Crichtonpelta
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''Crichtonpelta'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
of extinct herbivorous
ankylosaurid Ankylosauridae () is a family of armored dinosaurs within Ankylosauria, and is the sister group to Nodosauridae. The oldest known Ankylosaurids date to around 122 million years ago and went extinct 66 million years ago during the Cretaceous–Pal ...
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
from the
Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', ...
( Cenomanian) of China.


Discovery and naming

In 2007,
Lü Junchang Lü Junchang (; 1965 – 9 October 2018) was a Chinese palaeontologist and professor at the Institute of Geology, Chinese Academy of Geological Sciences. An expert on Mesozoic reptiles, he described and named dozens of dinosaur and pterosaur taxa ...
, Ji Qiang, Gao Yubo and Li Zhixin named and described a second species of ''
Crichtonsaurus ''Crichtonsaurus'' (meaning "Crichton's lizard") is a genus of herbivorous ankylosaurid dinosaur that lived during the Late Cretaceous in what is now China. It was named after Michael Crichton, the author of the dinosaur novel ''Jurassic Park''. ...
'': ''Crichtonsaurus benxiensis''. The specific name refers to the Benxi Geological Museum. The
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of sever ...
, BXGMV0012, is a skull found near Beipiao in a layer of the
Sunjiawan Formation The Sunjiawan Formation () is a geological formation in Liaoning, China, with strata possibly dating back to the early Late Cretaceous, specifically the Cenomanian.Xi et al., 2018 Dinosaur remains are among the fossils that have been recovered from ...
probably dating from the early Cenomanian (~99–95 Ma). Specimen BXGMV0012-1, a skeleton lacking the skull, discovered in the same quarry as the holotype, was referred to the species. Furthermore, a skeleton with skull, displayed by the Sihetun Fossil Museum as a ''Crichtonsaurus bohlini'' specimen, was in 2014 referred to ''Crichtonpelta''. In 2017, a fourth specimen was described, from the same quarry as the holotype, G20090034, consisting of a skull lacking the front snout.YANG Jingtao, YOU Hailu, XIE Li & ZHOU Hongrui, 2017, "A New Specimen of ''Crichtonpelta benxiensis'' (Dinosauria: Ankylosaurinae) from the Mid-Cretaceous of Liaoning Province, China", ''Acta Geologica Sinica'' 91(3): 781-790 In 2014,
Victoria Arbour Victoria Megan Arbour is a Canadian evolutionary biologist and vertebrate palaeontologist at Royal BC Museum, where she is Curator of Palaeontology. An "expert on the armoured dinosaurs known as ankylosaurs", Arbour analyzes fossils and creates ...
concluded that ''Crichtonsaurus'' were a '' nomen dubium''. Therefore, she named a separate genus for its second species: ''Crichtonpelta''. The generic name combines a reference to Michael Crichton, the author of '' Jurassic Park'', with a Greek πέλτη, ''peltè'', "small shield". At the time this was an invalid ''nomen ex dissertatione''. However, in 2015, ''Crichtonpelta'' was validly named by Arbour and
Philip John Currie Philip John Currie (born March 13, 1949) is a Canadian palaeontologist and museum curator who helped found the Royal Tyrrell Museum of Palaeontology in Drumheller, Alberta and is now a professor at the University of Alberta in Edmonton. In the ...
. The
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specime ...
is ''Crichtonsaurus benxiensis''; the '' combinatio nova'' is ''Crichtonpelta benxiensis''. There was a possibility that, though ''Crichtonsaurus bohlini'' was a ''nomen dubium'', its fossil material in fact belonged to ''Crichtonpelta''. Arbour however, noted clear differences in the scapula and humerus between BXGMV0012-1 and LPM 101, a specimen previously referred to ''Crichtonsaurus bohlini'': the scapula of the former has a tab-like
acromion In human anatomy, the acromion (from Greek: ''akros'', "highest", ''ōmos'', "shoulder", plural: acromia) is a bony process on the scapula (shoulder blade). Together with the coracoid process it extends laterally over the shoulder joint. The ac ...
and its humerus a much longer deltopectoral crest.


Description

Arbour established several distinguishing traits. One of these was an
autapomorphy In phylogenetics, an autapomorphy is a distinctive feature, known as a derived trait, that is unique to a given taxon. That is, it is found only in one taxon, but not found in any others or outgroup taxa, not even those most closely related to ...
, unique derived trait: the apex of the (quadrato)jugal, or cheek, horn, is pointing upwards. Also a unique combination of in themselves not unique traits is present. The upper snout armour forms an amorphous mass, not clearly separated into distinctive tiles. The
jugal bone The jugal is a skull bone found in most reptiles, amphibians and birds. In mammals, the jugal is often called the malar or zygomatic. It is connected to the quadratojugal and maxilla, as well as other bones, which may vary by species. Anato ...
is deeper than that of ''
Pinacosaurus ''Pinacosaurus'' (meaning "Plank lizard") is a genus of ankylosaurid thyreophoran dinosaur that lived in Asia during the Late Cretaceous (Santonian-Campanian, roughly 75 million to 71 million years ago), mainly in Mongolia and China. The first r ...
''. The skull roof is not notched at the
lacrimal bone The lacrimal bone is a small and fragile bone of the facial skeleton; it is roughly the size of the little fingernail. It is situated at the front part of the medial wall of the orbit. It has two surfaces and four borders. Several bony landmarks of ...
as in ''Pinacosaurus grangeri''. The squamosal horns are shorter than those of ''Pinacosaurus mephistocephalus''. However, these horns are longer and more pointed than those of ''
Gobisaurus ''Gobisaurus'' is an extinct genus of herbivorous basal ankylosaurid ankylosaur from the Upper Cretaceous (and possibly also the Lower Cretaceous) of China (''Nei Mongol Zizhiqu''). The genus is monotypic, containing only the species ''Gobisa ...
'' or ''
Shamosaurus ''Shamosaurus'' is an extinct genus of herbivorous basal ankylosaurid ankylosaur from Early Cretaceous ( Aptian to Albian stage) deposits of Höövör, Mongolia. Discovery and naming In 1977, a Soviet-Mongolian expedition discovered the ske ...
''. The point of the cheek horn is located on the rear edge. The transverse crest on the top of the rear skull has two points. The holotype of ''Crichtonpelta'' is somewhat larger than ''Crichtonsaurus'', itself about three to four metres long. It is uncertain whether ''Crichtonpelta'' already possessed a tail club.


Classification

''Crichtonpelta'' was, within the
Ankylosauridae Ankylosauridae () is a family of armored dinosaurs within Ankylosauria, and is the sister group to Nodosauridae. The oldest known Ankylosaurids date to around 122 million years ago and went extinct 66 million years ago during the Cretaceous–Pa ...
, placed in the
Ankylosaurinae Ankylosaurinae is a subfamily of ankylosaurid dinosaurs, existing from the Early Cretaceous about 105 million years ago until the end of the Late Cretaceous, about 66 mya. Many genera are included in the clade, such as ''Ankylosaurus'', ''Pina ...
, in a basal position. If correct, this makes it the oldest known ankylosaurine.


References

{{Taxonbar, from=Q20737863 Ankylosaurids Late Cretaceous dinosaurs of Asia Taxa named by Philip J. Currie Fossil taxa described in 2015 Paleontology in Liaoning Chaoyang, Liaoning Taxa named by Lü Junchang Ornithischian genera