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''Erebiola butleri'', or Butler's ringlet, is an elusive
New Zealand New Zealand () is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and List of islands of New Zealand, over 600 smaller islands. It is the List of isla ...
endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...
butterfly Butterflies are winged insects from the lepidopteran superfamily Papilionoidea, characterized by large, often brightly coloured wings that often fold together when at rest, and a conspicuous, fluttering flight. The oldest butterfly fossi ...
, discovered in 1879 by John Enys at the alpine pass at the head of the
Rakaia River The Rakaia River is in the Canterbury Plains in New Zealand's South Island. The Rakaia River is one of the largest braided rivers in New Zealand. The Rakaia River has a mean flow of and a mean annual seven-day low flow of . In the 1850s, Europ ...
. It is the only member of the genus ''Erebiola''. ''Erebiola'' is derived from
Erebus In Greek mythology, Erebus (; ), or Erebos, is the personification of darkness. In Hesiod's ''Theogony'', he is the offspring of Chaos, and the father of Aether and Hemera (Day) by Nyx (Night); in other Greek cosmogonies, he is the father of A ...
, the ancient Greek world of darkness between Earth and Hades, while the
specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
, ''butleri'', was after
Arthur Gardiner Butler Arthur Gardiner Butler F.L.S., F.Z.S. (27 June 1844 – 28 May 1925) was an English entomologist, arachnologist and ornithologist. He worked at the British Museum on the taxonomy of birds, insects, and spiders. Biography Arthur Gardiner B ...
of the
British Museum The British Museum is a Museum, public museum dedicated to human history, art and culture located in the Bloomsbury area of London. Its permanent collection of eight million works is the largest in the world. It documents the story of human cu ...
who played a major role in early descriptions of New Zealand butterflies. Its
Māori Māori or Maori can refer to: Relating to the Māori people * Māori people of New Zealand, or members of that group * Māori language, the language of the Māori people of New Zealand * Māori culture * Cook Islanders, the Māori people of the Co ...
name is ''pepe pouri'', which means dark moth, and shares the name with the black mountain ringlet and the forest ringlet butterfly.


Description

Butler's ringlet has a wingspan of 35–43 mm, with a 40 mm average for males and a 37 mm average for females. Both males and females are smoky brown, though males tend towards the richer browns while the females tend towards the paler browns. The underside of the hindwing has wedge-shaped silvery-white marks. Both the underside and the topside of the wings have eyespots at the distal-most ends, surrounded by reddish-brown shading. There is variation between individuals in the number of eyespots, the extent of the reddish-brown colouring around the eyespots, and the silvery-white markings on the undersides of the hindwings. The egg is ivory with vertical ribbing. The larvae are similar coloured from head to tail, being yellow brown with dark and light lateral striping. A fully grown larva is roughly 20 mm long. The pupa is grey and cream with fine black spotting along the abdomen, changing to a brown on the rest of the body. The style of pupation is unknown.


Life stages

The egg is laid singularly on a shrub and hatches after 14 days. The larvae grows from 3 mm to 20 mm over an unknown length of time, before pupating. Pupation lasts about 21 days. It is unknown how long adult ''E. butleri'' live for.


Distribution

Butler's ringlet is confined to the subalpine zone in the
South Island The South Island ( , 'the waters of Pounamu, Greenstone') is the largest of the three major islands of New Zealand by surface area, the others being the smaller but more populous North Island and Stewart Island. It is bordered to the north by ...
. It has been identified at only a few sites along the main divide of the
Southern Alps The Southern Alps (; officially Southern Alps / Kā Tiritiri o te Moana) are a mountain range extending along much of the length of New Zealand, New Zealand's South Island, reaching its greatest elevations near the range's western side. The n ...
. It favours subalpine terraces at altitudes of 900 to 1300 m in areas of snow-tussock with '' Hebe'' and '' Dracophyllum '' shrubs. Its preferred locations are often damp, almost boggy, or next to mountain lakes. It is very difficult to find even in areas where it has previously been seen, and so little is known about its actual range. Museum specimen records identified it as active from 27 December to 11 March. It is known to be able to remain aloft for long periods of time and to cover great distances.


Similar species

Butler's ringlet appears very similar to some species of the genus ''
Erebia ''Erebia'' is a Holarctic genus of brush-footed butterflies, family (biology), family Nymphalidae. Most of the about 90–100 species (see also #Taxonomy and systematics, below) are dark brown or black in color, with reddish-brown to orang ...
'', and was included in that genus until 1967, when ''Erebia butleri'' was reclassified as ''Erebiola butleri'' due to structural differences found between ''butleri'' and other members of the genus ''Erebia''. Butler's ringlet is visually similar to the black mountain ringlet, '' Percnodaimon pluto''. The two species may be differentiated by where the individual in question is sighted. Butler's ringlet prefers to fly over vegetation, settling among snow-tussock, subalpine shrubs and herbaceous flowers, whereas the black mountain ringlet tends to congregate over rock and scree.


See also

*
Butterflies of New Zealand Butterflies are winged insects from the lepidopteran superfamily Papilionoidea, characterized by large, often brightly coloured wings that often fold together when at rest, and a conspicuous, fluttering flight. The oldest butterfly fossil ...


References

* {{Taxonbar, from=Q1759024 Satyrini Taxa described in 1879 Monotypic butterfly genera Taxa named by Richard William Fereday