The emperor dragonfly
or blue emperor
(''Anax imperator'') is a large species of hawker
dragonfly
A dragonfly is a flying insect belonging to the infraorder Anisoptera below the order Odonata. About 3,000 extant species of dragonflies are known. Most are tropical, with fewer species in temperate regions. Loss of wetland habitat threat ...
of the family
Aeshnidae
Aeshnidae, also called aeshnids, hawkers, or darners, is a family of dragonflies, found nearly worldwide, with more than 50 genera and over 450 species.
The family includes some of the largest dragonflies.
Description
Common worldwide or nearl ...
. It is the largest dragonfly in most of Europe, including the United Kingdom,
although exceeded in some areas by other species.
Nomenclature
The generic name ''Anax'' is from the ancient Greek , "lord";
the specific epithet ''imperator'' is the Latin for "emperor", from ''imperare'', to command.
Distribution
This dragonfly has a wide distribution through
Afroeurasia; it is found throughout Africa and through most of Europe, the
Arabian Peninsula
The Arabian Peninsula (, , or , , ) or Arabia, is a peninsula in West Asia, situated north-east of Africa on the Arabian plate. At , comparable in size to India, the Arabian Peninsula is the largest peninsula in the world.
Geographically, the ...
, and south-western and central Asia.
Since the 1990s, its range has expanded in Europe, both northwards and to higher altitudes. For example, the first Scandinavian record was in 1994 in Denmark; in 2002 it was first recorded in Sweden and in 2004 first in Scotland; today it is regular in all three countries.
The species' northward expansion has been tied to
global warming
Present-day climate change includes both global warming—the ongoing increase in global average temperature—and its wider effects on Earth's climate system. Climate change in a broader sense also includes previous long-term changes ...
, and it is among the first
odonata
Odonata is an order of predatory flying insects that includes the dragonflies and damselflies (as well as the '' Epiophlebia'' damsel-dragonflies). The two major groups are distinguished with dragonflies (Anisoptera) usually being bulkier with ...
to do so.
Identification
The emperor dragonfly is a large and bulky species. It is long, with average being and males growing larger than females.
The average wingspan is .
[ When they first emerge, both sexes appear pale green with brown markings. The legs are brown with a yellow like base. Wings are born black but grow yellow-brown when they grow. Males have a bright sky blue or turquoise abdomen marked with a diagnostic black dorsal stripe.] However, their blue colour may be faded during cold weather spells. The thorax and head of a male is apple green and their prominent eyes are blue. Females have similar markings but they are mainly a duller green.[ As the females age, their wings become browner. Less immediately visible features for both sexes are the yellow costa and brown spots on the wings.] Emperor dragonflies can also be recognised by their flight patterns: they often fly with their abdomen hanging slightly downwards.
One of the largest species in Europe, the emperor dragonfly is exceeded by magnificent emperor, which occurs only marginally in the east Mediterranean and in length by females of the golden-ringed dragonfly, a species with an unusually long ovipositor
The ovipositor is a tube-like organ used by some animals, especially insects, for the laying of eggs. In insects, an ovipositor consists of a maximum of three pairs of appendages. The details and morphology of the ovipositor vary, but typica ...
. Thus, in most of Europe the emperor is the largest dragonfly species present.
File:Ai(loz)hydro.JPG, Male in side view
File:Anax imperator 2015 11 23 6807 (cropped).jpg, Male in flight
File:Anax imperator female.jpg, Female laying eggs
File:Anax imperator Exuvie MHNT Parc de la Maourine.jpg, Exuviae
In biology, exuviae are the remains of an exoskeleton and related structures that are left after ecdysozoans (including insects, crustaceans and arachnids) have molted. The exuviae of an animal can be important to biologists as they can often be ...
File:Anax Imperator 2(loz).JPG, Emerging
Behaviour
They frequently fly high up into the sky in search of prey, which includes butterflies
Butterflies are winged insects from the lepidopteran superfamily Papilionoidea, characterized by large, often brightly coloured wings that often fold together when at rest, and a conspicuous, fluttering flight. The oldest butterfly fossi ...
, other odonata
Odonata is an order of predatory flying insects that includes the dragonflies and damselflies (as well as the '' Epiophlebia'' damsel-dragonflies). The two major groups are distinguished with dragonflies (Anisoptera) usually being bulkier with ...
and tadpoles. If their hunt is successful, they eat their smaller prey while flying. The dragonflies breed in a variety of aquatic habitats from large ponds to dikes and slow-moving rivers, but require a plentiful supply of vegetation in the water. They do sometimes breed in brackish water. The females lay the eggs into plants such as pondweed
Pondweed refers to many species and genera of aquatic plants and green algae:
*''Potamogeton'', a diverse and worldwide genus
*''Elodea'', found in North America
*''Aponogeton'', in Africa, Asia and Australasia
*''Groenlandia'', a genus of aquatic ...
, and always lay alone. The aquatic larvae are very aggressive and are likely to influence the native species composition
Relative species abundance is a component of biodiversity and is a measure of how common or rare a species is relative to other species in a defined location or community.Hubbell, S. P. 2001. ''The unified neutral theory of biodiversity and biogeog ...
of freshwater ecosystems they arrive in. The larvae are also very large–around . The adult male is highly territorial, and difficult to approach.
Conservation
Emperor dragonflies are assessed as a least-concern species
A least-concern species is a species that has been evaluated and categorized by the International Union for Conservation of Nature (IUCN) as not being a focus of wildlife conservation because the specific species is still plentiful in the wil ...
by the IUCN. The species has widespread and has a stable population.
Mitochrondrial genome
The mitogenome of the emperor dragonfly is the longest of all known dragonfly sequences. It has 16,087 base pairs
A base pair (bp) is a fundamental unit of double-stranded nucleic acids consisting of two nucleobases bound to each other by hydrogen bonds. They form the building blocks of the DNA double helix and contribute to the folded structure of both DNA ...
. For comparison, the human mitogenome has 16,569 and the closely related dragonfly '' Anax parthenope'' has 15,366.
References
External links
Blue Emperor
Species text in the Online Atlas of South African Odonata
*
{{Taxonbar, from=Q527496
Aeshnidae
Odonata of Africa
Odonata of Asia
Dragonflies of Europe
Insects described in 1815
Taxa named by William Elford Leach