The smooth snake (''Coronella austriaca'')Street D (1979). ''The Reptiles of Northern and Central Europe''. London: B.T. Batsford Ltd. 268 pp. . is a

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

of non-

venomous snake Venomous snakes are species of the suborder Serpentes that are capable of producing venom, which they use for killing prey, for defense, and to assist with digestion of their prey. The venom is typically delivered by injection using hollow or g ...

in the

family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...

Colubridae. The species is found in northern and central

Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a subcontinent of Eurasia and it is located entirel ...

, but also as far east as northern

Iran Iran, officially the Islamic Republic of Iran, and also called Persia, is a country located in Western Asia. It is bordered by Iraq and Turkey to the west, by Azerbaijan and Armenia to the northwest, by the Caspian Sea and Turkmeni ...

. The

Reptile Database The Reptile Database is a scientific database that collects taxonomic information on all living reptile species (i.e. no fossil species such as dinosaurs). The database focuses on species (as opposed to higher ranks such as families) and has entrie ...

recognizes two subspecies as being valid, including the

nominotypical subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...

described here.

Description

Both sexes of ''C. austriaca'' grow to an average total length (including tail) of about to . Two specimens measuring have been recorded in Sweden, as well as one in

Russia Russia (, , ), or the Russian Federation, is a transcontinental country spanning Eastern Europe and Northern Asia. It is the largest country in the world, with its internationally recognised territory covering , and encompassing one-eig ...

that was . The head has a

rostral scale The rostral scale, or rostral, in snakes and other scaled reptiles is the median plate on the tip of the snout that borders the mouth opening. Wright AH, Wright AA (1957). ''Handbook of Snakes of the United States and Canada''. Ithaca and London: ...

that is at least as deep as it is wide, creating a triangular indentation between the

internasal scales In snakes, the internasal scales are those on top of the head between the scales that surround the nostrils. Wright AH, Wright AA (1957). ''Handbook of Snakes of the United States and Canada''. Ithaca and London: Comstock Publishing Associates. (7t ...

(rarely separating them). The top of the head is covered with nine large plates. The

nasal scale In reptiles, the nasal scale refers to the scale that encloses the nostril.Mallow D, Ludwig D, Nilson G. 2003. True Vipers: Natural History and Toxinology of Old World Vipers. Malabar, Florida: Krieger Publishing Company. 359 pp. . Sometimes this ...

is often divided. There is one (rarely two) preoculars and two postoculars. The temporals number 2+2 or 2+3 (rarely 1+2) . There are seven (rarely eight)

upper labials In reptiles, the supralabial scales, also called upper-labials, are those scales that border the mouth opening along the upper jaw. They do not include the median scaleWright AH, Wright AA. 1957. Handbook of Snakes. Comstock Publishing Associates ...

, of which the third and fourth or fourth and fifth border the eye. In the middle part of the body there are 19 (rarely 17 or 21) rows of

dorsal scales In snakes, the dorsal scales are the longitudinal series of plates that encircle the body, but do not include the ventral scales In snakes, the ventral scales or gastrosteges are the enlarged and transversely elongated scales that extend down t ...

. In contrast with many other snakes found in the region, these scales are flat (not keeled). This gives the snake a smooth texture to the touch, from which it gets its common name. The

ventral scales In snakes, the ventral scales or gastrosteges are the enlarged and transversely elongated scales that extend down the underside of the body from the neck to the anal scale. When counting them, the first is the anteriormost ventral scale that cont ...

number 150-164 in males and 162–200 in females. The

anal scale Anal may refer to: Related to the anus *Related to the anus of animals: ** Anal fin, in fish anatomy ** Anal vein, in insect anatomy ** Anal scale, in reptile anatomy *Related to the human anus: ** Anal sex, a type of sexual activity involv ...

is divided (rarely single) and the subcaudal scales are paired. Males have 54–70 subcaudal scales and females 40–76. The colour pattern consists of a brown, grey or reddish ground colourSteward JW (1971). ''The Snakes of Europe''. Cranbury, New Jersey: Associated University Press (Fairleigh Dickinson University Press). 238 pp. LCCCN 77-163307. . with two rows of small, rather indistinct dark spots running down the back towards the tail. In some cases, each pair of spots may be united toward the neck area, forming a series of cross-bars over the back. There is also a very indistinct series of dark spots running along each of the flanks. These four series of spots along the body overlay four parallel, rather shadowy stripes that also run down the back and flanks. On the top of the head is a dark marking which is often in the shape of a crown, giving rise to the

generic Generic or generics may refer to: In business * Generic term, a common name used for a range or class of similar things not protected by trademark * Generic brand, a brand for a product that does not have an associated brand or trademark, other ...

name ''Coronella'' (which means

coronet A coronet is a small crown consisting of ornaments fixed on a metal ring. A coronet differs from other kinds of crowns in that a coronet never has arches, and from a tiara in that a coronet completely encircles the head, while a tiara doe ...

). A relatively thick dark stripe extends from each nostril, through the eye, and along the side of the head to a little beyond the neck. The upper labials are whitish, greyish-white or light brown, sometimes with darker spots. The tongue is reddish brown or dark red.

Biology

The smooth snake feeds on smaller animals, especially other reptiles. It subdues larger prey by constriction, although unlike true constrictors it does not kill by this method. Smooth snakes are

ovoviviparous Ovoviviparity, ovovivipary, ovivipary, or aplacental viviparity is a term used as a "bridging" form of reproduction between egg-laying oviparous and live-bearing viviparous reproduction. Ovoviviparous animals possess embryos that develop insi ...

. The juveniles hatch out of eggs internally and are born live. In Britain it is restricted to

heathland A heath () is a shrubland habitat found mainly on free-draining infertile, acidic soils and characterised by open, low-growing woody vegetation. Moorland is generally related to high-ground heaths with—especially in Great Britain—a coole ...

habitats.

Geographic range

''Coronella austriaca'' is found from the south of

England England is a country that is part of the United Kingdom. It shares land borders with Wales to its west and Scotland to its north. The Irish Sea lies northwest and the Celtic Sea to the southwest. It is separated from continental Europe b ...

through

France France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of overseas regions and territories in the Americas and the Atlantic, Pacific and Indian Oceans. Its metropolitan area ...

and the

Low Countries The term Low Countries, also known as the Low Lands ( nl, de Lage Landen, french: les Pays-Bas, lb, déi Niddereg Lännereien) and historically called the Netherlands ( nl, de Nederlanden), Flanders, or Belgica, is a coastal lowland region in N ...

to northern

Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = ''Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , i ...

and

Portugal Portugal, officially the Portuguese Republic ( pt, República Portuguesa, links=yes ), is a country whose mainland is located on the Iberian Peninsula of Southwestern Europe, and whose territory also includes the Atlantic archipelagos of ...

Germany Germany,, officially the Federal Republic of Germany, is a country in Central Europe. It is the second most populous country in Europe after Russia, and the most populous member state of the European Union. Germany is situated betwe ...

Norway Norway, officially the Kingdom of Norway, is a Nordic country in Northern Europe, the mainland territory of which comprises the western and northernmost portion of the Scandinavian Peninsula. The remote Arctic island of Jan Mayen and the ...

and Sweden (as far north as latitude 63°), Latvia, Lithuania,

Estonia Estonia, formally the Republic of Estonia, is a country by the Baltic Sea in Northern Europe. It is bordered to the north by the Gulf of Finland across from Finland, to the west by the sea across from Sweden, to the south by Latvia, a ...

, Switzerland,

Austria Austria, , bar, Östareich officially the Republic of Austria, is a country in the southern part of Central Europe, lying in the Eastern Alps. It is a federation of nine states, one of which is the capital, Vienna, the most populous ...

Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical ...

and

Sicily (man) it, Siciliana (woman) , population_note = , population_blank1_title = , population_blank1 = , demographics_type1 = Ethnicity , demographics1_footnotes = , demographi ...

(but not in Corsica or

Sardinia Sardinia ( ; it, Sardegna, label=Italian, Corsican and Tabarchino ; sc, Sardigna , sdc, Sardhigna; french: Sardaigne; sdn, Saldigna; ca, Sardenya, label=Algherese and Catalan) is the second-largest island in the Mediterranean Sea, after ...

), the western

Balkans The Balkans ( ), also known as the Balkan Peninsula, is a geographical area in southeastern Europe with various geographical and historical definitions. The region takes its name from the Balkan Mountains that stretch throughout the who ...

and

Greece Greece,, or , romanized: ', officially the Hellenic Republic, is a country in Southeast Europe. It is situated on the southern tip of the Balkans, and is located at the crossroads of Europe, Asia, and Africa. Greece shares land borders ...

, and European Russia as far north as latitude 57°. In

Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an are ...

, it is found from

Turkey Turkey ( tr, Türkiye ), officially the Republic of Türkiye ( tr, Türkiye Cumhuriyeti, links=no ), is a transcontinental country located mainly on the Anatolian Peninsula in Western Asia, with a small portion on the Balkan Peninsula in ...

Azerbaijan Azerbaijan (, ; az, Azərbaycan ), officially the Republic of Azerbaijan, , also sometimes officially called the Azerbaijan Republic is a transcontinental country located at the boundary of Eastern Europe and Western Asia. It is a part of t ...

Georgia Georgia most commonly refers to: * Georgia (country), a country in the Caucasus region of Eurasia * Georgia (U.S. state), a state in the Southeast United States Georgia may also refer to: Places Historical states and entities * Related to the ...

Armenia Armenia (), , group=pron officially the Republic of Armenia,, is a landlocked country in the Armenian Highlands of Western Asia.The UNbr>classification of world regions places Armenia in Western Asia; the CIA World Factbook , , and ' ...

and northern

. It is oddly absent in Denmark despite the species being found just south of the German border as well as southern Sweden. The species is only known from Denmark from 6 specimens, all found between 1870 and 1914. In

Finland Finland ( fi, Suomi ; sv, Finland ), officially the Republic of Finland (; ), is a Nordic country in Northern Europe. It shares land borders with Sweden to the northwest, Norway to the north, and Russia to the east, with the Gulf of B ...

, the species is found only in

Åland Åland ( fi, Ahvenanmaa: ; ; ) is an autonomous and demilitarised region of Finland since 1920 by a decision of the League of Nations. It is the smallest region of Finland by area and population, with a size of 1,580 km2, and a populat ...

, and it is not common there.

Subspecies

References

Gallery

Image:Rézsikló (Coronella austriaca) feje.jpg, Head of ''C. austriaca''. Image:Rézsikló (Coronella austriaca)3.jpg, ''C. austriaca'' in

Hungary Hungary ( hu, Magyarország ) is a landlocked country in Central Europe. Spanning of the Pannonian Basin, Carpathian Basin, it is bordered by Slovakia to the north, Ukraine to the northeast, Romania to the east and southeast, Serbia to the ...

. File:Coronella austriaca s.jpg, ''C. austriaca'' in

Bosnia and Herzegovina Bosnia and Herzegovina ( sh, / , ), abbreviated BiH () or B&H, sometimes called Bosnia–Herzegovina and often known informally as Bosnia, is a country at the crossroads of south and southeast Europe, located in the Balkans. Bosnia and H ...

. Image:coronella_austriaca.jpg, ''C. austriaca'' curled up. Image:CoronellaAustriaca2.jpg, Subadult ''C. austriaca'' in northern

Description

Biology

Geographic range

Subspecies

See also

References

Further reading

Gallery

External links