''Enteroctopus'' is an
octopus
An octopus ( : octopuses or octopodes, see below for variants) is a soft-bodied, eight- limbed mollusc of the order Octopoda (, ). The order consists of some 300 species and is grouped within the class Cephalopoda with squids, cuttlefis ...
genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...
whose members are sometimes known as giant octopus.
Etymology
The generic name ''Enteroctopus'' was created by
Alphonse Tremeau de Rochebrune and
Jules François Mabille
Jules François Mabille (Tours, 5 December 1831 − 18 January 1904) was a French malacologist, biologist and zoologist who in many trips around the world discovered and studied many species of mollusc.
In 1882−83 Mabille participated in the F ...
in 1887 and published in 1889, joining
Ancient Greek
Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic p ...
'gut' and , thus 'octopus
ith arms similar to
The Ith () is a ridge in Germany's Central Uplands which is up to 439 m high. It lies about 40 km southwest of Hanover and, at 22 kilometres, is the longest line of crags in North Germany.
Geography
Location
The Ith is immedia ...
guts.'
Description
''Enteroctopus'' is a genus of generally temperate
octopus
An octopus ( : octopuses or octopodes, see below for variants) is a soft-bodied, eight- limbed mollusc of the order Octopoda (, ). The order consists of some 300 species and is grouped within the class Cephalopoda with squids, cuttlefis ...
es. Members of this genus are characterized by their large size and are often known as giant octopuses. ''Enteroctopus'' species have distinct longitudinal wrinkles or folds dorsally and laterally on their bodies. Their heads are distinctly narrower than the mantle width. The
hectocotylus
A hectocotylus (plural: ''hectocotyli'') is one of the arms of male cephalopod
A cephalopod is any member of the molluscan class Cephalopoda ( Greek plural , ; "head-feet") such as a squid, octopus, cuttlefish, or nautilus. These exclusive ...
of the males in this genus, found on the third right arm, is long and narrow in comparison with other genera in the family
Octopodidae
The Octopodidae comprise the family containing the majority of known octopus species.
Genera
The World Register of Marine Species lists these genera:
*'' Abdopus'' Norman & Finn, 2001
*'' Ameloctopus'' Norman, 1992
*'' Amphioctopus'' P. Fischer, ...
, often comprising one-fifth the length of the arm. Octopuses in this genus have large, paddle-like
papilla
Papilla (Latin, 'nipple') or papillae may refer to:
In animals
* Papilla (fish anatomy), in the mouth of fish
* Basilar papilla, a sensory organ of lizards, amphibians and fish
* Dental papilla, in a developing tooth
* Dermal papillae, part of ...
e instead of the more conical papillae in other octopus genera.
Species
Genus ''Enteroctopus'' at present consists of four species, tabulated below:
Type species
''E. membranaceus'' has often been regarded as
type species
In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen( ...
of the genus, not because it was designated as such by
Rochebrune and
Mabille when they erected the genus, but because it was the first named species in the genus. Robson in his 1929 monograph of octopods regarded ''E. membranaceus'' as a ''
species dubium'' because the original description was insufficient to identify an individual species, the
holotype
A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of seve ...
was an immature specimen, and the type specimen no longer existed.
As such, the genus was considered invalid until Hochberg resurrected it in 1998. Hochberg noted that Robson had considered ''E. membranaceus'' a
junior synonym of ''
E. megalocyathus'', the second species assigned to the genus by Rochebrune and Mabille in their 1889 description. Additionally, since Rochebrune and Mabille did not actually assign type status to ''E. membranaceus'', Hochberg concluded that ''Enteroctopus'' was indeed a valid genus and transferred type-species status to ''E. megalocyathus'' based on his conclusion that ''E. megalocyathus'' and ''E. membranaceus'' are the
same species.
[
]
Distribution
Species in the genus ''Enteroctopus'' are restricted to the temperate areas of the Northern and Southern Hemispheres. ''E. dofleini'' is the only member of the genus found in the Northern Hemisphere, and also the most widely distributed: It is found from Southern California
Southern California (commonly shortened to SoCal) is a geographic and cultural region that generally comprises the southern portion of the U.S. state of California. It includes the Los Angeles metropolitan area, the second most populous urban ...
, along the North Pacific Rim to Japan, including the Okhotsk
Okhotsk ( rus, Охотск, p=ɐˈxotsk) is an urban locality (a work settlement) and the administrative center of Okhotsky District of Khabarovsk Krai, Russia, located at the mouth of the Okhota River on the Sea of Okhotsk. Population:
Hi ...
and Bering Sea
The Bering Sea (, ; rus, Бе́рингово мо́ре, r=Béringovo móre) is a marginal sea of the Northern Pacific Ocean. It forms, along with the Bering Strait, the divide between the two largest landmasses on Earth: Eurasia and The Amer ...
s.[ The other three species are found in the Southern Hemisphere; ''E. megalocyathus'' occurs on the southeastern coast of ]South America
South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the souther ...
, ''E. magnificus'' on the southwestern coast of Africa
Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...
from Namibia
Namibia (, ), officially the Republic of Namibia, is a country in Southern Africa. Its western border is the Atlantic Ocean. It shares land borders with Zambia and Angola to the north, Botswana to the east and South Africa to the south and e ...
to Port Elizabeth
Gqeberha (), formerly Port Elizabeth and colloquially often referred to as P.E., is a major seaport and the most populous city in the Eastern Cape province of South Africa. It is the seat of the Nelson Mandela Bay Metropolitan Municipality, S ...
, South Africa
South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring count ...
, and ''E. zealandicus'' in temperate New Zealand
New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 List of islands of New Zealand, smaller islands. It is the ...
.
Size
The member of this genus that best embodies the common name "giant octopus" is ''Enteroctopus dofleini
The giant Pacific octopus (''Enteroctopus dofleini''), also known as the North Pacific giant octopus, is a large marine cephalopod belonging to the genus ''Enteroctopus''. Its spatial distribution includes the coastal North Pacific, along Mexico ...
'', which holds the record of being the world's largest octopus based on direct measurements of a individual, weighed live. This octopus had a total length near to . The remaining members of the genus are substantially smaller, with ''E. megalocyathus'' having an average mass of and reaching a total length of . ''E. magnificus'' reaches a total length of around .[
]
See also
*Gigantic octopus
The kraken () is a legendary sea monster of enormous size said to appear off the coasts of Norway.
Kraken, the subject of sailors' superstitions and mythos, was first described in the modern age at the turn of the 18th century, in a travel ...
*Giant squid
The giant squid (''Architeuthis dux'') is a species of deep-ocean dwelling squid in the family (biology), family Architeuthidae. It can grow to a tremendous size, offering an example of deep-sea gigantism, abyssal gigantism: recent estimates ...
*Colossal squid
The colossal squid (''Mesonychoteuthis hamiltoni'') is part of the family Cranchiidae. It is sometimes called the Antarctic squid or giant cranch squid and is believed to be the largest squid species in terms of mass. It is the only recognize ...
*Cephalopod size
Cephalopods vary enormously in size. The smallest are only about long and weigh less than at maturity, while the largest—the giant and colossal squids—can exceed in length and weigh close to half a tonne (), making them the largest livin ...
*Kraken
The kraken () is a legendary sea monster of enormous size said to appear off the coasts of Norway.
Kraken, the subject of sailors' superstitions and mythos, was first described in the modern age at the turn of the 18th century, in a travel ...
*Lusca
In Caribbean folklore, the Lusca is a name given to a sea monster said to exist in the region of the blue holes nearby Andros, an island in the Bahamas.
Description
It is described as a giant octopus, a giant cuttlefish, or a half dragon, half oc ...
Footnotes
References
{{Authority control
Enteroctopodidae
Taxa named by Jules François Mabille