Eotitanosuchus
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''Eotitanosuchus'' ("dawn giant crocodile") is an extinct
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of biarmosuchian therapsids whose
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
s were found in the town of Ochyor in
Perm Krai Perm Krai (, ; ) is a federal subjects of Russia, federal subject of Russia (a Krais of Russia, krai), located in Eastern Europe. Its administrative center is Perm, Russia, Perm. The population of the krai was 2,532,405 (2021 Russian census, 2021 ...
,
Russia Russia, or the Russian Federation, is a country spanning Eastern Europe and North Asia. It is the list of countries and dependencies by area, largest country in the world, and extends across Time in Russia, eleven time zones, sharing Borders ...
. It lived about 267 million
year A year is a unit of time based on how long it takes the Earth to orbit the Sun. In scientific use, the tropical year (approximately 365 Synodic day, solar days, 5 hours, 48 minutes, 45 seconds) and the sidereal year (about 20 minutes longer) ...
s ago. The only species is ''Eotitanosuchus olsoni''.


Description

''Eotitanosuchus'' is known from a single large skull without a lower jaw. The skull was , but the overall length may have been over , possibly up to and more than in weight for adult specimens. Like '' Biarmosuchus tener'', it was primitive in that, though it was a predator, the temple opening behind the eye was small, giving it a weak bite. The temple was, however, larger at the top than in other biarmosuchians.


Paleobiology

''Eotitanosuchus'' fossils were found in the Perm (or Cis-Urals) region of Russia. ''Eotitanosuchus'' was without doubt a dominant animal of its environment. Found preserved in flood deposits (once coastal bogs) containing many skeletons of estemmenosuchids, it has been suggested that this large predator was an excellent swimmer, possibly semi-aquatic or frequenting marshy ground. This, however, is just speculation.


Classification

''Eotitanosuchus'' is often grouped with the Phthinosuchidae and the Biarmosuchidae. In fact, Ivakhnenko (1999) argues that '' Biarmosuchus tener'' and ''Eotitanosuchus olsoni'' are the same organism, which would eliminate the Eotitanosuchia as a separate taxon, though this conclusion does not seem to have been widely accepted. Regardless of the eventual outcome of this debate, Ivakhnenko's paper does seem to show that ''Eotitanosuchus'' is very similar to ''Biarmosuchus''. Further, given the rather close similarity between ''Eotitanosuchus'' and later therapsids, this observation supports the view that Biarmosuchia is
paraphyletic Paraphyly is a taxonomic term describing a grouping that consists of the grouping's last common ancestor and some but not all of its descendant lineages. The grouping is said to be paraphyletic ''with respect to'' the excluded subgroups. In co ...
. Others view ''Eotitanosuchus'' as quite distinct from other basal therapsids and perhaps closer to the
Gorgonopsia Gorgonopsia (from the Greek Gorgon, a mythological beast, and 'aspect') is an extinct clade of Saber-toothed predator, sabre-toothed therapsids from the Middle Permian, Middle to the Upper Permian, roughly between 270 and 252 million years ago. ...
but gorgonopsian specializations are either not present in ''Eotitanosuchus'' or, as is more often the case, the state of the characters is unknown. This genus is characterized by many primitive features of the septomaxilla, the postorbital, the parietal, the interparietal, the basioccipital, the quadrate rami of the pterygoid and the vomers of the skull. The length of the dorsal process of the premaxilla (front jawbone) and the postorbital twisting (rear side of the skull) constitute specializations that indicate it is not a direct gorgonopsian ancestor. These features, however, are shared by the anteosaur and biarmosuchid lineages.


See also

*
List of therapsids This list of therapsids is an attempt to create a comprehensive listing of all Genus, genera that have ever been included in the Therapsida excluding mammals and purely vernacular terms. The list includes all commonly accepted genera, but also gene ...


References

* * * Patricia Vickers-Rich and Thomas H. Rich, '' The Great Russian Dinosaurs'', Gunter Graphics, 1993, pg. 28.


External links


Eotitanosuchidae
at Kheper



at Palaeos * ttp://palaeos.com/vertebrates/therapsida/biarmosuchia.html#Biarmosuchia Therapsida: Biarmosuchiaat Palaeos Biarmosuchia Prehistoric therapsid genera Wordian genera Guadalupian synapsids of Europe Permian Russia Fossils of Russia Fossil taxa described in 1960 {{paleo-therapsid-stub