Climacoceratidae is a family of superficially deer-like

artiodactyl The even-toed ungulates (Artiodactyla , ) are ungulates—hoofed animals—which bear weight equally on two (an even number) of their five toes: the third and fourth. The other three toes are either present, absent, vestigial, or pointing poster ...

ungulate Ungulates ( ) are members of the diverse clade Ungulata which primarily consists of large mammals with hooves. These include odd-toed ungulates such as horses, rhinoceroses, and tapirs; and even-toed ungulates such as cattle, pigs, giraffes, cam ...

s which lived in the

Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...

epoch in

Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...

. They are close to the ancestry of

giraffes The giraffe is a large African hoofed mammal belonging to the genus ''Giraffa''. It is the tallest living terrestrial animal and the largest ruminant on Earth. Traditionally, giraffes were thought to be one species, ''Giraffa camelopardalis ...

, with some genera, such as ''

Prolibytherium ''Prolibytherium'' is an extinct genus of prolibytheriid artiodactyl ungulate native to Middle Miocene North Africa and Pakistan, from around 16.9 to 15.97 million years ago. Fossils of ''Prolibytherium'' were found in the Marada Formation of ...

'', originally classified as giraffes. The climacoceratids, namely members of what is now the type genus ''

Climacoceras ''Climacoceras'' (from Greek for "ladder horns") is a genus of extinct artiodactyl ungulates that lived in Africa and Europe during the Miocene. The members of ''Climacoceras'' were related to giraffes, and the genus was formerly placed within th ...

'', were originally placed within the family

Palaeomerycidae The Palaeomerycidae are an extinct family of ruminants in the order Artiodactyla. Palaeomerycids lived in North America, Europe, Africa and Asia from 33 to 4.9 million years ago (from the Late Eocene to Pliocene epochs), existing for about 28 mi ...

, and then within

Giraffidae The Giraffidae are a family of ruminant artiodactyl mammals that share a common ancestor with deer and bovids. This family, once a diverse group spread throughout Eurasia and Africa, presently comprises only two extant genera, the giraffe (one or ...

. In 1978, W. R. Hamilton erected a new family, placing it close to Giraffidae within the superfamily

Giraffoidea Giraffoidea is a superfamily that includes the families Climacoceratidae, Antilocapridae, and Giraffidae. The only extant members in the superfamily are the pronghorn, giraffe, and okapi. The Climacoceratidae are also placed in the superfamily, ...

. They differ from giraffes in that their

antler Antlers are extensions of an animal's skull found in members of the Cervidae (deer) family. Antlers are a single structure composed of bone, cartilage, fibrous tissue, skin, nerves, and blood vessels. They are generally found only on male ...

-like

ossicone Ossicones are columnar or conical skin-covered bone structures on the heads of giraffes, male okapi, and some of their extinct relatives. Ossicones are distinguished from the superficially similar structures of horns and antlers by their unique ...

s are derived from different bones.

References

* MacInnes, D. G. 1936, "A new genus of fossil deer from the Miocene of Africa", Journal of the Linnean Society, Zoology 39: 521–530. * Hamilton, W. R., 1978, "Cervidae and Palaeomerycidae", 495–508, in Maglio, V. J. & Cooke, H. B. S., (eds.) "Evolution of African mammals", Harvard University Press, Cambridge, Massachusetts & London, England, 1978, xiv-641 Giraffoidea Miocene first appearances Miocene extinctions Prehistoric mammal families {{paleo-eventoedungulate-stub