Dromiacea is a group of

crab Crabs are decapod crustaceans of the infraorder Brachyura (meaning "short tailed" in Greek language, Greek), which typically have a very short projecting tail-like abdomen#Arthropoda, abdomen, usually hidden entirely under the Thorax (arthropo ...

s, ranked as a section. It contains 240 extant and nearly 300 extinct species. Dromiacea is the most basal grouping of Brachyura crabs, diverging the earliest in the evolutionary history, around the

Late Triassic The Late Triassic is the third and final epoch (geology), epoch of the Triassic geologic time scale, Period in the geologic time scale, spanning the time between annum, Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch a ...

Early Jurassic The Early Jurassic Epoch (geology), Epoch (in chronostratigraphy corresponding to the Lower Jurassic series (stratigraphy), Series) is the earliest of three epochs of the Jurassic Period. The Early Jurassic starts immediately after the Triassic� ...

. Below is a

cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not s ...

showing Dromiacea's placement within Brachyura: The

larva A larva (; : larvae ) is a distinct juvenile form many animals undergo before metamorphosis into their next life stage. Animals with indirect development such as insects, some arachnids, amphibians, or cnidarians typically have a larval phase ...

e of Dromiacea resemble those of the

Anomura Anomura (sometimes Anomala) is a group of decapod crustaceans, including hermit crabs and others. Although the names of many anomurans include the word ''crab'', all true crabs are in the sister group to the Anomura, the Brachyura (the two groups ...

more closely than those of other

s. This may simply reflect their basal position in the crab

phylogeny A phylogenetic tree or phylogeny is a graphical representation which shows the evolutionary history between a set of species or Taxon, taxa during a specific time.Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, M ...

. The superfamily Eocarcinoidea, containing '' Eocarcinus'' and '' Platykotta'', was previously considered to be a member of the Dromiacea, but has since been transferred to the

. The

fossil record A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...

of Dromiacea reaches back at least as far as the

Jurassic The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately 143.1 Mya. ...

, and, if '' Imocaris'' is indeed a member, into the

Carboniferous The Carboniferous ( ) is a Geologic time scale, geologic period and System (stratigraphy), system of the Paleozoic era (geology), era that spans 60 million years, from the end of the Devonian Period Ma (million years ago) to the beginning of the ...

. Dromiacea primarily consists of two groups of superfamilies - Dromioidea and Homoloidea. See the below

: Recent studies have found that some of the

families Family (from ) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictability, structure, and safety as ...

may not be

monophyletic In biological cladistics for the classification of organisms, monophyly is the condition of a taxonomic grouping being a clade – that is, a grouping of organisms which meets these criteria: # the grouping contains its own most recent co ...

, but rather

paraphyletic Paraphyly is a taxonomic term describing a grouping that consists of the grouping's last common ancestor and some but not all of its descendant lineages. The grouping is said to be paraphyletic ''with respect to'' the excluded subgroups. In co ...

. *The Dromioidea family Dromiidae may be paraphyletic with respect to Dynomenidae *The Homoloidea family

Homolidae The family Homolidae, known as carrier crabs or porter crabs, contains 14 genera of marine crabs. They mostly live on the continental slope and continental shelf, and are rarely encountered. Members of the Homolidae have their fifth pereiopod ...

is paraphyletic with respect to

Latreilliidae Latreilliidae is a small family of crabs. They are relatively small, long-legged crabs found on soft bottoms at depths of up in mostly tropical and subtemperate waters around the world. Their carapace is very small and doesn’t cover the bas ...

References

Crabs Extant Jurassic first appearances Taxa named by Wilhem de Haan {{Dromiacea-stub