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''Cteniogenys'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial no ...
of
choristodere Choristodera (from the Greek χωριστός ''chōristos'' + δέρη ''dérē'', 'separated neck') is an extinct order of semiaquatic diapsid reptiles that ranged from the Middle Jurassic, or possibly Triassic, to the late Miocene (168 ...
, a morphologically diverse group of aquatic reptiles. It is part of the monotypic family Cteniogenidae. The type and only named species, ''C. antiquus'', was named in 1928 by Charles W. Gilmore. The holotype, VP.001088, was collected in the
Morrison Formation The Morrison Formation is a distinctive sequence of Upper Jurassic sedimentary rock found in the western United States which has been the most fertile source of dinosaur fossils in North America. It is composed of mudstone, sandstone, siltstone, ...
(Como Bluff), Wyoming in 1881 by William H. Reed. More specimens have been discovered since then, including specimens from the Late Jurassic of Portugal and Middle Jurassic of Britain, which have not been assigned to species.


Description

''Cteniogenys'' was long, and probably weighed less than . The skull of this genus was long and slender, and the jaws had numerous conical teeth. ''Cteniogenys'' in the Morrison probably fed on insects and small fish. It is mostly known from freshwater sites (rivers and ponds), and is a rare find in the formation (only 60 specimens out of over 2,800 total vertebrate specimens known from the formation), mostly known from northern outcrops (particularly Wyoming); this may reflect a preservation bias against small animals in terrestrial settings, rather than an accurate reflection of ''Cteniogenys'' populations in the Morrison.


Distribution

Fossils of ''Cteniogenys'' are known from the upper Middle Jurassic ( Bathonian) aged
Forest Marble Formation The Forest Marble is a geological formation in England. Part of the Great Oolite Group, it dates to the late Bathonian stage of the Middle Jurassic.Weishampel, David B; et al. (2004). "Dinosaur distribution (Middle Jurassic, Europe)." In: Weisha ...
and Kilmaluag Formation of Britain, the
Late Jurassic The Late Jurassic is the third epoch of the Jurassic Period, and it spans the geologic time from 163.5 ± 1.0 to 145.0 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic strata.Owen 1987. In European lithostratigraphy, the nam ...
(
Kimmeridgian In the geologic timescale, the Kimmeridgian is an age in the Late Jurassic Epoch and a stage in the Upper Jurassic Series. It spans the time between 157.3 ± 1.0 Ma and 152.1 ± 0.9 Ma (million years ago). The Kimmeridgian follows the Oxford ...
) aged Alcobaça Formation of Portugal, and the Late Jurassic-age
Morrison Formation The Morrison Formation is a distinctive sequence of Upper Jurassic sedimentary rock found in the western United States which has been the most fertile source of dinosaur fossils in North America. It is composed of mudstone, sandstone, siltstone, ...
of western
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the C ...
, indeterminate similar remains are also known from the Callovian aged
Balabansai Formation The Balabansai Formation (sometimes transcribed Balabansay, also known as Balabansai Svita) is a geological formation in Kazakhstan, Kyrgyzstan and Uzbekistan whose strata date back to the Bathonian and Callovian stages of the Middle Jurassic. ...
of
Kyrgyzstan Kyrgyzstan,, pronounced or the Kyrgyz Republic, is a landlocked country in Central Asia. Kyrgyzstan is bordered by Kazakhstan to the north, Uzbekistan to the west, Tajikistan to the south, and the People's Republic of China to the east ...
and the Bathonian aged
Itat Formation The Itat Formation ( Russian: итатская свита) is a geologic formation in western Siberia. It was deposited in the Bajocian to Bathonian ages of the Middle Jurassic. Dinosaur remains are among the fossils that have been recovered from ...
of western Siberia, and the Berriasian aged Angeac-Charente bonebed in France.Ronan Allain, Romain Vullo, Lee Rozada, Jérémy Anquetin, Renaud Bourgeais, et al.
Vertebrate paleobiodiversity of the Early Cretaceous (Berriasian) Angeac-Charente Lagerstätte (southwestern France): implications for continental faunal turnover at the J/K boundary
Geodiversitas, Museum National d’Histoire Naturelle Paris, In press. ffhal-03264773f
A handful of skull and jaw fragments from the Late Cretaceous Oldman Formation and
Dinosaur Park Formation The Dinosaur Park Formation is the uppermost member of the Belly River Group (also known as the Judith River Group), a major geologic unit in southern Alberta. It was deposited during the Campanian stage of the Late Cretaceous, between about 7 ...
in Canada were assigned to ''Cteniogenys'' by Gao and Fox (1998). Given the long gap in time, however, Gao et al. (2005) cautioned that these remains could constitute a different, as-yet undetermined genus. In their description of new material of '' Khurendukhosaurus'', Matsumoto et al. (2009) agreed that the putative Cretaceous occurrence of ''Cteniogenys'' did not belong to that genus and represents an indeterminate choristoderan, possibly a new genus.


Systematics

The genus was named in 1928 by Charles W. Gilmore on the basis of a lower jaw collected during the late 19th century by
Othniel Charles Marsh Othniel Charles Marsh (October 29, 1831 – March 18, 1899) was an American professor of Paleontology in Yale College and President of the National Academy of Sciences. He was one of the preeminent scientists in the field of paleontology. Among h ...
's workers at Como Bluff. He tentatively described the genus as a lizard, noting that it could instead be a frog. New material of ''Cteniogenys'' from Europe prompted Susan E. Evans to reclassify it as an early choristodere in 1989. To date it is only known from fragmentary specimens. In phylogenic analyses it has been found to the basal most choristodere.


See also

*
Paleobiota of the Morrison Formation The Morrison Formation is a distinctive sequence of Late Jurassic sedimentary rock that is found in the western United States, which has a wide assortment of taxa represented in its fossil record, including dinosaur fossils in North America. It is ...


References

Choristodera Jurassic reptiles Late Jurassic reptiles of North America Jurassic Colorado Jurassic geology of Oklahoma Jurassic geology of South Dakota Jurassic geology of Utah Jurassic geology of Wyoming Late Cretaceous reptiles of North America Fossil taxa described in 1928 Prehistoric reptile genera {{cretaceous-reptile-stub