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''Durio graveolens,'' sometimes called the red-fleshed durian, orange-fleshed durian, or yellow durian, is a species of tree in the family
Malvaceae Malvaceae, or the mallows, is a family of flowering plants estimated to contain 244 genera with 4225 known species. Well-known members of economic importance include okra, cotton, cacao and durian. There are also some genera containing familia ...
. It is one of six species of
durian The durian (, ) is the edible fruit of several tree species belonging to the genus ''Durio''. There are 30 recognised ''Durio'' species, at least nine of which produce edible fruit. '' Durio zibethinus'', native to Borneo and Sumatra, is the on ...
named by Italian naturalist
Odoardo Beccari Odoardo Beccari (16 November 1843 – 25 October 1920) was an Italian botanist famous for his discoveries in Indonesia, particularly New Guinea, and Australia. He has been called the greatest botanist to ever study Malesia. His author abbre ...
. The
specific epithet In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bo ...
('strong smelling' or 'rank') is due to the odor. Although most species of ''
Durio The durian (, ) is the edible fruit of several tree species belonging to the genus ''Durio''. There are 30 recognised ''Durio'' species, at least nine of which produce edible fruit. ''Durio zibethinus'', native to Borneo and Sumatra, is the onl ...
'' (most notably ''
Durio dulcis ''Durio dulcis'', known as durian marangang (or merangang), red durian, tutong, or lahung, is a fairly large tree in the genus '' Durio''. It can grow up to 40 m tall. The husk of its fruit is dark red to brown-red, and covered with slender ...
'') have a strong scent, the red-fleshed type of ''D. graveolens'' has a mild scent. It is native to Southeast Asia. ''D. graveolens'' is an edible durian, perhaps the most popular 'wild' species of durian, and it is sold commercially regionally. However, its congener ''
Durio zibethinus ''Durio zibethinus'' is the most common tree species in the genus '' Durio'' that are known as durian and have edible fruit also known as durian. As with most other durian species, the edible flesh emits a distinctive odour that is strong and pe ...
'' is the typical species eaten and dominates sales worldwide.


Names

This species should not be confused with the popular durian clones from Malaysia known as 'Red Flesh' (D164) and 'Red Prawn' (D175), as both of those belong to ''D. zibethinus''. However, ''D. graveolens'' does have one registered variety, 'DQ2 nyekak (DK8)'. The color of the fruit's flesh denotes other varieties–an orange-fleshed, a red-fleshed one, and yellow-fleshed. These varieties may be different species, but currently there is no consensus. The yellow-fleshed kind is sometimes called ''durian simpor''. In the scientific name ''Durio graveolens'', '' graveolens'' means 'strong smelling', although it has been described as having a "mild" or "slight" odor or even, in a book published by the US National Research Council, as "odorless". In Malay, the fruit is called ', ' (literally "bird durian"), ' ("jungle durian"), ' ("yellow durian"), ' ("red durian"), or ' ("crayfish brain durian"). In
Iban IBAN or Iban or Ibán may refer to: Banking * International Bank Account Number Ethnology * Iban culture * Iban language * Iban people Given name Cycling * Iban Iriondo (born 1984) * Iban Mayo (born 1977) * Iban Mayoz (born 1981) Football * ...
, it is '. The
Bidayuh Bidayuh is the collective name for several indigenous groups found in southern Sarawak, Malaysia and northern West Kalimantan, Indonesia, on the island of Borneo, which are broadly similar in language and culture (see also issues below). T ...
call it ''durian umot''. Among the
Kenyah The Kenyah people are an indigenous, Austronesian-speaking people of Borneo, living in the remote Baram Lio Matoh, Long Selaan, Long Moh, Long Anap, Long Mekaba, Long Jeeh, Long Belaong, Long San, Long Silat, Long Tungan, Data Kakus ...
and
Dayak people The Dayak (; older spelling: Dajak) or Dyak or Dayuh are one of the native groups of Borneo. It is a loose term for over 200 riverine and hill-dwelling ethnic groups, located principally in the central and southern interior of Borneo, each w ...
s, it is known as ''durian anggang'' ("hornbill durian"), ''durian ajan'', ''pesang'', ''tabela'' or ''ta-bela'', ''tabelak'', ''taula'', ''tuala'', ''tuwala''. On
Sumatra Sumatra is one of the Sunda Islands of western Indonesia. It is the largest island that is fully within Indonesian territory, as well as the sixth-largest island in the world at 473,481 km2 (182,812 mi.2), not including adjacent i ...
, the
Batak Batak is a collective term used to identify a number of closely related Austronesian ethnic groups predominantly found in North Sumatra, Indonesia, who speak Batak languages. The term is used to include the Karo, Pakpak, Simalungun, Tob ...
call it ''tinambela''. In
Karo Karo may refer to: Ethnic groups * Karo people (East Africa), a group of tribes in East Africa * Karo people (Ethiopia), an ethnic group from Ethiopia * Karo people (Indonesia), the indigenous people of the Karo Plateau in North Sumatra Languag ...
, it is called '. In
Thailand Thailand ( ), historically known as Siam () and officially the Kingdom of Thailand, is a country in Southeast Asia, located at the centre of the Indochinese Peninsula, spanning , with a population of almost 70 million. The country is b ...
, it is referred to as ( th, ทุเรียนรากขา) or (; sources differ on which name refers to this species, with the other being attributed to ''D. kutejensis''). In
Aceh Tamiang Regency Aceh Tamiang Regency ( id, Kabupaten Aceh Tamiang) is a regency in the east of Aceh province of Indonesia, bordering on North Sumatra Province to the east. It is located on the island of Sumatra. The regency covers an area of 1,957.02 square k ...
, it can be called ''durian batu'' ("stone durian"), and elsewhere in Sumatra, it known as ''durian adjan''. Other regional names include ''durian dalit'' (but this can apply also to ''
Durio oxleyanus ''Durio oxleyanus'' is a perennial plant species of tree in the family Malvaceae. It was once placed in the family Bombacaceae. The IUCN list the species as near threatened. It is a popular plant for food among humans and orangutans. Nam ...
'') ''alau'', ''dujen'', ''durian alau'', ''durian daun dungoh'', ''durian hutan'' ("forest durian"), ''durian pipit'', ''lai bengang'', ''merang kunyit'', ''pasang'', and ''tongkai''. A natural hybrid of ''D. graveolens'' and ''D. zibethinus'' is called ''durian siunggong'' or ''durian suluk''. It has the texture and flavour of the popular ''D. zibethinus'' and the burnt
caramel Caramel ( or ) is an orange-brown confectionery product made by heating a range of sugars. It can be used as a flavoring in puddings and desserts, as a filling in bonbons, or as a topping for ice cream and custard. The process of carameli ...
overtones of ''D. graveolens''. After its initial description in 1889 by
Odoardo Beccari Odoardo Beccari (16 November 1843 – 25 October 1920) was an Italian botanist famous for his discoveries in Indonesia, particularly New Guinea, and Australia. He has been called the greatest botanist to ever study Malesia. His author abbre ...
, in 1924, Dutch botanist Reinier Cornelis Bakhuizen Van Den Brink reduced it to a
synonym A synonym is a word, morpheme, or phrase that means exactly or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are al ...
of '' D. conicus''. British botanist
John Wyatt-Smith John is a common English name and surname: * John (given name) * John (surname) John may also refer to: New Testament Works * Gospel of John, a title often shortened to John * First Epistle of John, often shortened to 1 John * Seco ...
combined it all under ''D. dulcis'' in 1953. Indonesian botanists
André Joseph Guillaume Henri Kostermans Dr. André Joseph Guillaume Henri 'Dok' Kostermans (Purworejo, 1 July 1906 – Jakarta, 10 July 1994) was an Indonesian botanist of Dutch ancestry. He was born in Purworejo, Java, Dutch East Indies, and educated at Utrecht University, taking h ...
and Wertit Soegeng-Reksodihardjo separated ''D. graveolens'' back to its own species in 1958.


Distribution

Wild ''D. graveolens'' grows in
Peninsular Malaysia Peninsular Malaysia ( ms, Semenanjung Malaysia; Jawi: سمننجڠ مليسيا), or the States of Malaya ( ms, Negeri-negeri Tanah Melayu; Jawi: نڬري-نڬري تانه ملايو), also known as West Malaysia or the Malaysian Peninsula, ...
(states of
Johor Johor (; ), also spelled as Johore, is a state of Malaysia in the south of the Malay Peninsula. Johor has land borders with the Malaysian states of Pahang to the north and Malacca and Negeri Sembilan to the northwest. Johor shares mariti ...
,
Kedah Kedah (), also known by its honorific Darul Aman and historically as Queda, is a state of Malaysia, located in the northwestern part of Peninsular Malaysia. The state covers a total area of over 9,000 km2, and it consists of the mainland ...
,
Kelantan Kelantan (; Jawi: ; Kelantanese Malay: ''Klate'') is a state in Malaysia. The capital is Kota Bharu and royal seat is Kubang Kerian. The honorific name of the state is ''Darul Naim'' (Jawi: ; "The Blissful Abode"). Kelantan is located in t ...
,
Malacca Malacca ( ms, Melaka) is a States and federal territories of Malaysia, state in Malaysia located in the southern region of the Malay Peninsula, next to the Strait of Malacca. Its capital is Malacca City, dubbed the Historic City, which has bee ...
,
Penang Penang ( ms, Pulau Pinang, is a Malaysian state located on the northwest coast of Peninsular Malaysia, by the Malacca Strait. It has two parts: Penang Island, where the capital city, George Town, is located, and Seberang Perai on the M ...
,
Perak Perak () is a state of Malaysia on the west coast of the Malay Peninsula. Perak has land borders with the Malaysian states of Kedah to the north, Penang to the northwest, Kelantan and Pahang to the east, and Selangor to the south. Thailand' ...
,
Selangor Selangor (; ), also known by its Arabic honorific Darul Ehsan, or "Abode of Sincerity", is one of the 13 Malaysian states. It is on the west coast of Peninsular Malaysia and is bordered by Perak to the north, Pahang to the east, Negeri Sem ...
, and
Terengganu Terengganu (; Terengganu Malay: ''Tranung'', Jawi: ), formerly spelled Trengganu or Tringganu, is a sultanate and constitutive state of federal Malaysia. The state is also known by its Arabic honorific, ''Dāru l- Īmān'' ("Abode of Faith") ...
),
Borneo Borneo (; id, Kalimantan) is the third-largest island in the world and the largest in Asia. At the geographic centre of Maritime Southeast Asia, in relation to major Indonesian islands, it is located north of Java, west of Sulawesi, and e ...
,
Sumatra Sumatra is one of the Sunda Islands of western Indonesia. It is the largest island that is fully within Indonesian territory, as well as the sixth-largest island in the world at 473,481 km2 (182,812 mi.2), not including adjacent i ...
,
Palawan Palawan (), officially the Province of Palawan ( cyo, Probinsya i'ang Palawan; tl, Lalawigan ng Palawan), is an archipelagic province of the Philippines that is located in the region of Mimaropa. It is the largest province in the country in t ...
, and
Southern Thailand Southern Thailand, Southern Siam or Tambralinga is a southernmost cultural region of Thailand, separated from Central Thailand region by the Kra Isthmus. Geography Southern Thailand is on the Malay Peninsula, with an area of around , bounde ...
. It is cultivated in
Brunei Brunei ( , ), formally Brunei Darussalam ( ms, Negara Brunei Darussalam, Jawi: , ), is a country located on the north coast of the island of Borneo in Southeast Asia. Apart from its South China Sea coast, it is completely surrounded by th ...
,
Sarawak Sarawak (; ) is a state of Malaysia. The largest among the 13 states, with an area almost equal to that of Peninsular Malaysia, Sarawak is located in northwest Borneo Island, and is bordered by the Malaysian state of Sabah to the northeast, ...
,
Sabah Sabah () is a state of Malaysia located in northern Borneo, in the region of East Malaysia. Sabah borders the Malaysian state of Sarawak to the southwest and the North Kalimantan province of Indonesia to the south. The Federal Territory ...
, and the
Northern Territory The Northern Territory (commonly abbreviated as NT; formally the Northern Territory of Australia) is an Australian territory in the central and central northern regions of Australia. The Northern Territory shares its borders with Western Aust ...
of Australia. In Brunei, its popularity outshines ''D. zibethinus'', which is not cultivated in the country. It is occasionally grown outside the tropics. In
Florida Florida is a state located in the Southeastern region of the United States. Florida is bordered to the west by the Gulf of Mexico, to the northwest by Alabama, to the north by Georgia, to the east by the Bahamas and Atlantic Ocean, and ...
, it has been seen to survive two consecutive nights at , albeit shrouded in cloth.


Description

''D. graveolens'' is a large tree, sharing many features with '' D. dulcis''. It inhabits the upper
canopy Canopy may refer to: Plants * Canopy (biology), aboveground portion of plant community or crop (including forests) * Canopy (grape), aboveground portion of grapes Religion and ceremonies * Baldachin or canopy of state, typically placed over an ...
, growing up to tall. The trunk is in diameter and can have no branches until about high. The trunk will be smooth or flaky, grey/
mauve Mauve (, ; , ) is a pale purple color named after the mallow flower (French: ''mauve''). The first use of the word ''mauve'' as a color was in 1796–98 according to the ''Oxford English Dictionary'', but its use seems to have been rare befo ...
to ruddy brown with steep
buttress root Buttress roots also known as plank roots are large, wide roots on all sides of a shallowly rooted tree. Typically, they are found in nutrient-poor tropical forest soils that may not be very deep. They prevent the tree from falling over (hen ...
s. The buttresses reach and extend out . The oblong leaves are long without the petiole (leaf stalk), and wide. They are perfectly rounded on both ends, rigid, and slightly
coriaceous This glossary of botanical terms is a list of definitions of terms and concepts relevant to botany and plants in general. Terms of plant morphology are included here as well as at the more specific Glossary of plant morphology and Glossary o ...
(leather-like in feel or texture). On the top, they are
glabrous Glabrousness (from the Latin '' glaber'' meaning "bald", "hairless", "shaved", "smooth") is the technical term for a lack of hair, down, setae, trichomes or other such covering. A glabrous surface may be a natural characteristic of all or part of ...
(smooth and hairless) and crisp, almost vernicose (varnished). Underneath, the leaves are copper-brown and lepidote (scaly), with large scales of up to in diameter, which are not very noticeable, at least when dry. The leaf scales are
peltate The following is a list of terms which are used to describe leaf morphology in the description and taxonomy of plants. Leaves may be simple (a single leaf blade or lamina) or compound (with several leaflets). The edge of the leaf may be regular ...
(shield-shaped),
ciliate The ciliates are a group of alveolates characterized by the presence of hair-like organelles called cilia, which are identical in structure to eukaryotic flagella, but are in general shorter and present in much larger numbers, with a differen ...
-radiated (fringed), and deeply-lobed in three to five parts. In addition to the scales, long strands of stellate hairs and other
trichome Trichomes (); ) are fine outgrowths or appendages on plants, algae, lichens, and certain protists. They are of diverse structure and function. Examples are hairs, glandular hairs, scales, and papillae. A covering of any kind of hair on a p ...
s of varying size form a soft
tomentose Trichomes (); ) are fine outgrowths or appendages on plants, algae, lichens, and certain protists. They are of diverse structure and function. Examples are hairs, glandular hairs, scales, and papillae. A covering of any kind of hair on a pl ...
(fuzzy) surface. The leaf
midrib This glossary of botanical terms is a list of definitions of terms and concepts relevant to botany and plants in general. Terms of plant morphology are included here as well as at the more specific Glossary of plant morphology and Glossary o ...
is very prominent on the underside and forms a crease on top. The leaf
stipule In botany, a stipule is an outgrowth typically borne on both sides (sometimes on just one side) of the base of a leafstalk (the petiole). Stipules are considered part of the anatomy of the leaf of a typical flowering plant, although in many speci ...
s are
caducous Dehiscence is the splitting of a mature plant structure along a built-in line of weakness to release its contents. This is common among fruits, anthers and sporangia. Sometimes this involves the complete detachment of a part; structures that op ...
(drop early). Leaves have 10-12 lateral veins per side (with some smaller ones intermixed), which are tiny and superficial above and more distinct, but still barely visible. The petiole is very large, long, and tumescent (swollen) from the middle up. Flowers grow on the branches on short cymes and a thin
calyx Calyx or calyce (plural "calyces"), from the Latin ''calix'' which itself comes from the Ancient Greek ''κάλυξ'' (''kálux'') meaning "husk" or "pod", may refer to: Biology * Calyx (anatomy), collective name for several cup-like structures ...
. The base is sac-like with three to five connate lobes. It has white,
spatulate The following is a list of terms which are used to describe leaf morphology in the description and taxonomy of plants. Leaves may be simple (a single leaf blade or lamina) or compound (with several leaflets). The edge of the leaf may be regular ...
(spoon-shaped) petals that are long. Inside are five separate bundles of
staminode In botany, a staminode is an often rudimentary, sterile or abortive stamen, which means that it does not produce pollen.Jackson, Benjamin, Daydon; ''A Glossary of Botanic Terms with their Derivation and Accent''; Published by Gerald Duckworth & C ...
s and
stamen The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the fila ...
s, fused for less than half of their length. The
anther The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filam ...
has small clusters of four or five elongated
pollen Pollen is a powdery substance produced by seed plants. It consists of pollen grains (highly reduced microgametophytes), which produce male gametes (sperm cells). Pollen grains have a hard coat made of sporopollenin that protects the gametop ...
locule A locule (plural locules) or loculus (plural loculi) (meaning "little place" in Latin) is a small cavity or compartment within an organ or part of an organism (animal, plant, or fungus). In angiosperms (flowering plants), the term ''locule'' usu ...
s that open with longitudinal slits.
Ovaries The ovary is an organ in the female reproductive system that produces an ovum. When released, this travels down the fallopian tube into the uterus, where it may become fertilized by a sperm. There is an ovary () found on each side of the body. T ...
are
ovoid An oval () is a closed curve in a plane which resembles the outline of an egg. The term is not very specific, but in some areas ( projective geometry, technical drawing, etc.) it is given a more precise definition, which may include either o ...
to
globose A sphere () is a geometrical object that is a three-dimensional analogue to a two-dimensional circle. A sphere is the set of points that are all at the same distance from a given point in three-dimensional space.. That given point is the c ...
(roughly spherical) and possess a yellow
capitate The capitate bone is a bone in the human wrist found in the center of the carpal bone region, located at the distal end of the radius and ulna bones. It articulates with the third metacarpal bone (the middle finger) and forms the third carpomet ...
(shaped like a pinhead) stigma and white to greenish
style Style is a manner of doing or presenting things and may refer to: * Architectural style, the features that make a building or structure historically identifiable * Design, the process of creating something * Fashion, a prevailing mode of clothing ...
about . The pollen is
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