Eothyrididae
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Eothyrididae is an extinct family of very primitive, insectivorous
synapsid Synapsida is a diverse group of tetrapod vertebrates that includes all mammals and their extinct relatives. It is one of the two major clades of the group Amniota, the other being the more diverse group Sauropsida (which includes all extant rept ...
s. Only three genera are known, '' Eothyris'', '' Vaughnictis'' and '' Oedaleops'', all from the early
Permian The Permian ( ) is a geologic period and System (stratigraphy), stratigraphic system which spans 47 million years, from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.902 Mya. It is the s ...
of
North America North America is a continent in the Northern Hemisphere, Northern and Western Hemisphere, Western hemispheres. North America is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South Ameri ...
. Their main distinguishing feature is the large
caniniform In mammalian oral anatomy, the canine teeth, also called cuspids, dogteeth, eye teeth, vampire teeth, or fangs, are the relatively long, pointed tooth, teeth. In the context of the upper jaw, they are also known as ''fangs''. They can appear mo ...
tooth in front of the
maxilla In vertebrates, the maxilla (: maxillae ) is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The two maxil ...
. Eothyridids share with the
Caseidae Caseidae are an Extinction, extinct Family (biology), family of Basal (phylogenetics), basal synapsids that lived from the Late Carboniferous to Middle Permian between about 300 and 265 million years ago. Fossils of these animals come from the so ...
a number of specialised features associated with the morphology of the snout and external naris and it is likely that their common ancestor was close in build to ''Eothyris''. The two together form the clade
Caseasauria Caseasauria is one of the two main clades of early synapsids, the other being the Eupelycosauria. Caseasaurs are currently known only from the Late Carboniferous and the Permian, and include two superficially different families, the small insect ...
. ''Eothyris'' is known from a single skull specimen; ''Oedaleops'' is known from three partial skulls and some parts of some limbs. ''Vaughnictis'' is known from a partial skull, six dorsal vertebrae and some hind-limb bones. The skulls are approximately six centimeters in length, suggesting that the total length of the animals was under one meter. The species were found in the lower
Permian The Permian ( ) is a geologic period and System (stratigraphy), stratigraphic system which spans 47 million years, from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.902 Mya. It is the s ...
in what is today
North America North America is a continent in the Northern Hemisphere, Northern and Western Hemisphere, Western hemispheres. North America is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South Ameri ...
. In modern cladistics, the Eothyrididae are considered to be a basal group within the Caseasauria. Caseasauria forms a sister group of the Eupelycosauria, out of which the
therapsids Therapsida is a clade comprising a major group of eupelycosaurian synapsids that includes mammals and their ancestors and close relatives. Many of the traits today seen as unique to mammals had their origin within early therapsids, including li ...
evolved.


See also

*
Evolution of mammals The evolution of mammals has passed through many stages since the first appearance of their synapsid ancestors in the Pennsylvanian (geology), Pennsylvanian sub-period of the late Carboniferous period. By the mid-Triassic, there were many synaps ...


References

* Carroll, R. L. (1988), ''Vertebrate Paleontology and Evolution'', WH Freeman & Co. * Reisz, R. R., 1986, ''Handbuch der Paläoherpetologie – Encyclopedia of Paleoherpetology, Part 17A Pelycosauria'' Verlag Dr. Friedrich Pfeil, * Romer, AS & Price L.I (1940), Review of the Pelycosauria. ''Geol. Soc. Amer. Spec. Papers'' 28: 1–538. * Brocklehurst, N., Reisz, R. R., Fernandez, V. & Fröbisch, J. (2016) ''A Re-Description of 'Mycterosaurus' smithae, an Early Permian Eothyridid, and Its Impact on the Phylogeny of Pelycosaurian-Grade Synapsids. ''PlosOne'' 11:e0156810.


External links


Tree of LifeThe Paleobiology Database
{{Taxonbar, from=Q136601 Caseasauria Prehistoric synapsid families Taxa named by Llewellyn Ivor Price Taxa named by Alfred Romer Permian synapsids of North America