of zoology as is usually regarded as nonstandard, though it is not uncommon. is the branch of biology that studies the animal kingdom, including the structure, embryology, evolution, classification, habits, an ...
at the University at Jena, where he remained for 47 years, from 1862 to 1909. Between 1859 and 1866 Haeckel worked on many phyla, such as radiolarians, poriferans (
sponge Sponges, the members of the phylum Porifera (; meaning 'pore bearer'), are a basal animal clade as a sister of the diploblasts. They are multicellular organisms that have bodies full of pores and channels allowing water to circulate throu ...
s) and annelids (segmented worms). During a trip to the
Mediterranean The Mediterranean Sea is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean Basin and almost completely enclosed by land: on the north by Western and Southern Europe and Anatolia, on the south by North Africa, and on ...
, Haeckel named nearly 150 new species of radiolarians. In 1864, his beloved first wife, Anna Sethe, died. Haeckel dedicated some species of jellyfish of particular beauty (such as ''Desmonema annasethe'') to his unforgettable wife. From 1866 to 1867 Haeckel made an extended journey to the
Canary Islands The Canary Islands (; es, :es:Canarias, Canarias, ), also known informally as the Canaries, are a Spanish Autonomous communities of Spain, autonomous community and archipelago in the Atlantic Ocean, in Macaronesia. At their closest point to ...
with Hermann Fol. On 17 October 1866 he arrived in London. Over the next few days he met Charles Lyell, and visited
Thomas Huxley Thomas Henry Huxley (4 May 1825 – 29 June 1895) was an English biologist and anthropologist specialising in comparative anatomy. He has become known as "Darwin's Bulldog" for his advocacy of Charles Darwin's theory of evolution. The stori ...
and family at their home. On 21 October he visited
Charles Darwin Charles Robert Darwin ( ; 12 February 1809 – 19 April 1882) was an English naturalist, geologist, and biologist, widely known for his contributions to evolutionary biology. His proposition that all species of life have descended ...
at Down House in Kent. In 1867 he married Agnes Huschke. Their son Walter was born in 1868, their daughters Elizabeth in 1871 and Emma in 1873. In 1869 he traveled as a researcher to
Norway Norway, officially the Kingdom of Norway, is a Nordic countries, Nordic country in Northern Europe, the mainland territory of which comprises the western and northernmost portion of the Scandinavian Peninsula. The remote Arctic island of ...
, in 1871 to
Croatia , image_flag = Flag of Croatia.svg , image_coat = Coat of arms of Croatia.svg , anthem = " Lijepa naša domovino"("Our Beautiful Homeland") , image_map = , map_caption = , capi ...
(where he lived on the island of Hvar in a monastery), and in 1873 to
Egypt Egypt ( ar, مصر , ), officially the Arab Republic of Egypt, is a List of transcontinental countries, transcontinental country spanning the North Africa, northeast corner of Africa and Western Asia, southwest corner of Asia via a land bridg ...
,
Turkey Turkey ( tr, Türkiye ), officially the Republic of Türkiye ( tr, Türkiye Cumhuriyeti, links=no ), is a transcontinental country located mainly on the Anatolian Peninsula in Western Asia, with a small portion on the Balkan Peninsula ...
, and
Greece Greece,, or , romanized: ', officially the Hellenic Republic, is a country in Southeast Europe. It is situated on the southern tip of the Balkans, and is located at the crossroads of Europe, Asia, and Africa. Greece shares land borders wi ...
. In 1907 he had a museum built in
Jena Jena () is a German city and the second largest city in Thuringia. Together with the nearby cities of Erfurt and Weimar, it forms the central metropolitan area of Thuringia with approximately 500,000 inhabitants, while the city itself has a po ...
to teach the public about
evolution Evolution is change in the heritable characteristics of biological populations over successive generations. These characteristics are the expressions of genes, which are passed on from parent to offspring during reproduction. Variation ...
. Haeckel retired from teaching in 1909, and in 1910 he withdrew from the
Evangelical Church of Prussia The Prussian Union of Churches (known under multiple other names) was a major Protestant church body which emerged in 1817 from a series of decrees by Frederick William III of Prussia that united both Lutheran and Reformed denominations in Pru ...
. On the occasion of his 80th birthday celebration he was presented with a two-volume work entitled ''Was wir Ernst Haeckel verdanken (What We Owe to Ernst Haeckel)'', edited at the request of the German Monistenbund by Heinrich Schmidt of Jena. Haeckel's wife, Agnes, died in 1915, and he became substantially frailer, breaking his leg and arm. He sold his "Villa Medusa" in
Jena Jena () is a German city and the second largest city in Thuringia. Together with the nearby cities of Erfurt and Weimar, it forms the central metropolitan area of Thuringia with approximately 500,000 inhabitants, while the city itself has a po ...
in 1918 to the Carl Zeiss foundation, which preserved his library. Haeckel died on 9 August 1919. Haeckel became the most famous proponent of
Monism Monism attributes oneness or singleness (Greek: μόνος) to a concept e.g., existence. Various kinds of monism can be distinguished: * Priority monism states that all existing things go back to a source that is distinct from them; e.g., i ...
in Germany.


Politics

Haeckel's affinity for the German Romantic movement, coupled with his acceptance of a form of
Lamarckism Lamarckism, also known as Lamarckian inheritance or neo-Lamarckism, is the notion that an organism can pass on to its offspring physical characteristics that the parent organism acquired through use or disuse during its lifetime. It is also calle ...
, influenced his political beliefs. Rather than being a strict Darwinian, Haeckel believed that the characteristics of an organism were acquired through interactions with the environment and that
ontogeny Ontogeny (also ontogenesis) is the origination and development of an organism (both physical and psychological, e.g., moral development), usually from the time of fertilization of the egg to adult. The term can also be used to refer to the s ...
reflected
phylogeny A phylogenetic tree (also phylogeny or evolutionary tree Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, MA.) is a branching diagram or a tree showing the evolutionary relationships among various biological s ...
. He saw the
social science Social science is one of the branches of science, devoted to the study of societies and the relationships among individuals within those societies. The term was formerly used to refer to the field of sociology, the original "science of s ...
s as instances of "applied biology", and that phrase was picked up and used for Nazi propaganda."Ernst Haeckel" (biography), UC Berkeley, 2004, webpage
BerkeleyEdu-Haeckel
In 1906 Haeckel belonged to the founders of the Monist League ( Deutscher Monistenbund), which took a stance against philosophical materialism and promote a "natural Weltanschauung". This organization lasted until 1933 and included such notable members as Wilhelm Ostwald, Georg von Arco (1869–1940),
Helene Stöcker Helene Stöcker (13 November 1869 – 24 February 1943) was a German feminist, pacifist and gender activist. She successfully campaigned keep same sex relationships between women legal, but she was unsuccessful in her campaign to legalise abortion ...
and
Walter Arthur Berendsohn Walter Arthur Berendsohn (10 September 1884, in Hamburg – 30 January 1984, in Stockholm) was a German literary scholar. He was an active member of the Deutsche Liga fur Menschenrechte ( League for Men's Rights), a spinoff of the pacifist Bund ...
. He was the first person to use the term "first world war". However, Haeckel's books were banned by the Nazi Party, which refused Monism and Haeckel's freedom of thought. Moreover, it is worth mentioning that Haeckel had often overtly recognized the great contribution of educated Jews to the German culture.


Research

Haeckel was a zoologist, an accomplished artist and illustrator, and later a professor of comparative anatomy. Although Haeckel's ideas are important to the history of evolutionary theory, and although he was a competent
invertebrate Invertebrates are a paraphyletic group of animals that neither possess nor develop a vertebral column (commonly known as a ''backbone'' or ''spine''), derived from the notochord. This is a grouping including all animals apart from the chorda ...
anatomist most famous for his work on radiolaria, many speculative concepts that he championed are now considered incorrect. For example, Haeckel described and named hypothetical ancestral
microorganism A microorganism, or microbe,, ''mikros'', "small") and ''organism'' from the el, ὀργανισμός, ''organismós'', "organism"). It is usually written as a single word but is sometimes hyphenated (''micro-organism''), especially in old ...
s that have never been found. He was one of the first to consider
psychology Psychology is the science, scientific study of mind and behavior. Psychology includes the study of consciousness, conscious and Unconscious mind, unconscious phenomena, including feelings and thoughts. It is an academic discipline of immens ...
as a branch of
physiology Physiology (; ) is the scientific study of functions and mechanisms in a living system. As a sub-discipline of biology, physiology focuses on how organisms, organ systems, individual organs, cells, and biomolecules carry out the chemic ...
. He also proposed the kingdom ''
Protista A protist () is any eukaryotic organism (that is, an organism whose cells contain a cell nucleus) that is not an animal, plant, or fungus. While it is likely that protists share a common ancestor (the last eukaryotic common ancestor), the e ...
'' in 1866. His chief interests lay in
evolution Evolution is change in the heritable characteristics of biological populations over successive generations. These characteristics are the expressions of genes, which are passed on from parent to offspring during reproduction. Variation ...
and life development processes in general, including development of nonrandom form, which culminated in the beautifully illustrated ''
Kunstformen der Natur (known in English as ''Art Forms in Nature'') is a book of lithographic and halftone prints by German biologist Ernst Haeckel. ...
'' (''Art forms of nature''). Haeckel did not support
natural selection Natural selection is the differential survival and reproduction of individuals due to differences in phenotype. It is a key mechanism of evolution, the change in the heritable traits characteristic of a population over generations. Cha ...
, rather believing in
Lamarckism Lamarckism, also known as Lamarckian inheritance or neo-Lamarckism, is the notion that an organism can pass on to its offspring physical characteristics that the parent organism acquired through use or disuse during its lifetime. It is also calle ...
. Haeckel advanced a version of the earlier recapitulation theory previously set out by
Étienne Serres Antoine Étienne Renaud Augustin Serres (12 September 1786, Clairac – 22 January 1868, Paris) was a French physician and embryologist. He was among the first to formulate the recapitulation theory. Background Étienne Serres was the son ...
in the 1820s and supported by followers of
Étienne Geoffroy Saint-Hilaire Étienne Geoffroy Saint-Hilaire (15 April 177219 June 1844) was a French naturalist who established the principle of "unity of composition". He was a colleague of Jean-Baptiste Lamarck and expanded and defended Lamarck's evolutionary theories ...
including Robert Edmond Grant. It proposed a link between
ontogeny Ontogeny (also ontogenesis) is the origination and development of an organism (both physical and psychological, e.g., moral development), usually from the time of fertilization of the egg to adult. The term can also be used to refer to the s ...
(development of form) and
phylogeny A phylogenetic tree (also phylogeny or evolutionary tree Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, MA.) is a branching diagram or a tree showing the evolutionary relationships among various biological s ...
(evolutionary descent), summed up by Haeckel in the phrase "
ontogeny recapitulates phylogeny Ontogeny (also ontogenesis) is the origination and development of an organism (both physical and psychological, e.g., moral development), usually from the time of fertilization of the egg to adult. The term can also be used to refer to the stu ...
". His concept of recapitulation has been refuted in the form he gave it (now called "strong recapitulation"), in favour of the ideas first advanced by Karl Ernst von Baer. The strong recapitulation hypothesis views ontogeny as repeating forms of adult ancestors, while weak recapitulation means that what is repeated (and built upon) is the ancestral embryonic development process. Haeckel supported the theory with embryo drawings that have since been shown to be oversimplified and in part inaccurate, and the theory is now considered an oversimplification of quite complicated relationships, however comparison of embryos remains a powerful way to demonstrate that all animals are related. Haeckel introduced the concept of
heterochrony In evolutionary developmental biology, heterochrony is any genetically controlled difference in the timing, rate, or duration of a developmental process in an organism compared to its ancestors or other organisms. This leads to changes in the ...
, the change in timing of embryonic development over the course of evolution. Haeckel was a flamboyant figure, who sometimes took great, non-scientific leaps from available evidence. For example, at the time when Darwin published ''
On the Origin of Species by Means of Natural Selection On, on, or ON may refer to: Arts and entertainment Music * On (band), a solo project of Ken Andrews * ''On'' (EP), a 1993 EP by Aphex Twin * ''On'' (Echobelly album), 1995 * ''On'' (Gary Glitter album), 2001 * ''On'' (Imperial Teen album), 200 ...
'' (1859), Haeckel postulated that evidence of human evolution would be found in the Dutch East Indies (now
Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania between the Indian and Pacific oceans. It consists of over 17,000 islands, including Sumatra, Java, Sulawesi, and parts of Borneo and New Gui ...
). At that time, no remains of human ancestors had yet been identified. He described these theoretical remains in great detail and even named the as-yet unfound species, ''Pithecanthropus alalus'', and instructed his students such as Richard and Oskar Hertwig to go and find it. One student did find some remains: a Dutchman named Eugène Dubois searched the
East Indies The East Indies (or simply the Indies), is a term used in historical narratives of the Age of Discovery. The Indies refers to various lands in the East or the Eastern hemisphere, particularly the islands and mainlands found in and around ...
from 1887 to 1895, discovering the remains of
Java Man Java Man (''Homo erectus erectus'', formerly also ''Anthropopithecus erectus'', ''Pithecanthropus erectus'') is an early human fossil discovered in 1891 and 1892 on the island of Java (Dutch East Indies, now part of Indonesia). Estimated to be b ...
in 1891, consisting of a skullcap, thighbone, and a few teeth. These remains are among the oldest hominid remains ever found. Dubois classified Java Man with Haeckel's ''Pithecanthropus'' label, though they were later reclassified as '' Homo erectus''. Some scientists of the day suggested Dubois' Java Man as a potential intermediate form between modern humans and the common ancestor we share with the other great apes. The current consensus of anthropologists is that the direct ancestors of modern humans were African populations of '' Homo erectus'' (possibly '' Homo ergaster''), rather than the Asian populations exemplified by Java Man and Peking Man. (Ironically, a new human species, Homo floresiensis, a dwarf human type, has recently been discovered in the island of Flores).


Polygenism and racial theory

The creationist polygenism of
Samuel George Morton Samuel George Morton (January 26, 1799 – May 15, 1851) was an American physician, natural scientist, and writer who argued against the single creation story of the Bible, monogenism, instead supporting a theory of multiple racial creations, poly ...
and Louis Agassiz, which presented human races as separately created
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriat ...
, was rejected by
Charles Darwin Charles Robert Darwin ( ; 12 February 1809 – 19 April 1882) was an English naturalist, geologist, and biologist, widely known for his contributions to evolutionary biology. His proposition that all species of life have descended ...
, who argued for the monogenesis of the
human Humans (''Homo sapiens'') are the most abundant and widespread species of primate, characterized by bipedalism and exceptional cognitive skills due to a large and complex brain. This has enabled the development of advanced tools, cultu ...
species and the African origin of modern humans. In contrast to most of Darwin's supporters, Haeckel put forward a doctrine of evolutionary polygenism based on the ideas of the linguist August Schleicher, in which several different language groups had arisen separately from speechless prehuman ''Urmenschen'' (german: proto-humans), which themselves had evolved from simian ancestors. These separate languages had completed the transition from animals to man, and under the influence of each main branch of languages, humans had evolved – in a kind of Lamarckian use-inheritance – as separate species, which could be subdivided into races. From this, Haeckel drew the implication that languages with the most potential yield the human races with the most potential, led by the Semitic and Indo-Germanic groups, with Berber, Jewish, Greco-Roman and Germanic varieties to the fore. As Haeckel stated: ) had split into several species or kinds. With each of these human species, language developed on its own and independently of the others. At least this is the view of Schleicher, one of the foremost authorities on this subject. ... If one views the origin of the branches of language as the special and principal act of becoming human, and the species of humankind as distinguished according to their language stem, then one can say that the different species of men arose independently of one another. Haeckel's view can be seen as a forerunner of the views of Carleton Coon, who also believed that human races evolved independently and in parallel with each other. These ideas eventually fell from favour. Haeckel also applied the hypothesis of polygenism to the modern diversity of human groups. He became a key figure in social darwinism and leading proponent of scientific racism, stating for instance: Haeckel divided human beings into ten races, of which the Caucasian was the highest and the primitives were doomed to extinction. In his view, 'Negroes' were savages and Whites were the most civilised: for instance, he claimed that ' e Negro' had stronger and more freely movable toes than any other race, which, he argued, was evidence of their being less evolved, and which led him to compare them to four-handed" Apes'.
Gustav Jahoda Gustav Jahoda, FBA, FRSE (11 October 1920 – 12 December 2016) was an Austrian psychologist and writer. He was educated in Vienna, then subsequently in Paris and London. He studied sociology and psychology at London University before obtaining a ...
, ''Images of Savages: Ancient Roots of Modern Prejudice in Western Culture'', 1999, p. 83
In his '' Ontogeny and Phylogeny'' Harvard paleontologist
Stephen Jay Gould Stephen Jay Gould (; September 10, 1941 – May 20, 2002) was an American paleontologist, evolutionary biologist, and historian of science. He was one of the most influential and widely read authors of popular science of his generation. Goul ...
wrote: " aeckel'sevolutionary racism; his call to the German people for racial purity and unflinching devotion to a 'just' state; his belief that harsh, inexorable laws of evolution ruled human civilization and nature alike, conferring upon favored races the right to dominate others ... all contributed to the rise of Nazism." In his introduction to the
Nazi party The Nazi Party, officially the National Socialist German Workers' Party (german: Nationalsozialistische Deutsche Arbeiterpartei or NSDAP), was a far-right political party in Germany active between 1920 and 1945 that created and supported t ...
ideologue Alfred Rosenberg's 1930 book, '' he Myth of the Twentieth Century', Peter Peel affirms that Rosenberg had indeed read Haeckel. In the same line of thought, historian Daniel Gasman states that Haeckel's ideology stimulated the birth of Fascist ideology in Italy and France. However,
Robert J. Richards Robert J. Richards (born 1942) is an author and the Morris Fishbein Distinguished Service Professor of the History of Science and Medicine at the University of Chicago. He has written or edited seven books about the history of science as well as ...
notes: "Haeckel, on his travels to Ceylon and Indonesia, often formed closer and more intimate relations with natives, even members of the untouchable classes, than with the European colonials." and says the Nazis rejected Haeckel, since he opposed antisemitism, while supporting ideas they disliked (for instance atheism, feminism, internationalism, pacifism etc.).


Asia hypothesis

Haeckel claimed the origin of humanity was to be found in Asia: he believed that
Hindustan ''Hindūstān'' ( , from '' Hindū'' and ''-stān''), also sometimes spelt as Hindōstān ( ''Indo-land''), along with its shortened form ''Hind'' (), is the Persian-language name for the Indian subcontinent that later became commonly used b ...
(Indian subcontinent) was the actual location where the first humans had evolved. Haeckel argued that humans were closely related to the
primate Primates are a diverse order of mammals. They are divided into the strepsirrhines, which include the lemurs, galagos, and lorisids, and the haplorhines, which include the tarsiers and the simians ( monkeys and apes, the latter includin ...
s of Southeast Asia and rejected Darwin's hypothesis of Africa. Haeckel later claimed that the missing link was to be found on the lost continent of
Lemuria Lemuria (), or Limuria, was a continent proposed in 1864 by zoologist Philip Sclater, theorized to have sunk beneath the Indian Ocean, later appropriated by occultists in supposed accounts of human origins. The theory was discredited with the d ...
located in the Indian Ocean. He believed that Lemuria was the home of the first humans and that Asia was the home of many of the earliest
primate Primates are a diverse order of mammals. They are divided into the strepsirrhines, which include the lemurs, galagos, and lorisids, and the haplorhines, which include the tarsiers and the simians ( monkeys and apes, the latter includin ...
s; he thus supported that Asia was the cradle of hominid evolution. Haeckel also claimed that Lemuria connected Asia and Africa, which allowed the migration of humans to the rest of the world. In Haeckel's book ''The History of Creation'' (1884) he included migration routes which he thought the first humans had used outside of Lemuria.


Embryology and recapitulation theory

When Haeckel was a student in the 1850s he showed great interest in
embryology Embryology (from Greek ἔμβρυον, ''embryon'', "the unborn, embryo"; and -λογία, '' -logia'') is the branch of animal biology that studies the prenatal development of gametes (sex cells), fertilization, and development of embr ...
, attending the rather unpopular lectures twice and in his notes sketched the visual aids: textbooks had few illustrations, and large format plates were used to show students how to see the tiny forms under a reflecting microscope, with the translucent tissues seen against a black background. Developmental series were used to show stages within a species, but inconsistent views and stages made it even more difficult to compare different species. It was agreed by all European evolutionists that all vertebrates looked very similar at an early stage, in what was thought of as a common ideal type, but there was a continuing debate from the 1820s between the Romantic recapitulation theory that human embryos developed through stages of the forms of all the major groups of adult animals, literally manifesting a sequence of organisms on a linear chain of being, and Karl Ernst von Baer's opposing view, stated in von Baer's laws of embryology, that the early general forms diverged into four major groups of specialised forms without ever resembling the adult of another species, showing affinity to an
archetype The concept of an archetype (; ) appears in areas relating to behavior, historical psychology, and literary analysis. An archetype can be any of the following: # a statement, pattern of behavior, prototype, "first" form, or a main model that ...
but no relation to other types or any
transmutation of species Transmutation of species and transformism are unproven 18th and 19th-century evolutionary ideas about the change of one species into another that preceded Charles Darwin's theory of natural selection. The French ''Transformisme'' was a term used ...
. By the time Haeckel was teaching he was able to use a textbook with woodcut illustrations written by his own teacher
Albert von Kölliker Albert von Kölliker (born Rudolf Albert Kölliker'';'' 6 July 18172 November 1905) was a Swiss anatomist, physiologist, and histologist. Biography Albert Kölliker was born in Zurich, Switzerland. His early education was carried on in Zurich, ...
, which purported to explain human development while also using other mammalian embryos to claim a coherent sequence. Despite the significance to ideas of transformism, this was not really polite enough for the new popular science writing, and was a matter for medical institutions and for experts who could make their own comparisons.


Darwin, Naturphilosophie and Lamarck

Darwin's ''
On the Origin of Species ''On the Origin of Species'' (or, more completely, ''On the Origin of Species by Means of Natural Selection, or the Preservation of Favoured Races in the Struggle for Life''),The book's full original title was ''On the Origin of Species by Me ...
'', which made a powerful impression on Haeckel when he read it in 1864, was very cautious about the possibility of ever reconstructing the history of life, but did include a section reinterpreting von Baer's embryology and revolutionising the field of study, concluding that "Embryology rises greatly in interest, when we thus look at the embryo as a picture, more or less obscured, of the common parent-form of each great class of animals." It mentioned von Baer's 1828 anecdote (misattributing it to Louis Agassiz) that at an early stage embryos were so similar that it could be impossible to tell whether an unlabelled specimen was of a mammal, a bird, or of a reptile, and Darwin's own research using embryonic stages of barnacles to show that they are
crustacean Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapoda, decapods, ostracoda, seed shrimp, branchiopoda, branchiopods, argulidae, fish lice, krill, remipedes, isopoda, isopods, barnacles, copepods, ...
s, while cautioning against the idea that one organism or embryonic stage is "higher" or "lower", or more or less evolved. Haeckel disregarded such caution, and in a year wrote his massive and ambitious ''Generelle Morphologie'', published in 1866, presenting a revolutionary new synthesis of Darwin's ideas with the German tradition of '' Naturphilosophie'' going back to
Goethe Johann Wolfgang von Goethe (28 August 1749 – 22 March 1832) was a German poet, playwright, novelist, scientist, statesman, theatre director, and critic. His works include plays, poetry, literature, and aesthetic criticism, as well as tr ...
and with the progressive evolutionism of Lamarck in what he called ''Darwinismus''. He used morphology to reconstruct the evolutionary history of life, in the absence of fossil evidence using embryology as evidence of ancestral relationships. He invented new terms, including
ontogeny Ontogeny (also ontogenesis) is the origination and development of an organism (both physical and psychological, e.g., moral development), usually from the time of fertilization of the egg to adult. The term can also be used to refer to the s ...
and
phylogeny A phylogenetic tree (also phylogeny or evolutionary tree Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, MA.) is a branching diagram or a tree showing the evolutionary relationships among various biological s ...
, to present his evolutionised recapitulation theory that "ontogeny recapitulated phylogeny". The two massive volumes sold poorly, and were heavy going: with his limited understanding of German, Darwin found them impossible to read. Haeckel's publisher turned down a proposal for a "strictly scholarly and objective" second edition.


Embryological drawings

Haeckel's aim was a reformed morphology with evolution as the organising principle of a cosmic synthesis unifying science, religion, and art. He was giving successful "popular lectures" on his ideas to students and townspeople in
Jena Jena () is a German city and the second largest city in Thuringia. Together with the nearby cities of Erfurt and Weimar, it forms the central metropolitan area of Thuringia with approximately 500,000 inhabitants, while the city itself has a po ...
, in an approach pioneered by his teacher Rudolf Virchow. To meet his publisher's need for a popular work he used a student's transcript of his lectures as the basis of his ''Natürliche Schöpfungsgeschichte'' of 1868, presenting a comprehensive presentation of evolution. In the Spring of that year he drew figures for the book, synthesising his views of specimens in Jena and published pictures to represent types. After publication he told a colleague that the images "are completely exact, partly copied from nature, partly assembled from all illustrations of these early stages that have hitherto become known". There were various styles of embryological drawings at that time, ranging from more schematic representations to "naturalistic" illustrations of specific specimens. Haeckel believed privately that his figures were both exact and synthetic, and in public asserted that they were schematic like most figures used in teaching. The images were reworked to match in size and orientation, and though displaying Haeckel's own views of essential features, they support von Baer's concept that vertebrate embryos begin similarly and then diverge. Relating different images on a grid conveyed a powerful evolutionary message. As a book for the general public, it followed the common practice of not citing sources. The book sold very well, and while some anatomical experts hostile to Haeckel's evolutionary views expressed some private concerns that certain figures had been drawn rather freely, the figures showed what they already knew about similarities in embryos. The first published concerns came from Ludwig Rütimeyer, a professor of zoology and comparative anatomy at the
University of Basel The University of Basel (Latin: ''Universitas Basiliensis'', German: ''Universität Basel'') is a university in Basel, Switzerland. Founded on 4 April 1460, it is Switzerland's oldest university and among the world's oldest surviving universiti ...
who had placed fossil mammals in an evolutionary lineage early in the 1860s and had been sent a complimentary copy. At the end of 1868 his review in the ''Archiv für Anthropologie'' wondered about the claim that the work was "popular and scholarly", doubting whether the second was true, and expressed horror about such public discussion of man's place in nature with illustrations such as the evolutionary trees being shown to non-experts. Though he made no suggestion that embryo illustrations should be directly based on specimens, to him the subject demanded the utmost "scrupulosity and conscientiousness" and an artist must "not arbitrarily model or generalise his originals for speculative purposes" which he considered proved by comparison with works by other authors. In particular, "one and the same, moreover incorrectly interpreted woodcut, is presented to the reader three times in a row and with three different captions as heembryo of the dog, the chick, ndthe turtle". He accused Haeckel of "playing fast and loose with the public and with science", and failing to live up to the obligation to the truth of every serious researcher. Haeckel responded with angry accusations of bowing to religious prejudice, but in the second (1870) edition changed the duplicated embryo images to a single image captioned "embryo of a mammal or bird". Duplication using galvanoplastic stereotypes ( clichés) was a common technique in textbooks, but not on the same page to represent different eggs or embryos. In 1891 Haeckel made the excuse that this "extremely rash foolishness" had occurred in undue haste but was "bona fide", and since repetition of incidental details was obvious on close inspection, it is unlikely to have been intentional deception. The revised 1870 second edition of 1,500 copies attracted more attention, being quickly followed by further revised editions with larger print runs as the book became a prominent part of the optimistic, nationalist, anticlerical "culture of progress" in Otto von Bismarck's new
German Empire The German Empire (),Herbert Tuttle wrote in September 1881 that the term "Reich" does not literally connote an empire as has been commonly assumed by English-speaking people. The term literally denotes an empire – particularly a hereditary ...
. The similarity of early vertebrate embryos became common knowledge, and the illustrations were praised by experts such as Michael Foster of the
University of Cambridge , mottoeng = Literal: From here, light and sacred draughts. Non literal: From this place, we gain enlightenment and precious knowledge. , established = , other_name = The Chancellor, Masters and Schola ...
. In the introduction to his 1871 '' The Descent of Man, and Selection in Relation to Sex'', Darwin gave particular praise to Haeckel, writing that if ''Natürliche Schöpfungsgeschichte'' "had appeared before my essay had been written, I should probably never have completed it". The first chapter included an illustration: "As some of my readers may never have seen a drawing of an embryo, I have given one of man and another of a dog, at about the same early stage of development, carefully copied from two works of undoubted accuracy" with a footnote citing the sources and noting that "Häckel has also given analogous drawings in his ''Schöpfungsgeschichte.''" The fifth edition of Haeckel's book appeared in 1874, with its frontispiece a heroic portrait of Haeckel himself, replacing the previous controversial image of the heads of apes and humans.


Controversy

Later in 1874, Haeckel's simplified embryology textbook ''Anthropogenie'' made the subject into a battleground over Darwinism aligned with Bismarck's '' Kulturkampf'' ("culture struggle") against the Catholic Church. Haeckel took particular care over the illustrations, changing to the leading zoological publisher Wilhelm Engelmann of Leipzig and obtaining from them use of illustrations from their other textbooks as well as preparing his own drawings including a dramatic double page illustration showing "early", "somewhat later" and "still later" stages of 8 different vertebrates. Though Haeckel's views had attracted continuing controversy, there had been little dispute about the embryos and he had many expert supporters, but Wilhelm His revived the earlier criticisms and introduced new attacks on the 1874 illustrations. Others joined in: both expert anatomists and Catholic priests and supporters were politically opposed to Haeckel's views. While it has been widely claimed that Haeckel was charged with fraud by five professors and convicted by a university court at Jena, there does not appear to be an independently verifiable source for this claim. Recent analyses (Richardson 1998, Richardson and Keuck 2002) have found that some of the criticisms of Haeckel's embryo drawings were legitimate, but others were unfounded. There were multiple versions of the embryo drawings, and Haeckel rejected the claims of fraud. It was later said that "there is evidence of sleight of hand" on both sides of the feud between Haeckel and Wilhelm His.
Robert J. Richards Robert J. Richards (born 1942) is an author and the Morris Fishbein Distinguished Service Professor of the History of Science and Medicine at the University of Chicago. He has written or edited seven books about the history of science as well as ...
, in a paper published in 2008, defends the case for Haeckel, shedding doubt against the fraud accusations based on the material used for comparison with what Haeckel could access at the time.


Awards and honors

Haeckel was elected as a member to the
American Philosophical Society The American Philosophical Society (APS), founded in 1743 in Philadelphia, is a scholarly organization that promotes knowledge in the sciences and humanities through research, professional meetings, publications, library resources, and communit ...
in 1885. He was awarded the title of Excellency by Kaiser Wilhelm II in 1907 and the
Linnean Society of London The Linnean Society of London is a learned society dedicated to the study and dissemination of information concerning natural history, evolution, and taxonomy. It possesses several important biological specimen, manuscript and literature coll ...
's prestigious Darwin-Wallace Medal in 1908. In the United States, ''
Mount Haeckel Mount Haeckel is a mountain summit located on the crest of the Sierra Nevada mountain range in northern California, United States. It is situated on the shared boundary of Kings Canyon National Park with John Muir Wilderness, and along the com ...
'', a summit in the Eastern Sierra Nevada, overlooking the Evolution Basin, is named in his honour, as is another ''Mount Haeckel'', a summit in
New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island coun ...
; and the asteroid 12323 Haeckel. In Jena he is remembered with a monument at Herrenberg (erected in 1969), an exhibition at Ernst-Haeckel-Haus, and at the
Jena Phyletic Museum The Phyletic Museum in 2012 Jena Phyletisches Museum (or Jena Phyletic Museum) is a museum in the German town of Jena. It was established by the scientist Ernst Haeckel, as an institute dedicated to explaining evolution to the public. Exhibits i ...
, which continues to teach about evolution and share his work to this day. The ratfish, '' Harriotta haeckeli'' is named in his honor. The research vessel ''Ernst Haeckel'' is named in his honor. In 1981, a botanical journal called ''Ernstia'' was started being published in the city of Maracay, Venezuela. In 2013, '' Ernstia'', a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
of calcareous sponges in the family Clathrinidae. The genus was erected to contain five species previously assigned to '' Clathrina''. The genus name honors Ernst Haeckel for his contributions towards sponge taxonomy and phylogeny.


Publications

Darwin's 1859 book ''
On the Origin of Species ''On the Origin of Species'' (or, more completely, ''On the Origin of Species by Means of Natural Selection, or the Preservation of Favoured Races in the Struggle for Life''),The book's full original title was ''On the Origin of Species by Me ...
'' had immense popular influence, but although its sales exceeded its publisher's hopes it was a technical book rather than a work of popular science: long, difficult and with few illustrations. One of Haeckel's books did a great deal to explain his version of "
Darwinism Darwinism is a theory of biological evolution developed by the English naturalist Charles Darwin (1809–1882) and others, stating that all species of organisms arise and develop through the natural selection of small, inherited variations tha ...
" to the world. It was a bestselling, provocatively illustrated book in German, titled ''Natürliche Schöpfungsgeschichte'', published in Berlin in 1868, and translated into English as ''The History of Creation'' in 1876. Until 1909, eleven editions had appeared, as well as 25 translations into other languages. The ''Natürliche Schöpfungsgeschichte'' cemented Haeckel's reputation as one of Germany's most forceful popularizers of science. His ''Welträthsel'' were reprinted ten times after the book's first publication in 1899; ultimately, over 400,000 copies were sold. Haeckel argued that human evolution consisted of precisely 22 phases, the 21st – the " missing link" – being a halfway step between apes and humans. He even formally named this missing link ''Pithecanthropus alalus'', translated as "ape man without speech". Haeckel's literary output was extensive, including many books, scientific papers, and illustrations.


Monographs

* ''Radiolaria'' (1862) * ''Siphonophora'' (1869) * ''Monera'' (1870) * ''Calcareous Sponges'' (1872)


''Challenger'' reports

* ''Deep-Sea Medusae'' (1881) * ''Siphonophora'' (1888) * ''Deep-Sea Keratosa'' (1889) * ''Radiolaria'' (1887)


Books on biology and its philosophy

* ''Generelle Morphologie der Organismen: allgemeine Grundzüge der organischen Formen-Wissenschaft, mechanisch begründet durch die von Charles Darwin reformirte Descendenz-Theorie.'' (1866) Berlin (''General morphology of organisms: general foundations of form-science, mechanically grounded by the descendance theory reformed by Charles Darwin'')
''Natürliche Schöpfungsgeschichte''
(1868); in Englis

(1876; 6th ed.: New York, D. Appleton and Co., 1914, 2 volumes) * ''Freie Wissenschaft und freie Lehre'' (1877), in English, ''Free Science and Free Teaching'' * ''Die systematische Phylogenie'' (1894) – ''Systematic Phylogeny'' * * ''Die Welträthsel'' (1895–1899), also spelled ''Die Welträtsel'' – in English '' The Riddle of the Universe: At the Close of the Nineteenth Century'', Translated by Joseph McCabe, New York and London: Harper & Brothers Publishers, 1900. * ''Über unsere gegenwärtige Kenntnis vom Ursprung des Menschen'' (1898) (''On our current understanding of the origin of man'') – in English ''The Last Link'', 1898 * ''Der Kampf um den Entwickelungsgedanken'' (1905) (''The struggle over thought on evolution'') – in English ''Last Words on Evolution: A Popular Retrospect and Summary'', Translated from the Second Edition by Joseph McCabe, New York: Peter Eckler, Publisher; London: A. Owen & Co., 1906. * ''Die Lebenswunder'' (1904) – in English
The Wonders of Life
' *
Kristallseelen : Studien über das anorganische Leben
' (1917) (''Crystal souls: studies on inorganic life'')


Travel books

* ''Indische Reisebriefe'' (1882) – ''Travel notes of India'' * ''Aus Insulinde: Malayische Reisebriefe'' (1901) – ''Travel notes of Malaysia'' * ''
Kunstformen der Natur (known in English as ''Art Forms in Nature'') is a book of lithographic and halftone prints by German biologist Ernst Haeckel. ...
'' (1904) – ''Art forms of Nature''
Digital Edition (1924)
* ''Wanderbilder'' (1905) – "Travel Images" *
A visit to Ceylon
' For a fuller list of works of and about Haeckel, see his entry in the German Wikisource.


Assessments of potential influence on Nazism

Some historians have seen Haeckel's social Darwinism as a forerunner to
Nazi Nazism ( ; german: Nazismus), the common name in English for National Socialism (german: Nationalsozialismus, ), is the far-right totalitarian political ideology and practices associated with Adolf Hitler and the Nazi Party (NSDAP) in ...
ideology. Others have denied the relationship altogether. The evidence is in some respects ambiguous. On one hand, Haeckel was an advocate of scientific racism. He held that evolutionary biology had definitively proven that races were unequal in intelligence and ability, and that their lives were also of unequal value, e.g., "These lower races (such as the Veddahs or Australian negroes) are psychologically nearer to the mammals (apes or dogs) than to civilised Europeans; we must therefore, assign a totally different value to their lives." As a result of the "struggle for existence", it followed that the "lower" races would eventually be exterminated. He was also a social Darwinist who believed that "survival of the fittest" was a natural law, and that struggle led to improvement of the race. As an advocate of eugenics, he also believed that about 200,000 mentally and congenitally ill should be killed by a medical control board. This idea was later put into practice by
Nazi Germany Nazi Germany (lit. "National Socialist State"), ' (lit. "Nazi State") for short; also ' (lit. "National Socialist Germany") (officially known as the German Reich from 1933 until 1943, and the Greater German Reich from 1943 to 1945) was ...
, as part of the Aktion T4 program. Alfred Ploetz, founder of the German Society for Racial Hygiene, praised Haeckel repeatedly, and invited him to become an honorary member. Haeckel accepted the invitation. Haeckel also believed that Germany should be governed by an authoritarian political system, and that inequalities both within and between societies were an inevitable product of evolutionary law. Haeckel was also an extreme German nationalist who believed strongly in the superiority of German culture. On the other hand, Haeckel was not an anti-Semite. In the racial hierarchies he constructed Jews tended to appear closer to the top, rather than closer to the bottom as in Nazi racial thought. He was also a pacifist until the First World War, when he wrote propaganda in favor of the war. The principal arguments of historians who deny a meaningful connection between Haeckel and Nazism are that Haeckel's ideas were very common at the time, that Nazis were much more strongly influenced by other thinkers, and that Haeckel is properly classified as a 19th-century German liberal, rather than a forerunner to Nazism. They also point to incompatibilities between evolutionary biology and Nazi ideology. Nazis themselves divided on the question of whether Haeckel should be counted as a pioneer of their ideology. SS captain and biologist Heinz Brücher wrote a biography of Haeckel in 1936, in which he praised Haeckel as a "pioneer in biological state thinking". This opinion was also shared by the scholarly journal, ''Der Biologe'', which celebrated Haeckel's 100th birthday, in 1934, with several essays acclaiming him as a pioneering thinker of Nazism. Other Nazis kept their distance from Haeckel. Nazi propaganda guidelines issued in 1935 listed books which popularized Darwin and evolution on an "expunged list". Haeckel was included by name as a forbidden author. Gunther Hecht, a member of the Nazi Department of Race Politics, also issued a memorandum rejecting Haeckel as a forerunner of Nazism. Kurt Hildebrandt, a Nazi political philosopher, also rejected Haeckel. Eventually Haeckel was rejected by Nazi bureaucrats.


See also

* Dysteleology *
Embryology Embryology (from Greek ἔμβρυον, ''embryon'', "the unborn, embryo"; and -λογία, '' -logia'') is the branch of animal biology that studies the prenatal development of gametes (sex cells), fertilization, and development of embr ...
* Haeckelites * '' Haeckel's Tale'' *
Heinrich Schmidt (philosopher) Heinrich Schmidt (December 18, 1874 – May 2, 1935) was a German archivist, naturalist, philosopher, professor and a student of Ernst Haeckel. Early life and education Schmidt was born in Heubach in the German State of Thuringia. From 1 ...
* Karl Blossfeldt *
List of wildlife artists This list of wildlife artists is a list for any notable wildlife artist, wildlife painter, wildlife photographer, other wildlife artist, society of wildlife artists, museum, or exhibition of wildlife art, worldwide. A * Jackson Miles Abbott * Jo ...
* '' Magosphaera planula'' * ''Proteus'' (2004 film)


Footnotes


Sources

* * * *


External links


E. Haeckel: Natürliche Schöpfungsgeschichte 1868 (front page of 1st edition, German)

E. Haeckel: Die Welträthsel 1899 (front page of 1st edition, German)


– biography
Ernst Haeckel – Evolution's controversial artist.
A slide-show essay

(fro
biolib.de
*
Kunstformen der Natur
' (Digitization from Phaidra)
PNG alpha-transparencies of Haeckel's "Kustformen der natur"


– Animated documentary film on Haeckel's life and work

and Museum in
Jena Jena () is a German city and the second largest city in Thuringia. Together with the nearby cities of Erfurt and Weimar, it forms the central metropolitan area of Thuringia with approximately 500,000 inhabitants, while the city itself has a po ...
* View works b
Haeckel
at the Biodiversity Heritage Library
aDiatomea: artificial life experiment with 3d generated diatoms, influenced by Haeckel

Images from ''Anthropogenie, oder, Entwickelungsgeschichte des menschen''
* * * *
Ernst Haeckel's Radiolarians and Medusa
– article on Haeckel in Villefranche-sur-Mer {{DEFAULTSORT:Haeckel, Ernst 1834 births 1919 deaths Alldeutscher Verband members Evolutionary biologists German atheists 19th-century German biologists German ecologists German humanists German male writers 19th-century German philosophers 19th-century German zoologists Lemuria (continent) Natural history illustrators Scientists from Potsdam People from the Province of Brandenburg Protistologists Proponents of scientific racism University of Jena faculty Spinozists Members of the American Philosophical Society People involved in scientific misconduct incidents