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''Dendrobium speciosum'', commonly known as the rock orchid or cane orchid, is a species of highly variable Australian
orchid Orchids are plants that belong to the family Orchidaceae (), a diverse and widespread group of flowering plants with blooms that are often colourful and fragrant. Orchids are cosmopolitan plants that are found in almost every habitat on Eart ...
. Its varieties can be found in a range of
habitat In ecology, habitat refers to the array of resources, biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species' habitat can be seen as the physical manifestation of its ...
s as
epiphyte An epiphyte is a plant or plant-like organism that grows on the surface of another plant and derives its moisture and nutrients from the air, rain, water (in marine environments) or from debris accumulating around it. The plants on which epiphyt ...
s (on branches or trunks of trees) or lithophytes. It has a continuous distribution along the east coast of Australia and in distinct populations along the Tropic of Capricorn. As a lithophyte, it forms gigantic spreading colonies on rocks and cliff faces, often exposed to full sun, with its roots forming dense, matted beds across the rock that anchor the plant. It can be found at altitudes from sea level to .


Description

''Dendrobium speciosum'' is an epiphytic or lithophytic herb with spreading roots and cylindrical or tapered
pseudobulb In botany, a pseudobulb is a storage organ found in many epiphytic and terrestrial sympodial orchids. It is derived from a thickening of the part of a stem between leaf nodes and may be composed of just one internode or several, termed ''heter ...
s long and wide. Each pseudobulb has up to seven, usually thick, leathery leaves originating from its top, the leaves long and wide. The leaves can remain on the plant for up to twelve years. The flowers vary in colour from white to bright yellows and there is considerable variation in the length of the flowering
raceme A raceme () or racemoid is an unbranched, indeterminate growth, indeterminate type of inflorescence bearing flowers having short floral stalks along the shoots that bear the flowers. The oldest flowers grow close to the base and new flowers are ...
, the number of flowers on it and the size of the flowers. The length of the flowering stem ranges from long and the number of flowers from two to two hundred. The flowers are long and wide. The
dorsal Dorsal (from Latin ''dorsum'' ‘back’) may refer to: * Dorsal (anatomy), an anatomical term of location referring to the back or upper side of an organism or parts of an organism * Dorsal, positioned on top of an aircraft's fuselage The fus ...
sepal A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 Etymology The term ''sepalum'' ...
is longer than the
lateral Lateral is a geometric term of location which may also refer to: Biology and healthcare * Lateral (anatomy), a term of location meaning "towards the side" * Lateral cricoarytenoid muscle, an intrinsic muscle of the larynx * Lateral release ( ...
sepals but narrower and the
petal Petals are modified leaves that form an inner whorl surrounding the reproductive parts of flowers. They are often brightly coloured or unusually shaped to attract pollinators. All of the petals of a flower are collectively known as the ''corol ...
s are about the same length as the lateral sepals but only half as wide. The labellum has reddish purple spots or streaks and three lobes, the sides lobes erect and curved and the middle lobe pointed, rounded or more or less square. Flowering occurs between August and October for most varieties but some flower as early as May and others as late as November.


Taxonomy and naming

''Dendrobium speciosum'' was first formally described in 1804 by James Edward Smith from a specimen found at Port Jackson and sent to him by John White. The description was published in '' Exotic Botany''. The
specific epithet In Taxonomy (biology), taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin gramm ...
(''speciosum'') is a
Latin Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...
word meaning "beautiful", "splendid" or "showy". ''Dendrobium speciosum'' is a variable species and the following varieties are recognised by the World Checklist of Selected Plant Families: * ''Dendrobium speciosum'' var. ''blackdownense'' P.B.Adams which is usually a lithophyte, with between 14 and 113 off-white to deep golden yellow flowers from August to November and occurring on the Blackdown Tableland; * ''Dendrobium speciosum'' var. ''boreale'' P.B.Adams, Jac.M.Burke & S.D.Lawson which is epiphytic or lithophytic, with between 9 and 125, star-like, widely opening, off-white to cream-coloured flowers from July to September and has the most northerly distribution of the species, from the Annan River to near Townsville; * ''Dendrobium speciosum'' var. ''capricornicum'' Clemesha – commonly known as the Capricorn rock orchid, which is usually a lithophyte with between 30 and 50 white flowers between May and August and is found between the Byfield Range, Yeppoon and
Rockhampton Rockhampton is a city in the Rockhampton Region of Central Queensland, Australia. In the , the population of Rockhampton was 79,293. A common nickname for Rockhampton is "Rocky", and the demonym of Rockhampton is Rockhamptonite. The Scottish- ...
; * ''Dendrobium speciosum'' var. ''carnarvonense'' P.B.Adams – commonly known as the gorge pink rock orchid, which grows in gorges near streams and has between 2 and 7 relatively small, pale pink flowers between August and November and is only found between Carnarvon Gorge and Isla Gorge; * ''Dendrobium speciosum'' var. ''curvicaule'' F.M.Bailey – commonly known as the rainforest rock orchid, which is an epiphyte or lithophyte growing in or near rainforest and has between 25 and 45 white or cream-coloured flowers from July to September and grows between Mossman and Paluma in Queensland; * ''Dendrobium speciosum'' var. ''grandiflorum'' F.M.Bailey – commonly known as the golden king orchid and which grows on trees or rocks and has between 50 and 150 relatively large, creamy yellow to bright yellow flowers from August to October and is found from the Calliope Range to the Bunya Mountains in Queensland; * ''Dendrobium speciosum'' var. ''hillii'' Mast. – commonly known as the pale king orchid and which grows on trees and rocks and has between 70 and 200 crowde white or cream-coloured flowers from August to October and grows between Maleny in Queensland to Mangrove Mountain in New South Wales; * ''Dendrobium speciosum'' var. ''pedunculatum'' Clemesha – commonly known as the dwarf rock orchid which grows on rocks and has between 10 and 30 crowded white or cream-coloured flowers from July to August and occurs between the Mount Windsor National Park and the Evelyn Tableland in Queensland; * ''Dendrobium speciosum'' Sm. var. ''speciosum'' – commonly known as the Sydney rock orchid or rock lily which is a lithophyte with between 30 and 100 crowded cream-coloured or dull yellow flowers between August and October and occurs between Bulahdelah and near the Victorian border and as far inland as Mudgee. This species complex has been previously described by Alick Dockrill in 1969 and reviewed by several authors ( Stephen Clemesha 1981a, 1981b, 1986, Banks & Clemesha 1990 and Adams 1991. The 1889 book ''The Useful Native Plants of Australia'' records that common names included "rock lily" and that "the large pseudo-bulbs have been eaten by the aborigines, they, however, contain but little nutritive matter."


Pollination

Potential pollinators of ''Dendrobium speciosum'', such as the stingless bee '' Tetragonula carbonaria'', are attracted to the plant by large, cream to yellow, finely segmented, aromatic inflorescences. Flowers vary in size within the six recognised varieties of ''D. speciosum'' and are pollinated when visited by bees of appropriate size.


Natural hybrids

''Dendrobium speciosum'' forms a natural hybrid with ''Dendrobium gracilicaule'' : ''Dendrobium'' × ''gracillimum''. This hybrid has also been described as ''Dendrobium speciosum'' var. ''nitidum '' , ''Dendrobium '' × ''nitidum '' , ''Dendrobium speciosum'' var. ''bancroftianum'' and ''Dendrobium jonesii'' subsp. ''bancroftianum'' .


Use in horticulture

This orchid is popular in cultivation, growing into a large specimen that does well outdoors in climates with a mild winter. It may be grown in an open, coarse orchid growing medium, on a sturdy tree with an open canopy, or as a terrestrial in a well-drained position. It requires very bright light to full sun. Watering is year round in moderation. It is very temperature tolerant as long as it receives good warmth during the growing season. Frost, however, can cause extensive defoliation, an event the plant may require years to recover from. In cultivation ''D. speciosum'' can develop extremely large pseudobulbs, and benefits from regular fertilisation. Even in ideal cultivation conditions it may not flower every year, especially so in plants from more southern populations.


References


External links

* {{DEFAULTSORT:Dendrobium Speciosum speciosum Endemic orchids of Australia Epiphytic orchids Plants described in 1804 Lithophytic orchids