
A subshrub (
Latin
Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...
''suffrutex'') or undershrub is either a small
shrub
A shrub or bush is a small to medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees by their multiple ...
(e.g.
prostrate shrubs) or a
perennial
In horticulture, the term perennial ('' per-'' + '' -ennial'', "through the year") is used to differentiate a plant from shorter-lived annuals and biennials. It has thus been defined as a plant that lives more than 2 years. The term is also ...
that is largely
herbaceous
Herbaceous plants are vascular plants that have no persistent woody stems above ground. This broad category of plants includes many perennials, and nearly all annuals and biennials.
Definitions of "herb" and "herbaceous"
The fourth edition of ...
but slightly
woody at the base (e.g.
garden pink and
florist's chrysanthemum). The term is often interchangeable with "bush".
[Jackson, Benjamin, Daydon; A Glossary of Botanic Terms with their Derivation and Accent; Published by Gerald Duckworth & Co. London, 4th ed 1928]
Because the criteria are matters of degree (normally of height) rather than of kind, the definition of a subshrub is not sharply distinguishable from that of a shrub; examples of reasons for describing plants as subshrubs include ground-hugging stems or low growth habit. Subshrubs may be largely herbaceous though still classified as woody, with overwintering perennial woody growth much lower-growing than deciduous summer growth. Some plants described as subshrubs are only weakly woody and some persist for only a few years. Others, such as ''
Oldenburgia paradoxa'' live indefinitely (though is still vulnerable to external effects), rooted in rocky cracks.
Small, low shrubs such as
lavender
''Lavandula'' (common name lavender) is a genus of 47 known species of perennial flowering plants in the sage family, Lamiaceae. It is native plant, native to the Old World, primarily found across the drier, warmer regions of the Mediterranean ...
,
periwinkle, and
thyme
Thyme () is a culinary herb consisting of the dried aerial parts of some members of the genus ''Thymus (plant), Thymus'' of flowering plants in the mint family Lamiaceae. Thymes are native to Eurasia and north Africa. Thymes have culinary, medici ...
, and many shrub-like members of the family
Ericaceae
The Ericaceae () are a Family (biology), family of flowering plants, commonly known as the heath or heather family, found most commonly in acidic and infertile growing conditions. The family is large, with about 4,250 known species spread acros ...
, such as
cranberries and small species of ''
Erica
Erica or ERICA may refer to:
* Erica (given name)
* Erica (plant), ''Erica'' (plant), a flowering plant genus
* Erica (chatbot), a service of Bank of America
* Erica (video game), ''Erica'' (video game), a 2019 FMV video game
* Erica (spider), ' ...
'', are often classed as subshrubs.
Definition
A chamaephyte, subshrub or dwarf-shrub is a plant that bears hibernating buds on persistent shoots near the ground – usually woody plants with
perennating buds borne close to the ground, usually less than above the soil surface. The significance of the closeness to the ground is that the buds remain within the soil surface layer and are thus somewhat protected from various adverse external influences. Accordingly, the chamaephyte
habit
A habit (or wont, as a humorous and formal term) is a routine of behavior that is repeated regularly and tends to occur subconsciously.
A 1903 paper in the '' American Journal of Psychology'' defined a "habit, from the standpoint of psychology, ...
is especially common in stressful environments, for example:
* ecosystems on nutrient-poor soils or
rock
Rock most often refers to:
* Rock (geology), a naturally occurring solid aggregate of minerals or mineraloids
* Rock music, a genre of popular music
Rock or Rocks may also refer to:
Places United Kingdom
* Rock, Caerphilly, a location in Wale ...
* exposed alpine or arctic ecosystems where seasonal or perennial wind and freezing conditions are prone to kill vulnerable growing shoots
* ecosystems subject to frequent fires and burning, where many species of e.g. ''
Banksia
''Banksia'' is a genus of around 170 species of flowering plants in the family Proteaceae. These Australian wildflowers and popular garden plants are easily recognised by their characteristic flower spikes, and woody fruiting "cones" and head ...
'' or ''
Eucalyptus
''Eucalyptus'' () is a genus of more than 700 species of flowering plants in the family Myrtaceae. Most species of ''Eucalyptus'' are trees, often Mallee (habit), mallees, and a few are shrubs. Along with several other genera in the tribe Eucalyp ...
'' regrow from a
lignotuber
A lignotuber is a woody swelling of the root crown possessed by some plants as a protection against destruction of the plant stem, such as by fire. Other woody plants may develop basal burls as a similar survival strategy, often as a respons ...
or
caudex
A caudex (: caudices) of a plant is a stem, but the term is also used to mean a rootstock and particularly a basal stem structure from which new growth arises.pages 456 and 695
In the strict sense of the term, meaning a stem, "caudex" is most ...
.
* heavily grazed or overgrazed ecosystems, such as
tortoise turf
Examples of chamaephytes
The term chamaephyte is most formally used within the context of
Raunkiær plant life-forms' classification. Examples of chamaephytes are many of the species living in the
maquis shrubland
220px, Low maquis in Corsica
220px, High ''macchia'' in Sardinia
( , , ) or ( , ; often in Italian; , ; ; ; ) is a savanna-like shrubland biome in the Mediterranean region, typically consisting of densely growing evergreen shrubs.
Maquis ...
and other plants of
submediterranean dry ecosystems (species such as thyme, ''
Thymus vulgaris
''Thymus vulgaris'' (common thyme, German thyme, garden thyme or just thyme) is a species of flowering plant in the mint family (biology), family Lamiaceae, native plant, native to southern Europe from the western Mediterranean to southern Ital ...
'', and rosemary, ''
Salvia rosmarinus''); others include heather species (e.g. ''
Calluna vulgaris
''Calluna vulgaris'', common heather, ling, or simply heather, is the sole species in the genus ''Calluna'' in the flowering plant family Ericaceae. It is a low-growing evergreen shrub growing to tall, or rarely to and taller, and is found wide ...
'' and ''
Erica
Erica or ERICA may refer to:
* Erica (given name)
* Erica (plant), ''Erica'' (plant), a flowering plant genus
* Erica (chatbot), a service of Bank of America
* Erica (video game), ''Erica'' (video game), a 2019 FMV video game
* Erica (spider), ' ...
s''), African wild olive (''
Olea europaea'' ssp. ''cuspidata'') and edelweiss (''
Leontopodium alpinum''). Chamaephytes also include
cushion plant
A cushion plant is a compact, low-growing, mat-forming plant that is found in alpine, subalpine, arctic, or subarctic environments around the world. The term "cushion" is usually applied to woody plants that grow as spreading mats, are limited i ...
s.
See also
*
Raunkiær plant life-form
The Raunkiær system is a system for categorizing plants using life-form categories, devised by Danish botanist Christen C. Raunkiær and later extended by various authors.
History
It was first proposed in a talk to the ''Danish Botanical Soci ...
*
Shrub
A shrub or bush is a small to medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees by their multiple ...
*
Prostrate shrub
*
Woody plant
A woody plant is a plant that produces wood as its structural tissue and thus has a hard stem. In cold climates, woody plants further survive winter or dry season above ground, as opposed to Herbaceous plant, herbaceous plants that die back to t ...
References
{{botany
Plant life-forms
Plant morphology
Mediterranean forests, woodlands, and scrub
Plants by habit