''Eschrichtioides'' is an extinct
genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of
baleen whale
Baleen whales (), also known as whalebone whales, are marine mammals of the order (biology), parvorder Mysticeti in the infraorder Cetacea (whales, dolphins and porpoises), which use baleen plates (or "whalebone") in their mouths to sieve plankt ...
known from the early
Pliocene
The Pliocene ( ; also Pleiocene) is the epoch (geology), epoch in the geologic time scale that extends from 5.33 to 2.58[Northern Italy
Northern Italy (, , ) is a geographical and cultural region in the northern part of Italy. The Italian National Institute of Statistics defines the region as encompassing the four Northwest Italy, northwestern Regions of Italy, regions of Piedmo ...](_blank)
. Its type species, ''E. gastaldii'', had a complex taxonomic history, starting as a
cetothere, then as an extinct member of ''
Balaenoptera'', before being finally recognized as a relative of the
gray whale
The gray whale (''Eschrichtius robustus''), also known as the grey whale,Britannica Micro.: v. IV, p. 693. is a baleen whale that migrates between feeding and breeding grounds yearly. It reaches a length of , a weight of up to and lives between ...
.
Distribution
''Eschrichtioides'' is one of two ''
Eschrichtius'' relatives known from the
Neogene
The Neogene ( ,) is a geologic period and system that spans 20.45 million years from the end of the Paleogene Period million years ago ( Mya) to the beginning of the present Quaternary Period million years ago. It is the second period of th ...
of Italy, the other being ''
Archaeschrichtius''. Its
holotype
A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...
, MRSN 13802, comes from the early Pliocene-age Sabbie d'Asti Formation of the
Piedmont
Piedmont ( ; ; ) is one of the 20 regions of Italy, located in the northwest Italy, Northwest of the country. It borders the Liguria region to the south, the Lombardy and Emilia-Romagna regions to the east, and the Aosta Valley region to the ...
region in Italy
and it is currently exposed in
Asti
Asti ( , ; ; ) is a ''comune'' (municipality) of 74,348 inhabitants (1–1–2021) located in the Italy, Italian region of Piedmont, about east of Turin, in the plain of the Tanaro, Tanaro River. It is the capital of the province of Asti and ...
's paleontological museum "Museo Paleontologico Territoriale dell'Astigiano".
Classification

Like other cetaceans from the Italian Pliocene, the classification of ''Eschrichtioides'' was highly contentious.
described several species of ''
Cetotherium'' from the Pliocene of northern Italy, including ''C. cortesi'' based on a specimen (
MSRN 13802) found by Italian geologist
Giuseppe Cortesi in 1816 near the town
Cortandone
Cortandone is a ''comune'' (municipality) in the Province of Asti in the Italy, Italian region Piedmont, located about southeast of Turin and about northwest of Asti. As of 31 December 2019, it had a population of 308 and an area of .All demograp ...
(: paleocoordinates )
described how he, after a day of searching for fossils, found what he first thought was a small piece of wood in a small river. After a closer inspection it proved to be a bone, and Cortesi and his companion found more and more of them in the banks of the river. After four days of digging they had unveiled a giant skeleton easily identifiable as a whale 12 feet 5 inches long. The three feet long head and its dentaries had no signs of either teeth nor
tooth sockets. Cortesi noted that at that time few naturalists could assign cetacean fossils to individual species, and he therefore never named his specimen.
thought the "Baleine de Cortesi" represented a distinctive species because it was a very small adult individual and because the curvature of the maxillary branches was less convex than in any other known whale. Other French naturalists named it "Le rorqual de Cortési"; arguing that both Cortesi's fossil whales ("rorqual de Cortési" and "de Cuvier") were closely related to extant rorquals and the only difference between them was the much smaller size of the former.
assigned the "baleine de Cortesi" to his own genus ''
Cetotherium Cortesii''.
In a preliminary note pointed out differences in cranial morphology in two skulls both described as "''Cetotherium cortesii''" (a species named and described by ), and he therefore proposed the new specific name "''Cetotherium gastaldii''" (in honour of Italian palaeontologist
Bartolomeo Gastaldi) for one of the specimens.
described the specimen and reiterated his arguments from 1875.
Originally misidentified as the balaenopterid "Balaena" ''cortesii'', described "B." ''cortesii'' as a species of ''Cetotherium''.
finally named it ''C. gastaldii''. However, several authors considered it to be referrable to ''
Balaenoptera'', a taxonomic arrangement which was accepted for more than a century.
However, re-examination of MRSN 13802 revealed that "C." gastaldii was not a balaenopterid and instead constituted its own genus within
Eschrichtiidae
Eschrichtiidae or the gray whales is a family (taxonomy), family of baleen whale (Parvorder Mysticeti) with a single extant species, the gray whale (''Eschrichtius robustus''), as well as four described fossil genera: ''Archaeschrichtius'' (Mioce ...
.
References
Notes
Sources
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Plates 1, 2, 5**
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{{Taxonbar, from=Q15701238
Pliocene mammals of Europe
Pliocene cetaceans
Fossil taxa described in 2008
Eschrichtiidae
Monotypic prehistoric cetacean genera