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''Cuvieronius'' is an extinct
New World The term ''New World'' is often used to mean the majority of Earth's Western Hemisphere, specifically the Americas."America." ''The Oxford Companion to the English Language'' (). McArthur, Tom, ed., 1992. New York: Oxford University Press, p. ...
genus of
gomphothere Gomphotheres are any members of the diverse, extinct taxonomic family Gomphotheriidae. Gomphotheres were elephant-like proboscideans, but do not belong to the family Elephantidae. They were widespread across Afro-Eurasia and North America dur ...
, named after the French naturalist
Georges Cuvier Jean Léopold Nicolas Frédéric, Baron Cuvier (; 23 August 1769 – 13 May 1832), known as Georges Cuvier, was a French naturalist and zoologist, sometimes referred to as the "founding father of paleontology". Cuvier was a major figure in na ...
. Alive, specimens typically stood about tall at the shoulder, weighed about and would have superficially resembled a modern
elephant Elephants are the largest existing land animals. Three living species are currently recognised: the African bush elephant, the African forest elephant, and the Asian elephant. They are the only surviving members of the family Elephantida ...
with spiral-shaped tusks.


Taxonomy

The species now known as ''Cuvieronius hyodon'' was among the first
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
animals from the New World to be studied. The first remains of this species were recovered from Ecuador by
Alexander von Humboldt Friedrich Wilhelm Heinrich Alexander von Humboldt (14 September 17696 May 1859) was a German polymath, geographer, naturalist, explorer, and proponent of Romantic philosophy and science. He was the younger brother of the Prussian minister ...
, at a location the local population referred to as the "Field of Giants".Mayor, A. (2005). Fossil legends of the first Americans. Princeton University Press. Humboldt recognized that, rather than being bones of giant humans as had been thought by the local population and previous Spanish colonists, they were similar to the giant elephants (''
Mastodon A mastodon ( 'breast' + 'tooth') is any proboscidean belonging to the extinct genus ''Mammut'' (family Mammutidae). Mastodons inhabited North and Central America during the late Miocene or late Pliocene up to their extinction at the end of the ...
'') being described from
Ohio Ohio () is a U.S. state, state in the Midwestern United States, Midwestern region of the United States. Of the List of states and territories of the United States, fifty U.S. states, it is the List of U.S. states and territories by area, 34th-l ...
. Humboldt sent teeth that he had collected from Mexico, Ecuador, and Chile to French anatomist
Georges Cuvier Jean Léopold Nicolas Frédéric, Baron Cuvier (; 23 August 1769 – 13 May 1832), known as Georges Cuvier, was a French naturalist and zoologist, sometimes referred to as the "founding father of paleontology". Cuvier was a major figure in na ...
, who classified the teeth into two species, which he referred to as the "mastodonte des cordilières" and the "mastodonte humboldtien", in an 1806 paper.Lucas, S.G. (2009)
Case 3479 ''Cuvieronius'' Osborn, 1923 (Mammalia, Proboscidea): Proposed conservation.
Bull. Zool. Nomen, 66, 1-6.
It was not until 1824 that Cuvier formally named the species. He referred both to the genus ''
Mastodon A mastodon ( 'breast' + 'tooth') is any proboscidean belonging to the extinct genus ''Mammut'' (family Mammutidae). Mastodons inhabited North and Central America during the late Miocene or late Pliocene up to their extinction at the end of the ...
'', calling them ''M. andium'' and ''M. humboldtii''. Unknown to Cuvier, Fischer had, in 1814, already named the two species based on Cuvier's original description, in the new genus ''Mastotherium'' as ''M. hyodon'' and ''M. humboldtii''. The idea of two distinct species continued to be accepted into the 20th century, usually using Cuvier's names, though Fischer's names were older. In 1923,
Henry Fairfield Osborn Henry Fairfield Osborn, Sr. (August 8, 1857 – November 6, 1935) was an American paleontologist, geologist and eugenics advocate. He was the president of the American Museum of Natural History for 25 years and a cofounder of the American Euge ...
recognized that these species were distinct from ''Mastodon'', and assigned each to its own new genus, as ''Cuvieronius humboldtii'' and ''Cordillerion andium''. However, by the 1930s, general agreement had shifted to regard both forms as representing a single, geographically widespread species, with ''Cuvieronius humboldtii'' considered to be the correct name. During the 1950s, the nomenclature of this species became increasingly tangled, as various scientists regarded the
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specim ...
of the genus ''Cuvieronius'' to be Fischer's first published name ''Mastotherium hyodon'', rather than the originally designated ''Mastodon humboldtii''. This situation went unaddressed until 2009, when Spencer Lucas petitioned the
International Commission on Zoological Nomenclature The International Commission on Zoological Nomenclature (ICZN) is an organization dedicated to "achieving stability and sense in the scientific naming of animals". Founded in 1895, it currently comprises 26 commissioners from 20 countries. Orga ...
to officially change the type species of ''Cuvieronius'' to ''M. hyodon'' as had been followed for over 50 years by that time, rather than abandoning the well-known ''Cuvieronius'' as a synonym. In 2011, Opinion 2276 of the ICZN ruled to conserve the names.


Evolution

''Cuvieronius'' initially evolved in North America during the Late
Pliocene The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58derived from, ''
Rhynchotherium ''Rhynchotherium'' is an extinct genus of proboscidea endemic to North America and Central America during the Miocene through Pliocene from 13.650 to 3.6 Ma, living for approximately . This gomphothere had two tusks and may have evolved fr ...
,'' a North American gomphothere genus known from the Late Miocene and Pliocene.Lucas, S.P., 2008a
Cuvieronius (Mammalia, Proboscidea) from the Neogene of Florida
In: Lucas, S.P., Morgan, G.S., Spielmann, J.A., Prothero, D.R. (Eds.), Neogene Mammals, vol. 44. N. M. Mus. Nat. Hist. Sci. Bull., pp. 31e38
''Cuvieronius'' was
extirpated Local extinction, also known as extirpation, refers to a species (or other taxon) of plant or animal that ceases to exist in a chosen geographic area of study, though it still exists elsewhere. Local extinctions are contrasted with global extinct ...
from most of North America during
Irvingtonian The Irvingtonian North American Land Mammal Age on the geologic timescale is the North American faunal stage according to the North American Land Mammal Ages chronology (NALMA), spanning from 1.9 million – 250,000 years BP.
after the arrival of
mammoth A mammoth is any species of the extinct elephantid genus ''Mammuthus'', one of the many genera that make up the order of trunked mammals called proboscideans. The various species of mammoth were commonly equipped with long, curved tusks an ...
s in North America around 1.3 Ma, presumably due to
competitive exclusion In ecology, the competitive exclusion principle, sometimes referred to as Gause's law, is a proposition that two species which compete for the same limited resource cannot coexist at constant population values. When one species has even the sligh ...
by mammoths and mastodons, but persisted in southern North America until the very end of the Pleistocene. During the
Great American Interchange The Great American Biotic Interchange (commonly abbreviated as GABI), also known as the Great American Interchange and the Great American Faunal Interchange, was an important late Cenozoic paleozoogeographic biotic interchange event in which lan ...
, ''Cuvieronius'' and a relative, ''
Notiomastodon ''Notiomastodon'' is an extinct proboscidean genus of gomphotheres (a distant relative to modern elephants) endemic to South America from the Pleistocene to the beginning of the Holocene. ''Notiomastodon'' specimens reached a size similar to th ...
'', dispersed into South America. ''Cuvieronius'' apparently reached South America considerably later than ''Notiomastodon'', with the oldest possible date being 0.76 ±0.03 Mya and the oldest confirmed date being 304 ±54 ka, and had a much more restricted range, confined mostly to the Andes. ''Cuvieronius'' remains have been identified as far south as
Chile Chile, officially the Republic of Chile, is a country in the western part of South America. It is the southernmost country in the world, and the closest to Antarctica, occupying a long and narrow strip of land between the Andes to the eas ...
, with remains initially suggested to represent ''Cuvieronius'' at the Quereo I site (Quereo Quebrada) dating to the
Late Pleistocene The Late Pleistocene is an unofficial age in the international geologic timescale in chronostratigraphy, also known as Upper Pleistocene from a stratigraphic perspective. It is intended to be the fourth division of the Pleistocene Epoch withi ...
11,600—11,400 BP. However, more recent studies suggest that all remains from the southern Andes actually belong to ''Notiomastodon'', and ''Cuvieronius'' was extirpated from South America by the end of the Late Pleistocene, before the arrival of people.


Distribution and habitat

The oldest fossil remains to date are of ''Cuvieronius'' species found in
Lincoln County, Nevada Lincoln County is a county located in the U.S. state of Nevada. As of the 2010 census, the population was 5,345. Its county seat is Pioche. Like many counties in Nevada, it is dry and sparsely populated, though notable for containing the Area ...
, which date to an accurate 4.6 million years ago (Mya). It was also found as far east as
South Carolina )''Animis opibusque parati'' ( for, , Latin, Prepared in mind and resources, links=no) , anthem = " Carolina";" South Carolina On My Mind" , Former = Province of South Carolina , seat = Columbia , LargestCity = Charleston , LargestMetro = ...
and
North Carolina North Carolina () is a U.S. state, state in the Southeastern United States, Southeastern region of the United States. The state is the List of U.S. states and territories by area, 28th largest and List of states and territories of the United ...
in
Pleistocene The Pleistocene ( , often referred to as the ''Ice age'') is the geological Epoch (geology), epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fina ...
rocks dating to 1.81 million to 126,000 years ago. In
Florida Florida is a state located in the Southeastern region of the United States. Florida is bordered to the west by the Gulf of Mexico, to the northwest by Alabama, to the north by Georgia, to the east by the Bahamas and Atlantic Ocean, and ...
, remains show both ''Cuvieronius'' living from 2 to 0.5 Mya. The most recent findings of ''Cuvieronius'' in North America are in Sonora, Mexico, that date to 13,390 years BP at a
Clovis culture The Clovis culture is a prehistoric Paleoamerican culture, named for distinct stone and bone tools found in close association with Pleistocene fauna, particularly two mammoths, at Blackwater Locality No. 1 near Clovis, New Mexico, in 1936 a ...
kill site. As well as remains found near the town of
Hockley Hockley is a large village and civil parish in Essex in the East of England located between Chelmsford and Southend-on-Sea, or, more specifically, between Rayleigh and Rochford. It came to prominence during the coming of the railway in the 189 ...
in Texas near
Austin Austin is the capital city of the U.S. state of Texas, as well as the seat and largest city of Travis County, with portions extending into Hays and Williamson counties. Incorporated on December 27, 1839, it is the 11th-most-populous city ...
, which date to around 24,000 years BP. According to a group of Brazilian mammalogists, many sites in South America referred to ''Cuvieronius'' actually refer to ''
Notiomastodon ''Notiomastodon'' is an extinct proboscidean genus of gomphotheres (a distant relative to modern elephants) endemic to South America from the Pleistocene to the beginning of the Holocene. ''Notiomastodon'' specimens reached a size similar to th ...
'', with many previous studies simply labeling fossils one or the other depending on location, with only localities definitely identified as ''Cuvieronius'', the range now extends in the high
Andes The Andes, Andes Mountains or Andean Mountains (; ) are the longest continental mountain range in the world, forming a continuous highland along the western edge of South America. The range is long, wide (widest between 18°S – 20°S ...
from Ecuador in the north, to Bolivia in the south, with the localities in the southern Andes in Chile and Argentina now thought to belong to ''Notiomastodon''. The same group maintains that all the South American specimens represent the single species ''C. hyodon'', and that no confirmed fossils of ''Cuvieronius'' are more recent than 44,000 years ago in South America, so the species would not have been in South America at the time of human arrival. By the end of the
Pleistocene The Pleistocene ( , often referred to as the ''Ice age'') is the geological Epoch (geology), epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fina ...
, the northern limit of the range of ''Cuvieronius'' was in
Mexico Mexico (Spanish language, Spanish: México), officially the United Mexican States, is a List of sovereign states, country in the southern portion of North America. It is borders of Mexico, bordered to the north by the United States; to the so ...
and Central Texas. Fossils of ''Cuvieronius'' have been found in the Ulloma and Tarija Formations of Bolivia, the
Sabana Formation The Sabana Formation ( es, Formación Sabana, Q1sa, QTs) is a geological formation of the Bogotá savanna, Altiplano Cundiboyacense, Eastern Ranges of the Colombian Andes. The formation consists mainly of shales with at the edges of the Bogotá ...
of Colombia,Correal Urrego, 1990, p.77 Costa Rica, Ecuador, El Salvador, Guatemala, Mexico, Nicaragua, Panama, United States (Florida, New Mexico, North Carolina, Oklahoma, South Carolina, Texas), and Venezuela.''Cuvieronius''
at
Fossilworks Fossilworks is a portal which provides query, download, and analysis tools to facilitate access to the Paleobiology Database, a large relational database assembled by hundreds of paleontologists from around the world. History Fossilworks was crea ...
.org


References


Bibliography

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External links


American Museum of Natural History
Includes images {{Taxonbar, from=Q139070 Gomphotheres Prehistoric placental genera Pleistocene proboscideans Zanclean first appearances Holocene extinctions Pliocene proboscideans Pliocene mammals of North America Pleistocene mammals of North America Rancholabrean Irvingtonian Blancan Neogene Mexico Neogene United States Pleistocene Costa Rica Pleistocene El Salvador Pleistocene Guatemala Pleistocene Mexico Pleistocene Nicaragua Pleistocene Panama Pleistocene United States Fossils of Costa Rica Fossils of El Salvador Fossils of Guatemala Fossils of Mexico Fossils of Nicaragua Fossils of Panama Fossils of the United States Pliocene mammals of South America Pleistocene mammals of South America Lujanian Ensenadan Uquian Chapadmalalan Montehermosan Neogene Chile Pleistocene Bolivia Pleistocene Chile Pleistocene Colombia Pleistocene Ecuador Pleistocene Venezuela Fossils of Bolivia Fossils of Chile Fossils of Colombia Fossils of Ecuador Fossils of Venezuela Fossil taxa described in 1923 Taxa named by Henry Fairfield Osborn