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The Dingy Skipper (''Erynnis tages'') is a
butterfly Butterflies are insects in the macrolepidopteran clade Rhopalocera from the order Lepidoptera, which also includes moths. Adult butterflies have large, often brightly coloured wings, and conspicuous, fluttering flight. The group compris ...
of the family
Hesperiidae Skippers are a family of the Lepidoptera (moths and butterflies) named the Hesperiidae. Being diurnal, they are generally called butterflies. They were previously placed in a separate superfamily, Hesperioidea; however, the most recent taxonomy ...
.


Description

''Erynnis tages'' is different from other skippers because of the predominantly monochrome, gray-brown wing coloration and the marbling, which is only present on the upper side of the forewings, as well as a series of small white dots on the wing edge. It is probably the most moth-like British butterfly and normally rests with its wings in a moth-like fashion. This well-camouflaged, brown and grey butterfly can be confused with the
grizzled skipper ''Pyrgus'' is a genus in the skippers butterfly family, Hesperiidae, known as the grizzled skippers. The name "checkered" or "chequered skipper" may also be applied to some species, but also refers to species in the genera '' Burnsius'' and '' ...
, the
Mother Shipton moth ''Callistege mi'', the Mother Shipton moth, is a moth of the family Erebidae. It was classified by Carl Alexander Clerck in 1759, and is also known under the name of ''Euclidia mi''. In Finnish it is known as ''piirtoyökkönen'' and in German ...
or the burnet companion moth. Faded specimens of '' Carcharodus alceae'' are distinguished by glass spots in the forewing and by a jagged rear wing edge. A special feature is the restraint of the imagos, which is reminiscent of an deltoid moth with its roof-shaped wings placed one on top of the other.


Range

The Dingy Skipper is widespread from the
Iberian Peninsula The Iberian Peninsula (), ** * Aragonese and Occitan: ''Peninsula Iberica'' ** ** * french: Péninsule Ibérique * mwl, Península Eibérica * eu, Iberiar penintsula also known as Iberia, is a peninsula in southwestern Europe, def ...
and
Ireland Ireland ( ; ga, Éire ; Ulster-Scots: ) is an island in the North Atlantic Ocean, in north-western Europe. It is separated from Great Britain to its east by the North Channel, the Irish Sea, and St George's Channel. Ireland is the s ...
in the west to the Pacific in East Asia. In the north, the area boundary follows roughly the 62°N. In Asia, the species penetrates south across the
Caucasus The Caucasus () or Caucasia (), is a region between the Black Sea and the Caspian Sea, mainly comprising Armenia, Azerbaijan, Georgia, and parts of Southern Russia. The Caucasus Mountains, including the Greater Caucasus range, have historica ...
to
Pakistan Pakistan ( ur, ), officially the Islamic Republic of Pakistan ( ur, , label=none), is a country in South Asia. It is the world's List of countries and dependencies by population, fifth-most populous country, with a population of almost 24 ...
, and in
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's List of countries and dependencies by population, most populous country, with a Population of China, population exceeding 1.4 billion, slig ...
the area boundary is far to the south. In
Bavaria Bavaria ( ; ), officially the Free State of Bavaria (german: Freistaat Bayern, link=no ), is a state in the south-east of Germany. With an area of , Bavaria is the largest German state by land area, comprising roughly a fifth of the total l ...
, the distribution shows a noticeable change from more or less densely populated areas to regions in which the species is absent over long stretches. In southern Bavaria, the species is concentrated in the Alpine region, the valleys of the Alpine rivers, in particular the Lech and
Isar The Isar is a river in Tyrol, Austria, and Bavaria, Germany, which is not navigable for watercraft above raft size. Its source is in the Karwendel range of the Alps in Tyrol; it enters Germany near Mittenwald and flows through Bad Tölz, Mu ...
, as well as the areas near the river in the Donaumoos. In Northern Bavaria, the focus is on the Franconian Alb, in parts of the Franconian Keuper-Lias-Land and the Mainfränkische Platten. A secondary focus in the Upper Main hill country leads to scattered evidence along the northeastern border. Here the species occurs from the
Vogtland Vogtland (; cz, Fojtsko) is a region spanning the German states of Bavaria, Saxony and Thuringia and north-western Bohemia in the Czech Republic. It overlaps with and is largely contained within Euregio Egrensis. The name alludes to the forme ...
over the
Fichtelgebirge The Fichtel MountainsRandlesome, C. et al. (2011). ''Business Cultures in Europe'', 2nd ed., Routledge, Abingdon and New York, p. 52. . (german: Fichtelgebirge, cs, Smrčiny), form a small horseshoe-shaped mountain range in northeastern Bavaria ...
to the
Upper Palatinate The Upper Palatinate (german: Oberpfalz, , ) is one of the seven administrative districts of Bavaria, Germany, and is located in the east of Bavaria. Geography The Upper Palatinate is a landscape with low mountains and numerous ponds and lak ...
and
Bavarian Forest The village of Zell in the Bavarian Forest The Bavarian Forest (German: ' or ''Bayerwald''; bar, Boarischa Woid) is a wooded, low-mountain region in Bavaria, Germany that is about 100 kilometres long. It runs along the Czech border and is co ...
. With the exception of the East Bavarian low mountain range, the occurrence of the Dingy Skipper coincides with the distribution of limestone areas (Jura and Muschelkalk) or geological features (basalt, gypsum, basic sandstones such as dolomitic arkose). It is widely but patchily distributed across Britain. It occurs further north than any other skipper in
Scotland Scotland (, ) is a country that is part of the United Kingdom. Covering the northern third of the island of Great Britain, mainland Scotland has a border with England to the southeast and is otherwise surrounded by the Atlantic Ocean to ...
with some isolated colonies in the
Inverness Inverness (; from the gd, Inbhir Nis , meaning "Mouth of the River Ness"; sco, Innerness) is a city in the Scottish Highlands. It is the administrative centre for The Highland Council and is regarded as the capital of the Highlands. Histor ...
region. It is also one of the two skippers to be found in Ireland, again with a patchy distribution but the main strongholds along the western side. It is on the decline in several European countries including the UK and
Armenia Armenia (), , group=pron officially the Republic of Armenia,, is a landlocked country in the Armenian Highlands of Western Asia.The UNbr>classification of world regions places Armenia in Western Asia; the CIA World Factbook , , and ''O ...
.Butterfly Conservation Armenia
/ref>


Habitat

''Erynnis tages'' favours open grassy habitats up to 2,000 metres above sea level. A variety of habitats are used including
chalk downland Downland, chalkland, chalk downs or just downs are areas of open chalk hills, such as the North Downs. This term is used to describe the characteristic landscape in southern England where chalk is exposed at the surface. The name "downs" is deriv ...
, woodland clearings, coastal dunes, railway lines and waste ground. The habitats of the Dingy Skipper are mainly dry and poor grasslands. Extensive grassland with one or two-tier meadows and pastures as well as habitats with little vegetation with raw soil and initial plant communities are also regularly populated. In forests, the imagos fly in very sparse wooded stands or on forest meadows, on the edges of paths and forest edges. The species is also found in fens. Suitable habitats are disturbances such as paths with a dry, warm microclimate.


Ecology

The imagos fly in two generations from May-June and July-August but in northern regions and at the high altitudes, there is only a single generation. The flight period in Bavaria extends from the middle of April to the beginning of September with a focus from the beginning of May to the end of June and a maximum in the last May decade. First and second generation overlap. The second generation, which occurs only irregularly or regionally, is always significantly weaker. The imagos prefer low-growing or very patchy vegetation and often stay on bare ground to sunbathe or to absorb moisture and minerals. On the ground or on low plants, the males also move from perch in order to track down the females by approaching insects of suitable size. The flight is usually very fast and low, whereby the locally loyal imagos usually only cover short distances of a few meters and soon settle again. There is very little information on flower visits. In Bavaria out of a total of 14 listed plant species, only three are mentioned several times, in addition to the most important egg-laying plants ''
Hippocrepis comosa ''Hippocrepis comosa'', the horseshoe vetch, is a species of perennial flowering plant belonging to the genus ''Hippocrepis'' in the family Fabaceae. Description The overall appearance depends on its habitat: sometimes it forms upright clumps ...
'' and ''
Lotus corniculatus ''Lotus corniculatus'' is a flowering plant in the pea family Fabaceae, native to grasslands in temperate Eurasia and North Africa. Common names include common bird's-foot trefoil, eggs and bacon, birdsfoot deervetch, and just bird's-foot trefoi ...
'', ''
Ajuga reptans ''Ajuga reptans'' is commonly known as bugle, blue bugle, bugleherb, bugleweed, carpetweed, carpet bugleweed, and common bugle, and traditionally but less commonly as St. Lawrence plant. It is an herbaceous flowering plant in the mint family Lami ...
'' also is visited. Other suction plants are yellow-flowered
Fabaceae The Fabaceae or Leguminosae,International Code of Nomenc ...
such as ''
Anthyllis vulneraria ''Anthyllis vulneraria'', the common kidneyvetch, kidney vetch or woundwort is a medicinal plant native to Europe. The name ''vulneraria'' means "wound healer". Description ''Anthyllis vulneraria'' reaches of height. The stem is simple or more ...
,
Chamaecytisus ratisbonensis ''Chamaecytisus'' is a genus of flowering plants in the legume family, Fabaceae. It belongs to the subfamily Faboideae. It may be a synonym of '' Cytisus''. '' C. palmensis'' is a tree from the Canary Islands used as a fodder crop around the worl ...
'' and ''
Medicago falcata ''Medicago falcata'' is a plant species of the genus '' Medicago''. It is native to much of Europe and Asia, but is found throughout the world. It forms a symbiotic relationship with the bacterium ''Sinorhizobium meliloti'', which is capable of n ...
'' as well as blue- and violet-flowered mint flowers such as ''
Salvia pratensis ''Salvia pratensis'', the meadow clary or meadow sage, is a species of flowering plant in the family Lamiaceae, native to Europe, western Asia and northern Africa. The Latin specific epithet ''pratensis'' means "of meadows", referring to its ...
'' and ''
Thymus The thymus is a specialized primary lymphoid organ of the immune system. Within the thymus, thymus cell lymphocytes or '' T cells'' mature. T cells are critical to the adaptive immune system, where the body adapts to specific foreign invaders ...
spp.'' Visits to flowers on taller perennials such as ''
Cirsium arvense ''Cirsium arvense'' is a perennial species of flowering plant in the family Asteraceae, native throughout Europe and western Asia, northern Africa and widely introduced elsewhere.Joint Nature Conservation Committee''Cirsium arvense'' The standa ...
'', ''
Eupatorium cannabinum ''Eupatorium cannabinum'', commonly known as hemp-agrimony, or holy rope, is a herbaceous plant in the family Asteraceae. It is a robust perennial native to many areas of Europe. The alkaloids may be present in the plant material as their N-oxide ...
'' or ''
Echium vulgare ''Echium vulgare'', known as viper's bugloss and blueweed,Dickinson, T.; Metsger, D.; Bull, J.; & Dickinson, R. (2004) ROM Field Guide to Wildflowers of Ontario. Toronto:Royal Ontario Museum, p. 203. is a species of flowering plant in the bora ...
'' are rare, as the imagos mainly stay near the ground. In addition to visiting flowers, the imagos regularly suckle on moist soil, carrion and excrement. Eggs are laid individually on the upper side of mostly terminal leaflets near the ground, with gaps or weak-growing, microclimatically favored locations being preferred. Observations of egg laying or of egg and caterpillar finds are only available in isolated cases. The following egg-laying and host plants have been reported from Bavaria: ''
Hippocrepis comosa ''Hippocrepis comosa'', the horseshoe vetch, is a species of perennial flowering plant belonging to the genus ''Hippocrepis'' in the family Fabaceae. Description The overall appearance depends on its habitat: sometimes it forms upright clumps ...
,
Lotus corniculatus ''Lotus corniculatus'' is a flowering plant in the pea family Fabaceae, native to grasslands in temperate Eurasia and North Africa. Common names include common bird's-foot trefoil, eggs and bacon, birdsfoot deervetch, and just bird's-foot trefoi ...
,
Securigera varia ''Securigera varia'' ( synonym ''Coronilla varia''), commonly known as crownvetch or purple crown vetch, is a low-growing legume vine. It is native to Africa, Asia and Europe and is commonly used throughout the United States and Canada for e ...
, Tetragonalobus maritimus''. Almost all reports relate to observations of oviposition or egg discoveries, only a single detection of an adult caterpillar was found in ''Lotus corniculatus''. In Great Britain eggs were found on the tender young leaves of bird's-foot trefoil (''Lotus corniculatus''), the favoured food plant here (although
horseshoe vetch ''Hippocrepis comosa'', the horseshoe vetch, is a species of perennial flowering plant belonging to the genus ''Hippocrepis'' in the family Fabaceae. Description The overall appearance depends on its habitat: sometimes it forms upright clumps ...
(''Hippocrepis comosa'') and greater bird's-foot trefoil (''Lotus pendunculatus'') are sometimes used). Other larval host plants in Europe are ''
Eryngium ''Eryngium'' is a genus of flowering plants in the family Apiaceae. There are about 250 species.Wang, P., et al. (2012)Phytochemical constituents and pharmacological activities of ''Eryngium'' L. (Apiaceae). ''Pharmaceutical Crops'' 3 99-120. The ...
'', ''
Coronilla The genus ''Coronilla'' contains 8 species of flowering plants native to Europe and North Africa. It cointained about 20 species before being split into ''Securigera''. Species include: * ''Coronilla coronata'' * ''Coronilla juncea'' * ''Coron ...
'', ''
Medicago ''Medicago'' is a genus of flowering plants, commonly known as medick or burclover, in the legume family (Fabaceae). It contains at least 87 species and is distributed mainly around the Mediterranean basin. The best-known member of the genus is ...
'', etc. The hemispherical egg with clear longitudinal ribs is initially light yellow and later orange-red in color and is easily recognizable on the green upper side of the leaf with a targeted search. This combination of features results in a further, reliable detection method for the species. During the day, the caterpillar hides in a hiding place made of spun leaves and mainly eats at night. The
caterpillar Caterpillars ( ) are the larval stage of members of the order Lepidoptera (the insect order comprising butterflies and moths). As with most common names, the application of the word is arbitrary, since the larvae of sawflies (suborder Sy ...
creates a shelter by spinning leaves together and feeds until fully grown in August. It then creates a larger tent to form a hibernaculum where it hibernates.
Pupa A pupa ( la, pupa, "doll"; plural: ''pupae'') is the life stage of some insects undergoing transformation between immature and mature stages. Insects that go through a pupal stage are holometabolous: they go through four distinct stages in thei ...
tion occurs the following spring without further feeding. The winter web is dense like parchment, so that no water can penetrate. Spun threads in summer, the pupae of which result in second generation butterflies, are of a loose texture.


Conservation

In accordance with its preference for low and gappy vegetation, grazing is particularly suitable for maintaining the habitats, which, in addition to the continuous shortening of the vegetation, also results in the formation of the open ground areas that are essential for the species due to the footfall. Extensive grazing with sheep or cattle must therefore be continued on lime lawns, the most important type of habitat in central Europe. If grazing is not possible on poor grass, mowing can also be considered. There are no restrictions with regard to the time of mowing, as the pre-imaginal stages almost always live close to the ground. For the occurrence in poor meadows, the extensive renouncement of fertilization is necessary for survival. Another contribution to the protection of the species is the preservation of small structures with patchy vegetation on fields, embankments and roadsides. It is essential to avoid the use of suction mowers or mulchers when caring for such structures. The creation of new habitats is also possible in sub-areas of the occurrence, for example on limestone gravel in the valleys of southern Bavaria, since the imagos often colonize anthropogenic secondary sites. Suitable areas are e.g. flood dams, embankments or disused mining sites, which are maintained in a low-growing and gappy state by largely dispensing with planting and corresponding subsequent care.


Etymology

"A grandson of Jupiter"


Taxonomy

Subspecies are little defined and include ''E. t. unicolor'' Freyer, 1852 found in
Transcaucasia The South Caucasus, also known as Transcaucasia or the Transcaucasus, is a geographical region on the border of Eastern Europe and Western Asia, straddling the southern Caucasus Mountains. The South Caucasus roughly corresponds to modern Arme ...
. ''Synonyms: Erynnis morio'' SCOPOLI, 1763; ''Erynnis geryon'' ROTTEMBURG, 1775


See also

*
List of butterflies of Great Britain This is a list of butterflies of Great Britain, including extinct, naturalised species and those of dubious origin. The list comprises butterfly species listed in ''The Moths and Butterflies of Great Britain and Ireland'' by Emmet ''et al.'' and '' ...


References


External links


Butterfly ConservationUK ButterfliesButterfly Conservation Armenia
{{Taxonbar, from=Q867291 Erynnis Butterflies of Asia Butterflies of Europe Butterflies described in 1758 Taxa named by Carl Linnaeus