Dartford warbler
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The Dartford warbler (''Curruca undata'') is a typical warbler from the warmer parts of western
Europe Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...
and northwestern
Africa Africa is the world's second-largest and second-most populous continent after Asia. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 20% of Earth's land area and 6% of its total surfac ...
. It is a small warbler with a long thin tail and a thin pointed bill. The adult male has grey-brown upperparts and is dull reddish-brown below except for the centre of the belly which has a dirty white patch. It has light speckles on the throat and a red eye-ring. The sexes are similar but the adult female is usually less grey above and paler below. Its breeding range lies west of a line from southern
England England is a Countries of the United Kingdom, country that is part of the United Kingdom. It is located on the island of Great Britain, of which it covers about 62%, and List of islands of England, more than 100 smaller adjacent islands. It ...
to the heel of
Italy Italy, officially the Italian Republic, is a country in Southern Europe, Southern and Western Europe, Western Europe. It consists of Italian Peninsula, a peninsula that extends into the Mediterranean Sea, with the Alps on its northern land b ...
(southern
Apulia Apulia ( ), also known by its Italian language, Italian name Puglia (), is a Regions of Italy, region of Italy, located in the Southern Italy, southern peninsular section of the country, bordering the Adriatic Sea to the east, the Strait of Ot ...
). The Dartford warbler is usually resident all year in its breeding range, but there is some limited
migration Migration, migratory, or migrate may refer to: Human migration * Human migration, physical movement by humans from one region to another ** International migration, when peoples cross state boundaries and stay in the host state for some minimum le ...
.


Taxonomy and systematics

The Dartford warbler was first described in 1776 by the Welsh naturalist,
Thomas Pennant Thomas Pennant (16 December 1798) was a Welsh natural history, naturalist, traveller, writer and antiquarian. He was born and lived his whole life at his family estate, Downing Hall, near Whitford, Flintshire, in Wales. As a naturalist he had ...
. He introduced the English name and based his description on two specimens that had been obtained by the ornithologist John Latham from Bexley Heath, near
Dartford Dartford is the principal town in the Borough of Dartford, Kent, England. It is located south-east of Central London and is situated adjacent to the London Borough of Bexley to its west. To its north, across the Thames Estuary, is Thurrock in ...
in
Kent Kent is a Ceremonial counties of England, ceremonial county in South East England. It is bordered by Essex across the Thames Estuary to the north, the Strait of Dover to the south-east, East Sussex to the south-west, Surrey to the west, and Gr ...
. In 1783 Latham included the warbler in his ''A General Synopsis of Birds'' but did not coin the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin grammatical forms, altho ...
, ''Sylvia dartfordiensis'', until the supplement to his work was published in 1787. However, in 1783, before the publication of Latham's supplement, the Dutch naturalist
Pieter Boddaert Pieter Boddaert (1730 – 6 May 1795) was a Dutch physician and naturalist. Early life, family and education Boddaert was the son of a Middelburg jurist and poet by the same name (1694–1760). The younger Pieter obtained his M.D. at the Univers ...
introduced the name, ''Motacilla undata'', based on a coloured plate of "Le Pitte-chou, de Provence" in Edmé-Louis Daubenton's ''Planches enluminées d'histoire naturelle''. The specific epithet ''undata'' is from
Medieval Latin Medieval Latin was the form of Literary Latin used in Roman Catholic Church, Roman Catholic Western Europe during the Middle Ages. It was also the administrative language in the former Western Roman Empire, Roman Provinces of Mauretania, Numidi ...
''undatus'' meaning "with wavy markings". The type locality is
Provence Provence is a geographical region and historical province of southeastern France, which stretches from the left bank of the lower Rhône to the west to the France–Italy border, Italian border to the east; it is bordered by the Mediterrane ...
in France. This species probably forms a
superspecies In biology, a species complex is a group of closely related organisms that are so similar in appearance and other features that the boundaries between them are often unclear. The taxa in the complex may be able to hybridize readily with each oth ...
with Tristram's warbler and this in turn seems close to Marmora's warbler and the Balearic warbler. Altogether, this group of typical warblers bears a resemblance to the
wrentit The wrentit (''Chamaea fasciata'') is a small bird that lives in chaparral, California oak woodland, oak woodlands, and bushland on the western coast of North America. It is the Monotypic taxon, only species in the genus ''Chamaea''. Its systema ...
, the only species of
Sylviidae Sylviidae is a family of passerine birds that includes the typical warblers ''Sylvia'', and closely related genus '' Curruca'', formerly included in ''Sylvia''. They are found in Eurasia and Africa, with the greatest diversity in the Mediterrane ...
from the
Americas The Americas, sometimes collectively called America, are a landmass comprising the totality of North America and South America.''Webster's New World College Dictionary'', 2010 by Wiley Publishing, Inc., Cleveland, Ohio. When viewed as a sing ...
. However, the wrentit is less closely related to the genus ''Sylvia'' than to the
parrotbill The parrotbills are a family, Paradoxornithidae, of passerine birds that are primarily native to East, Southeast and South Asia, with a single species in western North America, though feral populations exist elsewhere. They are generally small ...
s. Its visual similarity to the Dartford warbler group is an example of
convergent evolution Convergent evolution is the independent evolution of similar features in species of different periods or epochs in time. Convergent evolution creates analogous structures that have similar form or function but were not present in the last comm ...
. Three
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
are recognised: * ''C. u. dartfordiensis'' ( Latham, 1787) – south England and north-west France * ''C. u. toni'' (
Hartert Ernst Johann Otto Hartert (29 October 1859 – 11 November 1933) was a widely published German people, German ornithologist. Life and career Hartert was born in the Free and Hanseatic City of Hamburg on 29 October 1859. In July 1891, he married ...
, 1909) – north-west Africa * ''C. u. undata'' ( Boddaert, 1783) –
Iberian Peninsula The Iberian Peninsula ( ), also known as Iberia, is a peninsula in south-western Europe. Mostly separated from the rest of the European landmass by the Pyrenees, it includes the territories of peninsular Spain and Continental Portugal, comprisin ...
and south France


Description

The Dartford warbler is a small, ,
passerine A passerine () is any bird of the order Passeriformes (; from Latin 'sparrow' and '-shaped') which includes more than half of all bird species. Sometimes known as perching birds, passerines generally have an anisodactyl arrangement of their ...
bird, distinguished by its long tail compared with that of other warblers. Its plumage comprises unobtrusive and muted tones, which blend in with the dry dead plants, old wood or sunny greyish wood found in its preferred habitats. Like many typical warblers, the Dartford warbler has distinct male and female
plumage Plumage () is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, there can b ...
s. The male has a grey back and head, reddish underparts, and a red eye. The reddish throat is spotted with white. The sides are a dull greyish tone, being more clear about the abdomen. In some populations males have bluish-grey or brownish-grey backs and heads. The female is paler below, especially on the throat, and a browner grey above. The female's throat also has white spots, although they are smaller and less marked than in the male. Juvenile birds are similar to females.


Distribution and habitat

The species is naturally rare. The largest European populations of ''Curruca undata'' are in the Iberian peninsula, others in much of
France France, officially the French Republic, is a country located primarily in Western Europe. Overseas France, Its overseas regions and territories include French Guiana in South America, Saint Pierre and Miquelon in the Atlantic Ocean#North Atlan ...
, in
Italy Italy, officially the Italian Republic, is a country in Southern Europe, Southern and Western Europe, Western Europe. It consists of Italian Peninsula, a peninsula that extends into the Mediterranean Sea, with the Alps on its northern land b ...
and southern
England England is a Countries of the United Kingdom, country that is part of the United Kingdom. It is located on the island of Great Britain, of which it covers about 62%, and List of islands of England, more than 100 smaller adjacent islands. It ...
and south
Wales Wales ( ) is a Countries of the United Kingdom, country that is part of the United Kingdom. It is bordered by the Irish Sea to the north and west, England to the England–Wales border, east, the Bristol Channel to the south, and the Celtic ...
. In
Africa Africa is the world's second-largest and second-most populous continent after Asia. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 20% of Earth's land area and 6% of its total surfac ...
it can be found only in small areas in the north, wintering in northern
Morocco Morocco, officially the Kingdom of Morocco, is a country in the Maghreb region of North Africa. It has coastlines on the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to Algeria–Morocc ...
and northern
Algeria Algeria, officially the People's Democratic Republic of Algeria, is a country in the Maghreb region of North Africa. It is bordered to Algeria–Tunisia border, the northeast by Tunisia; to Algeria–Libya border, the east by Libya; to Alger ...
.


Behaviour and ecology


Breeding

Dartford warblers first breed when they are one year old. They are usually monogamous and the
pair bond In biology, a pair bond is the strong affinity that develops in some species between a mating pair, often leading to the production and rearing of young and potentially a lifelong bond. Pair-bonding is a term coined in the 1940s that is frequently ...
can persist from one year to the next. The male sometimes builds several simple nests (cock nests) of which one is chosen by the female, but it is more usual for both birds to participate in constructing the breeding nest. In southern England the birds breed on heathlands, sometimes near the coast, and nest in either common gorse (''Ulex europaeus'') or common heather (''Calluna vulgaris''). The compact cup-shaped nest is located in dense bushes, usually within of the ground. It is formed mainly of grasses and is lined with a layer of finer material that can include thin roots and feathers. The eggs are laid from early April in southern France and Spain, and from mid-April in southern England. The
clutch A clutch is a mechanical device that allows an output shaft to be disconnected from a rotating input shaft. The clutch's input shaft is typically attached to a motor, while the clutch's output shaft is connected to the mechanism that does th ...
is typically 3–5 eggs which are smooth and glossy, with a white or occasionally pale green ground and marked with brown speckles which are sometimes concentrated at the larger end. The average size of an egg is with a weight of of which 6 percent is shell. The eggs are incubated for 12–14 days mainly by the female. The chicks are fed
arthropod Arthropods ( ) are invertebrates in the phylum Arthropoda. They possess an arthropod exoskeleton, exoskeleton with a cuticle made of chitin, often Mineralization (biology), mineralised with calcium carbonate, a body with differentiated (Metam ...
s by both parents. The nestlings
fledge Fledging is the stage in a flying animal's life between egg, hatching or birth and becoming capable of flight. This term is most frequently applied to birds, but is also used for bats. For altricial birds, those that spend more time in vulnera ...
10–14 days after hatching and are then fed by their parents for a further two weeks. Usually two and occasionally three broods are raised in a year.


Feeding

It inhabits open fields with degraded scrub brush and is common in heather. In winter it may visit urban areas, but always feeds within shrubs in these areas. It nests in bushes with thorns and near the ground. These warblers are mostly
insectivorous A robber fly eating a hoverfly An insectivore is a carnivorous animal or plant which eats insects. An alternative term is entomophage, which can also refer to the human practice of eating insects. The first vertebrate insectivores we ...
, eating
caterpillar Caterpillars ( ) are the larval stage of members of the order Lepidoptera (the insect order comprising butterflies and moths). As with most common names, the application of the word is arbitrary, since the larvae of sawflies (suborder ...
s,
butterflies Butterflies are winged insects from the lepidopteran superfamily Papilionoidea, characterized by large, often brightly coloured wings that often fold together when at rest, and a conspicuous, fluttering flight. The oldest butterfly fossi ...
,
beetle Beetles are insects that form the Taxonomic rank, order Coleoptera (), in the superorder Holometabola. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 40 ...
s and
spider Spiders (order (biology), order Araneae) are air-breathing arthropods that have eight limbs, chelicerae with fangs generally able to inject venom, and spinnerets that extrude spider silk, silk. They are the largest order of arachnids and ran ...
s. The song of the Dartford warbler is a distinctive rattling warble.


Status and conservation

Dartford warblers almost died out in the
United Kingdom The United Kingdom of Great Britain and Northern Ireland, commonly known as the United Kingdom (UK) or Britain, is a country in Northwestern Europe, off the coast of European mainland, the continental mainland. It comprises England, Scotlan ...
in the severe winter of 1962/1963 when the national population dropped to just ten pairs. ''Curruca undata'' is also sensitive to drought affecting breeding success or producing heath fires, as occurred during 1975 and 1976 in England when virtually all juveniles failed to survive their first year.Bibby, Colin J.
‘Dartford Warblers in England’
''British Birds'', 72 (1979); pp. 10–22
However, this species can recover well in good quality habitat with favourable temperatures and rainfall, thanks to repeated nesting and a high survival rate for the young. Indeed, they recovered in some areas of the UK, but numbers are once again on the decline in other regions of their natural range. The range of the Dartford warbler is restricted to western and southern Europe. The total population in 2012 was estimated at 1.1–2.5 million breeding pairs. The largest numbers occur in Spain where there were believed to be 983,000–1,750,000 pairs. For reasons that probably include loss of suitable habitat, the Spanish population appears to be declining. The species is therefore classed by the
International Union for Conservation of Nature The International Union for Conservation of Nature (IUCN) is an international organization working in the field of nature conservation and sustainable use of natural resources. Founded in 1948, IUCN has become the global authority on the stat ...
as being
Near Threatened A near-threatened species is a species which has been Conservation status, categorized as "Near Threatened" (NT) by the International Union for Conservation of Nature (IUCN) as that may be vulnerable to Endangered species, endangerment in the ne ...
. A period of climatic warming since 1963 has seen the UK population increase to "more than 2,500 pairs in 2006 (Wotton et al. 2009). Expansion into patches of structurally suitable habitat (up to an altitude of 400m), more northerly areas and away from the core of the range, from Dorset and Hampshire to Derbyshire and Suffolk, is likely to have been facilitated by milder winter weather (Wotton et al. 2009, Bradbury et al. 2011). The Dartford warbler population in the UK is expected to continue to increase. However, future climate-based projections for the European range indicate that by 2080, more than 60% of the current European range may no longer be suitable (Huntley et al 2007). There is evidence that this is happening already, with severe declines in Spain and France (Green 2017). For this reason, the species is classified as Near Threatened on the IUCN Global Red List. If the declines in southern Europe continue, the UK will become increasingly important for global conservation of this species".


References


Sources

*
BirdLife International BirdLife International is a global partnership of non-governmental organizations that strives to conserve birds and their habitats. BirdLife International's priorities include preventing extinction of bird species, identifying and safeguarding i ...
(BLI) (2008)
2008 IUCN Redlist status changes
Retrieved 23 May 2008 *


External links


Ageing and sexing (PDF; 3.1 MB) by Javier Blasco-Zumeta & Gerd-Michael Heinze

Photos and videos from the Internet Bird Collection
* *
Vocalisations on Xeno-canto
{{Taxonbar, from=Q110257524, from2=Q752545
Dartford warbler The Dartford warbler (''Curruca undata'') is a typical warbler from the warmer parts of western Europe and northwestern Africa. It is a small warbler with a long thin tail and a thin pointed bill. The adult male has grey-brown upperparts and is d ...
Birds of Europe Birds of North Africa
Dartford warbler The Dartford warbler (''Curruca undata'') is a typical warbler from the warmer parts of western Europe and northwestern Africa. It is a small warbler with a long thin tail and a thin pointed bill. The adult male has grey-brown upperparts and is d ...