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Emballonuridae is a
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
of microbats, many of which are referred to as sac-winged or sheath-tailed bats. They are widely distributed in
tropical The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in the Southern Hemisphere at S. The tropics are also referred to ...
and subtropical regions around the world. The earliest fossil records are from the
Eocene The Eocene ( ) Epoch is a geological epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period in the modern Cenozoic Era. The name ''Eocene'' comes from the Ancient Greek (''ēṓs'', " ...
.


Description

The emballonurids include some of the smallest of all bats, and range from 3.5 to 10 cm in body length. They are generally brown or grey, although the species of genus ''
Diclidurus ''Diclidurus'' is a genus of bats whose common name is the ghost bats (not to be confused with the Australia ''Macroderma gigas''). ''Diclidurus'' all inhabit tropical South America, and ''D. albus'' is also found in Mexico and Central America.E ...
'' are white. The faces are said to be handsome, the heads being comparable to those of domestic dogs, and their wings are long and narrow. As with other microchiropteran families, they use ultrasonic echolocation to sense the surrounding environment and their prey; the signals of some species are unusual in being audible to humans. Possession of the postorbital processes, the reduced, noncontacting premaxillaries, and rather simple shoulder and elbow joints, which is similar to pteropodids, makes them rather a primitive group. However, they are more advanced in the reduction of the second digit phalanges and the flexion of the third digit proximal phalanges over the metacarpal dorsal side. The wing surface extends between the legs, a membrane known as a
uropatagium The patagium (plural: patagia) is a membranous body part that assists an animal in obtaining lift when gliding or flight. The structure is found in extant and extinct groups of flying and gliding animals including bats, birds, some dromaeosau ...
, and the structure of these is a characteristic in many of the genera. They have tails which are partially enclosed, a short part of which projects through the uropatagium to form a sheath. The usual arrangement of the uropatagium is as to be fixed to the tail, but the sheathtail feature is joined by an elastic component which allows greater flexibility; they are able to use the hind legs for locomotion and to adjust the membrane's surface while in flight. As a common name indicates, many species also possess sac-shaped glands in their wings (
propatagium The patagium (plural: patagia) is a membranous body part that assists an animal in obtaining lift when gliding or flight. The structure is found in extant and extinct groups of flying and gliding animals including bats, birds, some dromaeosaurs ...
), which are open to the air and may release
pheromone A pheromone () is a secreted or excreted chemical factor that triggers a social response in members of the same species. Pheromones are chemicals capable of acting like hormones outside the body of the secreting individual, to affect the behavio ...
s to attract mates. Other species have throat glands which produce strong-smelling secretions. They have the
dental formula Dentition pertains to the development of teeth and their arrangement in the mouth. In particular, it is the characteristic arrangement, kind, and number of teeth in a given species at a given age. That is, the number, type, and morpho-physiolog ...
These bats generally prefer to roost in better-illuminated areas than other species of bats. Their dwellings can often be found in hollow trees and entryways to caves or other structures. Some species, such as the genus ''
Taphozous ''Taphozous'' is a genus of the family Emballonuridae. The wide distribution of the genus includes several regions of Australia, Indonesia, Papua New Guinea and Africa. ''Taphozous'' comes from the Greek τάφος, meaning "a tomb". The common n ...
'', live in large colonies, but others are solitary. Species living away from the tropics may enter periods of
torpor Torpor is a state of decreased physiological activity in an animal, usually marked by a reduced body temperature and metabolic rate. Torpor enables animals to survive periods of reduced food availability. The term "torpor" can refer to the time ...
or extended hibernation during colder months. Emballonurids feed mainly on insects and occasionally on fruit. Most of these bats catch their meals while flying. The common name for some groups, 'sheath-tailed bats', is sometimes noted as sheathtails.


Distribution

Found in the
Neotropics The Neotropical realm is one of the eight biogeographic realms constituting Earth's land surface. Physically, it includes the tropical terrestrial ecoregions of the Americas and the entire South American temperate zone. Definition In bioge ...
, Afrotropics, southern
Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an are ...
, Australia and South Pacific islands.


Classification

Family Emballonuridae *Genus ''
Balantiopteryx ''Balantiopteryx'' is a genus of sac-winged bats. It comprises three species: *Ecuadorian sac-winged bat - ''B. infusca'' *Thomas's sac-winged bat - ''B. io'' *Gray sac-winged bat The gray sac-winged bat (''Balantiopteryx plicata'') is a spe ...
'' **
Ecuadorian sac-winged bat The Ecuadorian sac-winged bat (''Balantiopteryx infusca'') is a species of sac-winged bat in the family Emballonuridae. It is found in Colombia and Ecuador. According to the IUCN Red List of Threatened Species, the population trend is decreasing ...
, ''Balantiopteryx infusca'' **
Thomas's sac-winged bat Thomas's sac-winged bat (''Balantiopteryx io'') is a species of sac-winged bat in the family Emballonuridae. It is found in Belize, Guatemala, and Mexico. Taxonomy Oldfield Thomas named this bat in 1904. It is presumed that he named it after the ...
, ''Balantiopteryx io'' **
Gray sac-winged bat The gray sac-winged bat (''Balantiopteryx plicata'') is a species in the family Emballonuridae which comprises the 51 species of sac-winged bats. It is found in Mexico from Baja California Sur and Sonora to Guatemala, El Salvador, Honduras, Nica ...
, ''Balantiopteryx plicata'' *Genus ''
Centronycteris ''Centronycteris'' is a genus of sac-winged bat Emballonuridae is a family of microbats, many of which are referred to as sac-winged or sheath-tailed bats. They are widely distributed in tropical and subtropical regions around the world. The ...
'' **
Thomas's shaggy bat The Thomas's shaggy bat (''Centronycteris centralis'') is a bat species from Central and South America. It was previously included in the shaggy bat but Simmons and Handley (1998) showed that the species were distinct. Taxonomy It was describ ...
, ''Centronycteris centralis'' **
Shaggy bat The shaggy bat (''Centronycteris maximiliani'') is a bat species from northern South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in t ...
, ''Centronycteris maximiliani'' *Genus '' Coleura'' ** African sheath-tailed bat, ''Coleura afra'' ** Madagascar sheath-tailed bat, ''Coleura kibomalandy'' ** Seychelles sheath-tailed bat, ''Coleura seychellensis'' *Genus ''
Cormura The chestnut sac-winged bat, or Wagner's sac-winged bat (''Cormura brevirostris'') is a species of sac-winged bat native to South and Central America. It is monotypic taxon, the only species within its genus. Description The chestnut sac-winged ...
'' ** Chestnut sac-winged bat, ''Cormura brevirostris'' *Genus '' Cyttarops'' ** Short-eared bat, ''Cyttarops alecto'' *Genus ''
Diclidurus ''Diclidurus'' is a genus of bats whose common name is the ghost bats (not to be confused with the Australia ''Macroderma gigas''). ''Diclidurus'' all inhabit tropical South America, and ''D. albus'' is also found in Mexico and Central America.E ...
'' - ghost bats'' ** Northern ghost bat, ''Diclidurus albus'' **
Greater ghost bat The greater ghost bat (''Diclidurus ingens'') is a bat species found in northwestern Brazil, southeastern Colombia, Guyana, Peru, and Venezuela Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bol ...
, ''Diclidurus ingens'' **
Isabelle's ghost bat Isabelle's ghost bat (''Diclidurus isabella'') is a bat species found in northwestern Brazil, Guyana, and Venezuela, and possibly Colombia. It was discovered in October 1916 by Emilie Snethlage, and described by Oldfield Thomas Michael R ...
, ''Diclidurus isabella'' **
Lesser ghost bat The lesser ghost bat (''Diclidurus scutatus'') is a bat species found in South America. It is one of six bat species worldwide to have white fur. Taxonomy and etymology It was described by Wilhelm Peters in 1869 based on a specimen sent to the ...
, ''Diclidurus scutatus'' *Genus ''
Emballonura ''Emballonura'' (meaning: Erect tail) is a genus of sac-winged bats in the family Emballonuridae. It contains these species: * Small Asian sheath-tailed bat (''E. alecto'') * Beccari's sheath-tailed bat (''E. beccarii'') * Large-eared sheath-tai ...
'' ** Small Asian sheath-tailed bat, ''Emballonura alecto'' ** Beccari's sheath-tailed bat, ''Emballonura beccarii'' ** Large-eared sheath-tailed bat, ''Emballonura dianae'' ** Greater sheath-tailed bat, ''Emballonura furax'' ** Lesser sheath-tailed bat, ''Emballonura monticola'' ** Raffray's sheath-tailed bat, ''Emballonura raffrayana'' **
Pacific sheath-tailed bat The Pacific sheath-tailed bat or Polynesian sheath-tailed bat (''Emballonura semicaudata'') is a species of sac-winged bat in the family Emballonuridae found in American Samoa, Fiji, Guam, Micronesia, Palau, Samoa (where it is called ''pe'a vai ...
, ''Emballonura semicaudata'' ** Seri's Sheathtail-bat, ''Emballonura serii'' *Genus '' Mosia'' ** Dark sheath-tailed bat, ''Mosia nigrescens'' *Genus '' Paremballonura'' ** Western sheath-tailed bat'', Paremballonura tiavato'' ** Peters's sheath-tailed bat, ''Paremballonura atrata'' *Genus '' Peropteryx'' ** Greater dog-like bat, ''Peropteryx kappleri'' ** White-winged dog-like bat, ''Peropteryx leucoptera'' ** Lesser dog-like bat, ''Peropteryx macrotis'' ** Pale-winged dog-like bat, ''Peropteryx pallidoptera'' ** Trinidad dog-like bat, ''Peropteryx trinitatis'' *Genus '' Rhynchonycteris'' **
Proboscis bat The proboscis bat (''Rhynchonycteris naso'') is a species of bat found in South America and Central America. Other common names include Long-nosed proboscis bat, sharp-nosed bat, Brazilian long-nosed bat. and river bat It is the only species i ...
, ''Rhynchonycteris naso'' *Genus ''
Saccolaimus ''Saccolaimus'' is a genus of the family Emballonuridae, small insectivorous bats with distinctive sheathtails and pouches at the wrist. The species have been placed with genus ''Taphozous'', in subgenus ''Taphozous'', although an elevation to ...
'' ** Yellow-bellied pouched bat, ''Saccolaimus flaviventris'' ** Troughton's pouched bat, ''Saccolaimus mixtus'' ** Pel's pouched bat, ''Saccolaimus peli'' **
Naked-rumped pouched bat The naked-rumped pouched bat (''Saccolaimus saccolaimus''), also known as the pouched tomb bat, is a species of sac-winged bat in the family Emballonuridae. Taxonomy Described in 1838 by Coenraad Temminck. the author assigned the species to a n ...
, ''Saccolaimus saccolaimus'' *Genus ''
Saccopteryx ''Saccopteryx'' is a genus of sac-winged bats from Central and South America. The species within this genus are: * Antioquian sac-winged bat ''Saccopteryx antioquensis'' *Greater sac-winged bat The greater sac-winged bat (''Saccopteryx bili ...
'' ** Antioquian sac-winged bat, ''Saccopteryx antioquensis'' **
Greater sac-winged bat The greater sac-winged bat (''Saccopteryx bilineata'') is a bat of the family Emballonuridae native to Central and South America. They are the most common bats seen in the rainforest, as they often roost on the outside of large trees. They are ...
, ''Saccopteryx bilineata'' ** Frosted sac-winged bat, ''Saccopteryx canescens'' ** Amazonian sac-winged bat, ''Saccopteryx gymnura'' ** Lesser sac-winged bat, ''Saccopteryx leptura'' *Genus ''
Taphozous ''Taphozous'' is a genus of the family Emballonuridae. The wide distribution of the genus includes several regions of Australia, Indonesia, Papua New Guinea and Africa. ''Taphozous'' comes from the Greek τάφος, meaning "a tomb". The common n ...
'' ** Indonesian tomb bat (''Taphozous achates'') ** Coastal sheath-tailed bat (''Taphozous australis'') ** Common sheath-tailed bat (''Taphozous georgianus'') ** Hamilton's tomb bat (''Taphozous hamiltoni'') **
Hildegarde's tomb bat Hildegarde's tomb bat (''Taphozous hildegardeae'') is a species of sac-winged bat in the family Emballonuridae. It is found near the coast in Kenya and Tanzania where it feeds in tropical dry forests and roosts in caves. It is a diurnal species a ...
(''Taphozous hildegardeae'') ** Hill's sheath-tailed bat (''Taphozous hilli'') ** Arnhem sheath-tailed bat (''Taphozous kapalgensis'') ** Long-winged tomb bat (''Taphozous longimanus'') ** Mauritian tomb bat (''Taphozous mauritianus'') **
Black-bearded tomb bat The black-bearded tomb bat (''Taphozous melanopogon'') is a species of sac-winged bat found in South and South East Asia. Taxonomy and etymology It was described as a new species in 1841 by Dutch zoologist Coenraad Jacob Temminck. The holotype ...
(''Taphozous melanopogon'') ** Naked-rumped tomb bat (''Taphozous nudiventris'') ** Egyptian tomb bat (''Taphozous perforatus'') ** Theobald's tomb bat (''Taphozous theobaldi'') ** Troughton's sheath-tailed bat (''Taphozous troughtoni'')


References

{{Taxonbar, from=Q838530 Bat families