''Eleutherodactylus'' is a

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...

frog A frog is any member of a diverse and largely Carnivore, carnivorous group of short-bodied, tailless amphibians composing the order (biology), order Anura (ανοὐρά, literally ''without tail'' in Ancient Greek). The oldest fossil "proto-f ...

s in the family Eleutherodactylidae.Hedges, S. B., W. E. Duellman, and M. P. Heinicke . 2008. New World direct-developing frogs (Anura: Terrarana): molecular phylogeny, classification, biogeography, and conservation. Zootaxa 1737: 1-182. Many of the 200 species of the genus are commonly known as "rain frogs" or "robber frogs", due to their sharp, high-pitched, insect-like calls. The best-known species is the

common coquí The common coquí or coquí (''Eleutherodactylus coqui'') is a species of frog endemic to Puerto Rico belonging to the family Eleutherodactylidae. The species is named for the loud call the males make at night. This sound serves two purposes. "C ...

(''E. coqui''), which is both a national symbol of

Puerto Rico Puerto Rico (; abbreviated PR; tnq, Boriken, ''Borinquen''), officially the Commonwealth of Puerto Rico ( es, link=yes, Estado Libre Asociado de Puerto Rico, lit=Free Associated State of Puerto Rico), is a Caribbean island and unincorporated ...

and a notorious

invasive species An invasive species otherwise known as an alien is an introduced organism that becomes overpopulated and harms its new environment. Although most introduced species are neutral or beneficial with respect to other species, invasive species adv ...

Hawaii Hawaii ( ; haw, Hawaii or ) is a state in the Western United States, located in the Pacific Ocean about from the U.S. mainland. It is the only U.S. state outside North America, the only state that is an archipelago, and the only stat ...

. Two ''Eleutherodactylus'' species, '' E. limbatus'' and '' E. iberia'', are among the smallest known frogs, measuring only 8.5 mm in length (only slightly larger than ''

Paedophryne amauensis ''Paedophryne amauensis'' is a species of microhylid frog endemic to eastern Papua New Guinea. At in snout-to-vent length, it is considered the world's smallest known vertebrate. (See also Ecological guild.) The species was listed in the ...

'', which measures around 7.7 mm).

Etymology

The name "Eleutherodactylus" is derived from the

Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...

words for ‘free-toed’, composed of the Ancient Greek ' (, ‘free, unbound’) and ' (, ‘finger, toe’). Most species are small, slender, and cryptically colored, with three to five free toes. A few, such as the web-footed coquí (''E. karlschmidti'') of Puerto Rico, do have completely webbed feet.

Distribution and habitat

Species of ''Eleutherodactylus'' are found throughout the Neotropics, including the southern United States, Mexico, Central America, and the Caribbean. Additionally, the common coquí (''E. coqui'') has been introduced to several islands in the

an archipelago, as well as elsewhere in the Pacific. They can be terrestrial, arboreal, or aquatic, typically living in forests or

riparian A riparian zone or riparian area is the interface between land and a river or stream. Riparian is also the proper nomenclature for one of the terrestrial biomes of the Earth. Plant habitats and communities along the river margins and banks a ...

areas, and feeding primarily upon

arthropod Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and cuticle made of chiti ...

s. Many ''Eleutherodactylus'' species have highly restricted ranges and are found on only one island or in one or a few localities. Even some of these restricted species can occur at very high densities.

Reproduction and development

All species of ''Eleutherodactylus'' are characterized by direct development, in which eggs hatch directly into small frogs, completely bypassing the

tadpole A tadpole is the larval stage in the biological life cycle of an amphibian. Most tadpoles are fully aquatic, though some species of amphibians have tadpoles that are terrestrial. Tadpoles have some fish-like features that may not be found ...

stage. This adaptation may be largely responsible for their ecological and evolutionary success. Most species are characterized by parental behaviors, such as egg-guarding by either the male or female parent. In some cases, even young froglets are attended by parents. Another extinct Puerto Rican species, the golden coquí (''E. jasperi''), gave birth to live young. Many species (for example, Cook's robber frog, ''E. cooki''), also of Puerto Rico, exhibit

sexual dimorphism Sexual dimorphism is the condition where the sexes of the same animal and/or plant species exhibit different morphological characteristics, particularly characteristics not directly involved in reproduction. The condition occurs in most an ...

in size and color. Study on ''Eleutherodactylus'' and ''

Lithobates ''Lithobates'' is a genus of true frogs, of the family Ranidae. The name is derived from '' litho-'' (stone) and the Greek ' (, one that treads), meaning one that treads on rock, or rock climber. The name was defined by Hillis and Wilcox (2005) ...

'' amphibians shows that number of offsprings instead of body size may help to find which species require conservation from being extinct.

Phylogenetics

The basis of forming this genus has been morphological, but sequence comparisons of

protein Proteins are large biomolecules and macromolecules that comprise one or more long chains of amino acid residues. Proteins perform a vast array of functions within organisms, including catalysing metabolic reactions, DNA replication, res ...

-encoding DNA,

mitochondrial A mitochondrion (; ) is an organelle found in the cells of most Eukaryotes, such as animals, plants and fungi. Mitochondria have a double membrane structure and use aerobic respiration to generate adenosine triphosphate (ATP), which is used t ...

DNA, and

ribosomal Ribosomes ( ) are macromolecular machines, found within all cells, that perform biological protein synthesis (mRNA translation). Ribosomes link amino acids together in the order specified by the codons of messenger RNA (mRNA) molecules to for ...

RNA have shown geographic range is a much more consistent predictor of

cladistics Cladistics (; ) is an approach to biological classification in which organisms are categorized in groups (" clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is typically shared derived cha ...

for this group of frogs. The climbing habits of many species have

evolved independently Convergent evolution is the independent evolution of similar features in species of different periods or epochs in time. Convergent evolution creates analogous structures that have similar form or function but were not present in the last co ...

. All true members of the genus have been clustered into

subgenera In biology, a subgenus (plural: subgenera) is a taxonomic rank directly below genus. In the International Code of Zoological Nomenclature, a subgeneric name can be used independently or included in a species name, in parentheses, placed between ...

, but many less-related species require more genetic data before they are to be officially classified elsewhere. The theory that the eleutherodactyline colonization of

Central America Central America ( es, América Central or ) is a subregion of the Americas. Its boundaries are defined as bordering the United States to the north, Colombia to the south, the Caribbean Sea to the east, and the Pacific Ocean to the west. ...

and the

Caribbean The Caribbean (, ) ( es, El Caribe; french: la Caraïbe; ht, Karayib; nl, De Caraïben) is a region of the Americas that consists of the Caribbean Sea, its islands (some surrounded by the Caribbean Sea and some bordering both the Caribbean ...

from their origins in South America occurred during the

Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of ...

has fallen out of favor. The

fossil record A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...

, combined with

molecular clock The molecular clock is a figurative term for a technique that uses the mutation rate of biomolecules to deduce the time in prehistory when two or more life forms diverged. The biomolecular data used for such calculations are usually nucleo ...

analyses, indicate the subgenera were probably founded by small groups of individuals by flotsam dispersal during the

Eocene The Eocene ( ) Epoch is a geological epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period in the modern Cenozoic Era. The name ''Eocene'' comes from the Ancient Greek (''ēṓs'', ...

Oligocene The Oligocene ( ) is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that define the epoch are well identified but t ...

epochs. Land bridges would have been limited to facilitating dispersal between

West Indian A West Indian is a native or inhabitant of the West Indies (the Antilles and the Lucayan Archipelago). For more than 100 years the words ''West Indian'' specifically described natives of the West Indies, but by 1661 Europeans had begun to use it ...

islands, however, the Oligocene division of

Hispaniola Hispaniola (, also ; es, La Española; Latin and french: Hispaniola; ht, Ispayola; tnq, Ayiti or Quisqueya) is an island in the Caribbean that is part of the Greater Antilles. Hispaniola is the most populous island in the West Indies, and th ...

and

Cuba Cuba ( , ), officially the Republic of Cuba ( es, República de Cuba, links=no ), is an island country comprising the island of Cuba, as well as Isla de la Juventud and several minor archipelagos. Cuba is located where the northern Caribb ...

resulted in further speciation. The distribution of the subgenus '' Syrrhopus'' is most likely due to a secondary dispersal to Central America from the

Greater Antilles The Greater Antilles ( es, Grandes Antillas or Antillas Mayores; french: Grandes Antilles; ht, Gwo Zantiy; jam, Grieta hAntiliiz) is a grouping of the larger islands in the Caribbean Sea, including Cuba, Hispaniola, Puerto Rico, Jamaica, a ...

during the

Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...

. The formation of the

Panama Isthmus The Isthmus of Panama ( es, Istmo de Panamá), also historically known as the Isthmus of Darien (), is the narrow strip of land that lies between the Caribbean Sea and the Pacific Ocean, linking North and South America. It contains the countr ...

during the

Pliocene The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58 ''

Sensu stricto ''Sensu'' is a Latin word meaning "in the sense of". It is used in a number of fields including biology, geology, linguistics, semiotics, and law. Commonly it refers to how strictly or loosely an expression is used in describing any particular c ...

'', however, it should exclude clades with distributions south of the

Panama Canal The Panama Canal ( es, Canal de Panamá, link=no) is an artificial waterway in Panama that connects the Atlantic Ocean with the Pacific Ocean and divides North and South America. The canal cuts across the Isthmus of Panama and is a condui ...

Fossil record

Frog remains referred to ''Eleutherodactylus'' have been reported from Dominican amber.

Species

The following species are recognised in the genus ''Eleutherodactylus'': * '' Eleutherodactylus aporostegus'' Schwartz, 1965 * '' Eleutherodactylus bothroboans'' Schwartz, 1965 * ''

Eleutherodactylus campi ''Eleutherodactylus cystignathoides'', also known as the Rio Grande chirping frog, Mexican chirping frog, or lowland chirping frog, is a small Eleutherodactylidae, eleutherodactylid frog. It is found from the southern United States in Texas, and ...

'' (Stejneger, 1915) * ''

Eleutherodactylus counouspeus ''Eleutherodactylus counouspeus'', also known as Counou robber frog and yellow cave frog, is a species of frog in the family Eleutherodactylidae. It is endemic to the Massif de la Hotte, southwestern Haiti. The specific name ''counouspeus'' re ...

'' Schwartz, 1964 * ''

Eleutherodactylus diplasius ''Eleutherodactylus diplasius'', the patternless whistling frog, is a species of frog in the family Eleutherodactylidae. A native of Haiti, it can be found in the Tiburon Peninsula northwest, to the Massif de la Hotte The Massif de la Hotte is ...

West Indian A West Indian is a native or inhabitant of the West Indies (the Antilles and the Lucayan Archipelago). For more than 100 years the words ''West Indian'' specifically described natives of the West Indies, but by 1661 Europeans had begun to use it ...
(subgenus ''
Eleutherodactylus ''Eleutherodactylus'' is a genus of frogs in the family Eleutherodactylidae.Hedges, S. B., W. E. Duellman, and M. P. Heinicke . 2008. New World direct-developing frogs (Anura: Terrarana): molecular phylogeny, classification, biogeography, and c ...
'')

* '' E. (E.) abbotti'' Cochran, 1923 * '' E. (E.) amplinympha'' Kaiser, Green & Schmid, 1994 * '' E. (E.) antillensis'' Reinhardt & Lütken, 1863 * '' E. (E.) audanti'' Cochran, 1934 * '' E. (E.) auriculatoides'' Noble, 1923 * '' E. (E.) auriculatus'' Cope, 1862 * '' E. (E.) barlagnei'' Lynch, 1965 * '' E. (E.) bartonsmithi'' Schwartz, 1960 * '' E. (E.) brittoni'' Schmidt, 1920 * '' E. (E.) cochranae'' Grant, 1932 * '' E. (E.) cooki'' Grant, 1932 * '' E. (E.) coqui'' Thomas, 1966 * '' E. (E.) eileenae'' Dunn, 1926 * '' E. (E.) eneidae'' Rivero, 1959 * '' E. (E.) flavescens'' Noble, 1923 * '' E. (E.) fowleri'' Schwartz, 1973 * '' E. (E.) glamyrus'' Estrada & Hedges, 1997 * '' E. (E.) gryllus'' Schmidt, 1920 * '' E. (E.) guantanamera'' Hedges, Estrada & Thomas, 1992 * '' E. (E.) haitianus'' Barbour, 1942 * '' E. (E.) hedricki'' Rivero, 1963 * '' E. (E.) ionthus'' Schwartz, 1960 * '' E. (E.) jasperi'' Drewry & Jones, 1976 * '' E. (E.) johnstonei'' Barbour, 1914 * '' E. (E.) juanariveroi'' Ríos-López & Thomas, 2007 * '' E. (E.) karlschmidti'' Grant, 1931 * '' E. (E.) lamprotes'' Schwartz, 1973 * '' E. (E.) leberi'' Schwartz, 1965 * '' E. (E.) locustus'' Schmidt, 1920 * '' E. (E.) mariposa'' Hedges, Estrada & Thomas, 1992 * '' E. (E.) martinicensis'' Tschudi, 1838 * '' E. (E.) melacara'' Hedges, Estrada & Thomas, 1992 * '' E. (E.) minutus'' Noble, 1923 * '' E. (E.) montanus'' Schmidt, 1919 * '' E. (E.) parabates'' Schwartz, 1964 * '' E. (E.) patriciae'' Schwartz, 1965 * '' E. (E.) pinchoni'' Schwartz, 1967 * '' E. (E.) pituinus'' Schwartz, 1965 * '' E. (E.) poolei'' Cochran, 1938 * '' E. (E.) portoricensis'' Schmidt, 1927 * '' E. (E.) principalis'' Estrada & Hedges, 1997 * '' E. (E.) richmondi'' Stejneger, 1904 * '' E. (E.) ronaldi'' Schwartz, 1960 * '' E. (E.) schwartzi'' Thomas, 1966 * '' E. (E.) unicolor'' Stejneger, 1904 * ''E. (E.) varians'' Gundlach & Peters, 1864 * '' E. (E.) wetmorei'' Cochran, 1932 * '' E. (E.) wightmanae'' Schmidt, 1920

West Indian A West Indian is a native or inhabitant of the West Indies (the Antilles and the Lucayan Archipelago). For more than 100 years the words ''West Indian'' specifically described natives of the West Indies, but by 1661 Europeans had begun to use it ...
(subgenus '' Euhyas'')

* '' E. (Eu.) acmonis'' Schwartz, 1960 * '' E. (Eu.) adelus'' Diaz, Cadiz & Hedges, 2003 * '' E. (Eu.) albipes'' Barbour & Shreve, 1937 * '' E. (Eu.) alcoae'' Schwartz, 1971 * '' E. (Eu.) alticola'' Lynn, 1937 * '' E. (Eu.) amadeus'' Hedges, Thomas & Franz, 1987 * '' E. (Eu.) andrewsi'' Lynn, 1937 * '' E. (Eu.) apostates'' Schwartz, 1973 * '' E. (Eu.) armstrongi'' Noble & Hassler, 1933 * '' E. (Eu.) atknisi'' Dunn, 1925 * '' E. (Eu.) bakeri'' Cochran, 1935 * '' E. (Eu.) beguei'' Díaz and Hedges, 2015 * '' E. (Eu.) blairhedgesi'' Estrada, Díaz & Rodriguez, 1998 * '' E. (Eu.) bresslerae'' Schwartz, 1960 * '' E. (Eu.) brevirostris'' Shreve, 1936 * '' E. (Eu.) caribe'' Hedges & Thomas, 1992 * '' E. (Eu.) casparii'' Dunn, 1926 * '' E. (Eu.) cavernicola'' Lynn, 1954 * '' E. (Eu.) corona'' Hedges & Thomas, 1992 * '' E. (Eu.) cubanus'' Barbour, 1942 * '' E. (Eu.) cundalli'' Dunn, 1926 * '' E. (Eu.) cuneatus'' Cope, 1862 * '' E. (Eu.) darlingtoni'' Cochran, 1935 * '' E. (Eu.) dimidiatus'' Cope, 1862 * '' E. (Eu.) dolomedes'' Hedges & Thomas, 1992 * '' E. (Eu.) emiliae'' Dunn, 1926 * '' E. (Eu.) etheridgei'' Schwartz, 1958 * '' E. (Eu.) eunaster'' Schwartz, 1973 * '' E. (Eu.) furcyensis'' Shreve & Williams, 1963 * '' E. (Eu.) fuscus'' Lynn & Dent, 1943 * '' E. (Eu.) glandulifer'' Cochran, 1935 * '' E. (Eu.) glanduliferoides'' Shreve, 1936 * '' E. (Eu.) glaphycompus'' Schwartz, 1973 * '' E. (Eu.) glaucoreius'' Schwartz & Fowler, 1973 * '' E. (Eu.) goini'' Schwartz, 1960 * '' E. (Eu.) gossei'' Dunn, 1926 * '' E. (Eu.) grabhami'' Dunn, 1926 * '' E. (Eu.) grahami'' Schwartz, 1979 * '' E. (Eu.) greyi'' Dunn, 1926 * '' E. (Eu.) griphus'' Crombie, 1986 * '' E. (Eu.) guanahacabibes'' Estrada & Rodriguez, 1985 * '' E. (Eu.) gundlachi'' Schmidt, 1920 * '' E. (Eu.) heminota'' Shreve & Williams, 1963 * '' E. (Eu.) iberia'' Estrada & Hedges, 1996 * '' E. (Eu.) intermedius'' Barbour & Shreve, 1937 * '' E. (Eu.) jamaicensis'' Barbour, 1910 * '' E. (Eu.) jaumei'' Estrada & Alonso, 1997 * '' E. (Eu.) jugans'' Cochran, 1937 * '' E. (Eu.) junori'' Dunn, 1926 * '' E. (Eu.) klinikowskii'' Schwartz, 1959 * '' E. (Eu.) lentus'' Cope, 1862 * '' E. (Eu.) leoncei'' Shreve & Williams, 1963 * '' E. (Eu.) limbatus'' Cope, 1862 * '' E. (Eu.) lucioi'' Schwartz, 1980 * '' E. (Eu.) luteolus'' Gosse, 1851 * '' E. (Eu.) maestrensis'' Díaz, Cádiz & Navarro, 2005 * '' E. (Eu.) michaelschmidi'' Díaz, Cádiz & Navarro, 2007 * '' E. (Eu.) monensis'' Meerwarth, 1901 * '' E. (Eu.) nubicola'' Dunn, 1926 * '' E. (Eu.) orcutti'' Dunn, 1928 * '' E. (Eu.) orientalis'' Barbour & Shreve, 1937 * '' E. (Eu.) oxyrhyncus'' Duméril & Bibron, 1841 * '' E. (Eu.) pantoni'' Dunn, 1926 * '' E. (Eu.) paulsoni'' Schwartz, 1964 * '' E. (Eu.) pentasyringos'' Schwartz & Fowler, 1973 * '' E. (Eu.) pezopetrus'' Schwartz, 1960 * '' E. (Eu.) pictissimus'' Cochran, 1935 * '' E. (Eu.) pinarensis'' Dunn, 1926 * '' E. (Eu.) planirostris'' Cope, 1862 * '' E. (Eu.) probolaeus'' Schwartz, 1965 * '' E. (Eu.) rhodesi'' Schwartz, 1980 * '' E. (Eu.) ricordii'' Duméril & Bibron, 1841 * '' E. (Eu.) riparius'' Estrada & Hedges, 1998 * '' E. (Eu.) rivularis'' Diaz, Estrada & Hedges, 2001 * '' E. (Eu.) rogersi'' Goin, 1955 * '' E. (Eu.) rufifemroralis'' Noble & Hassler, 1933 * '' E. (Eu.) schmidti'' Noble, 1923 * '' E. (Eu.) sciagraphus'' Schwartz, 1973 * '' E. (Eu.) semipalmatus'' Shreve, 1936 * '' E. (Eu.) simulans'' Diaz & Fong, 2001 * '' E. (Eu.) sisyphodemus'' Crombie, 1977 * '' E. (Eu.) tetajulia'' Estrada & Hedges, 1996 * '' E. (Eu.) thomasi'' Schwartz, 1959 * '' E. (Eu.) thorectes'' Hedges, 1988 * '' E. (Eu.) toa'' Estrada & Hedges, 1991 * '' E. (Eu.) tonyi'' Estrada & Hedges, 1997 * '' E. (Eu.) turquinensis'' Barbour & Shreve, 1937 * '' E. (Eu.) varleyi'' Dunn, 1925 * '' E. (Eu.) ventrilineatus'' Shreve, 1936 * '' E. (Eu.) warreni'' Schwartz, 1976 * '' E. (Eu.) weinlandi'' Barbour, 1914 * '' E. (Eu.) zugi'' Schwartz, 1958

Hispaniola Hispaniola (, also ; es, La Española; Latin and french: Hispaniola; ht, Ispayola; tnq, Ayiti or Quisqueya) is an island in the Caribbean that is part of the Greater Antilles. Hispaniola is the most populous island in the West Indies, and th ...
n (subgenus '' Pelorius'')

* '' E. (P.) cattus'' Rodriguez, Dugo-Cota, Montero-Mendieta, Gonzalez-Voyer, Alonso Bosch, Vences, and Vilà, 2017 * '' E. (P.) chlorophenax'' Schwartz, 1976 * '' E. (P.) geitonos'' Díaz, Incháustegui, Marte, Köhler, Cádiz, and Rodríguez, 2018 * '' E. (P.) hypostenor'' Schwartz, 1965 * '' E. (P.) inoptatus'' Barbour, 1914 * '' E. (P.) ligiae'' Incháustegui, Díaz, and Marte, 2015 * '' E. (P.) neiba'' Incháustegui, Díaz, and Marte, 2015 * '' E. (P.) nortoni'' Schwartz, 1976 * '' E. (P.) parapelates'' Hedges & Thomas, 1987 * '' E. (P.) ruthae'' Noble, 1923

North North is one of the four compass points or cardinal directions. It is the opposite of south and is perpendicular to east and west. ''North'' is a noun, adjective, or adverb indicating direction or geography. Etymology The word ''north ...
/
Central America Central America ( es, América Central or ) is a subregion of the Americas. Its boundaries are defined as bordering the United States to the north, Colombia to the south, the Caribbean Sea to the east, and the Pacific Ocean to the west. ...
n and
Cuba Cuba ( , ), officially the Republic of Cuba ( es, República de Cuba, links=no ), is an island country comprising the island of Cuba, as well as Isla de la Juventud and several minor archipelagos. Cuba is located where the northern Caribb ...
n (subgenus '' Syrrhopus'')

* '' E. (S.) albolabris'' Lynch & Lescure, 1980 * '' E. (S.) angustidigitorum'' Taylor, 1940 * '' Eleutherodactylus colimotl'' Grünwald, Reyes-Velasco, Franz-Chávez, Morales-Flores, Ahumada-Carrillo, Jones, and Boissinot, 2018 * '' E. (S.) cystingnathoides'' Cope, 1877 * '' E. (S.) dennisi'' Lynch, 1970 * '' E. (S.) dilatus'' Davis & Dixon, 1955 * '' E. (S.) erendirae'' Grünwald, Reyes-Velasco, Franz-Chávez, Morales-Flores, Ahumada-Carrillo, Jones, and Boissinot, 2018 * '' E. (S.) floresvillelai'' Grünwald, Reyes-Velasco, Franz-Chávez, Morales-Flores, Ahumada-Carrillo, Jones, and Boissinot, 2018 * '' E. (S.) grandis'' Dixon, 1957 * '' E. (S.) grunwaldi'' Reyes-Velasco, Ahumada-Carrillo, Burkhardt, and Devitt, 2015 * '' E. (S.) guttilatus'' Cope, 1879 * '' E. (S.) interorbitalis'' Langebartel & Shannon, 1956 * '' E. (S.) jaliscoensis'' Grünwald, Reyes-Velasco, Franz-Chávez, Morales-Flores, Ahumada-Carrillo, Jones, and Boissinot, 2018 * '' E. (S.) leprus'' Cope, 1879 * '' E. (S.) longipes'' Baird, 1859 * '' E. (S.) manantlanensis'' Grünwald, Reyes-Velasco, Franz-Chávez, Morales-Flores, Ahumada-Carrillo, Jones, and Boissinot, 2018 * '' E. (S.) marnockii'' Cope, 1878 * '' E. (S.) maurus'' Hedges, 1989 * '' E. (S.) modestus'' Taylor, 1942 * '' E. (S.) nietoi'' Grünwald, Reyes-Velasco, Franz-Chávez, Morales-Flores, Ahumada-Carrillo, Jones, and Boissinot, 2018 * '' E. (S.) nitidus'' Peters, 1870 * '' E. (S.) pallidus'' Duellman, 1958 * '' E. (S.) pipilans'' Taylor, 1940 * '' E. (S.) rubrimaculatus'' Taylor & Smith, 1945 * '' E. (S.) rufescens'' (Duellman & Dixon, 1959) * '' E. (S.) saxatilis'' Webb, 1962 * '' E. (S.) symingtoni'' Schwartz, 1957 * '' E. (S.) syristes'' Hoyt, 1965 * '' E. (S.) teretistes'' Duellman, 1958 * '' E. (S.) verrucipes'' Cope, 1885 * '' E. (S.) verruculatus'' Peters, 1870 * '' E. (S.) wixarika'' Reyes-Velasco, Ahumada-Carrillo, Burkhardt, and Devitt, 2015 * '' E. (S.) zeus'' Schwartz, 1958

References

{{Taxonbar, from=Q223286 Eleutherodactylinae Amphibian genera Taxa named by André Marie Constant Duméril Taxa named by Gabriel Bibron