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Carnivora is a
monophyletic In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic gr ...
order of placental mammals consisting of the most recent common ancestor of all cat-like and dog-like animals, and all descendants of that ancestor. Members of this group are formally referred to as carnivorans, and have evolved to specialize in eating flesh. The order is the fifth largest order of mammals, comprising at least 279 species. Carnivorans live on every major landmass and in a variety of habitats, ranging from the cold polar regions to the hyper-arid region of the
Sahara Desert , photo = Sahara real color.jpg , photo_caption = The Sahara taken by Apollo 17 astronauts, 1972 , map = , map_image = , location = , country = , country1 = , ...
to the open seas. They come in a very large array of different body plans in contrasting shapes and sizes. Carnivora can be divided into two subclades: the cat-like
Feliformia Feliformia is a suborder within the order Carnivora consisting of "cat-like" carnivorans, including cats (large and small), hyenas, mongooses, viverrids, and related taxa. Feliformia stands in contrast to the other suborder of Carnivora, Canifor ...
and the dog-like
Caniformia Caniformia is a suborder within the order Carnivora consisting of "dog-like" carnivorans. They include dogs (wolves, foxes, etc.), bears, raccoons, and mustelids. The Pinnipedia ( seals, walruses and sea lions) are also assigned to this group. T ...
, which are differentiated based on the structure of their ear bones and cranial features. The feliforms include families such as the
cats The cat (''Felis catus'') is a domestic species of small carnivorous mammal. It is the only domesticated species in the family Felidae and is commonly referred to as the domestic cat or house cat to distinguish it from the wild members o ...
, the
hyena Hyenas, or hyaenas (from Ancient Greek , ), are feliform carnivoran mammals of the family Hyaenidae . With only four extant species (each in its own genus), it is the fifth-smallest family in the Carnivora and one of the smallest in the cl ...
s, the
mongoose A mongoose is a small terrestrial carnivorous mammal belonging to the family Herpestidae. This family is currently split into two subfamilies, the Herpestinae and the Mungotinae. The Herpestinae comprises 23 living species that are native to so ...
s and the
civets A civet () is a small, lean, mostly nocturnal mammal native to tropical Asia and Africa, especially the tropical forests. The term civet applies to over a dozen different species, mostly from the family Viverridae. Most of the species diversity ...
. The majority of feliform species are found in the
Old World The "Old World" is a term for Afro-Eurasia that originated in Europe , after Europeans became aware of the existence of the Americas. It is used to contrast the continents of Africa, Europe, and Asia, which were previously thought of by thei ...
, though the cats and one extinct genus of hyena have successfully diversified into the
Americas The Americas, which are sometimes collectively called America, are a landmass comprising the totality of North America, North and South America. The Americas make up most of the land in Earth's Western Hemisphere and comprise the New World. ...
. The caniforms include the
dogs The dog (''Canis familiaris'' or ''Canis lupus familiaris'') is a domesticated descendant of the wolf. Also called the domestic dog, it is derived from the extinct Pleistocene wolf, and the modern wolf is the dog's nearest living relative. ...
,
bear Bears are carnivoran mammals of the family Ursidae. They are classified as caniforms, or doglike carnivorans. Although only eight species of bears are extant, they are widespread, appearing in a wide variety of habitats throughout the No ...
s, raccoons,
weasels Weasels are mammals of the genus ''Mustela'' of the family Mustelidae. The genus ''Mustela'' includes the least weasels, polecats, stoats, ferrets and European mink. Members of this genus are small, active predators, with long and slend ...
, and
seals Seals may refer to: * Pinniped, a diverse group of semi-aquatic marine mammals, many of which are commonly called seals, particularly: ** Earless seal, or "true seal" ** Fur seal * Seal (emblem), a device to impress an emblem, used as a means of a ...
. Members of this group are found worldwide and with immense diversity in their diet, behavior, and morphology.


Etymology

The word ''carnivore'' is derived from
Latin Latin (, or , ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through ...
''carō'' (stem ''carn-'') 'flesh' and ''vorāre'' 'to devour', and refers to any meat-eating organism.


Phylogeny

The oldest known carnivoran line mammals (
Carnivoramorpha Carnivoramorpha ("carnivoran-like forms") is a clade of placental mammals that includes the modern order Carnivora and its extinct stem-relatives.Bryant, H.N., and M. Wolson (2004“Phylogenetic Nomenclature of Carnivoran Mammals.”''First Inte ...
) appeared in
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and th ...
6 million years after the
Cretaceous–Paleogene extinction event The Cretaceous–Paleogene (K–Pg) extinction event (also known as the Cretaceous–Tertiary extinction) was a sudden mass extinction of three-quarters of the plant and animal species on Earth, approximately 66 million years ago. With the ...
. These early ancestors of carnivorans would have resembled small weasel or genet-like mammals, occupying a nocturnal shift on the forest floor or in the trees, as other groups of mammals like the
mesonychian Mesonychia ("middle claws") is an extinct taxon of small- to large-sized carnivorous ungulates related to artiodactyls. Mesonychids first appeared in the early Paleocene, went into a sharp decline at the end of the Eocene, and died out entirel ...
s and later the creodonts were occupying the megafaunal faunivorous niche. However, following the extinction of mesonychians and the oxyaenid creodonts at the end of the Eocene, carnivorans quickly moved into this niche, with forms like the nimravids being the dominant large-bodied ambush predators during the Oligocene alongside the hyaenodont creodonts (which similarly produced larger, more open-country forms at the start of the Oligocene). By the time
Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...
epoch appeared, most if not all of the major lineages and families of carnivorans had diversified and become the most dominant group of large terrestrial predators in Eurasia and North America, with various lineages being successful in megafaunal faunivorous niches at different intervals during the Miocene and later epochs.


Systematics


Evolution

The order Carnivora belongs to a group of mammals known as
Laurasiatheria Laurasiatheria ("laurasian beasts") is a superorder of placental mammals that groups together true insectivores ( eulipotyphlans), bats ( chiropterans), carnivorans, pangolins ( pholidotes), even-toed ungulates ( artiodactyls), odd-toed ungulat ...
, which also includes other groups such as bats and
ungulate Ungulates ( ) are members of the diverse clade Ungulata which primarily consists of large mammals with hooves. These include odd-toed ungulates such as horses, rhinoceroses, and tapirs; and even-toed ungulates such as cattle, pigs, giraffes, ...
s. Within this group the carnivorans are placed in the clade Ferae. Ferae includes the closest extant relative of carnivorans, the
pangolin Pangolins, sometimes known as scaly anteaters, are mammals of the order Pholidota (, from Ancient Greek ϕολιδωτός – "clad in scales"). The one extant family, the Manidae, has three genera: '' Manis'', ''Phataginus'', and '' Smuts ...
s, as well as several extinct groups of mostly
Paleogene The Paleogene ( ; also spelled Palaeogene or Palæogene; informally Lower Tertiary or Early Tertiary) is a geologic period and system that spans 43 million years from the end of the Cretaceous Period million years ago ( Mya) to the beginning o ...
carnivorous placentals such as the creodonts, the arctocyonians, and
mesonychian Mesonychia ("middle claws") is an extinct taxon of small- to large-sized carnivorous ungulates related to artiodactyls. Mesonychids first appeared in the early Paleocene, went into a sharp decline at the end of the Eocene, and died out entirel ...
s. The creodonts were originally thought of as the sister taxon to the carnivorans, perhaps even ancestral to, based on the presence of the carnassial teeth, but the nature of the carnassial teeth is different between the two groups. In carnivorans the carnassials are positioned near the front of the molar row, while in the creodonts they are positioned near the back of the molar row, and this suggests a separate evolutionary history and an order-level distinction. In addition recent phylogenetic analysis suggests that creodonts are more closely related to pangolins while mesonychians might be the sister group to carnivorans and their stem-relatives. The closest stem-carnivorans are the miacoids. The miacoids include the families Viverravidae and Miacidae, and together the Carnivora and Miacoidea form the stem-clade
Carnivoramorpha Carnivoramorpha ("carnivoran-like forms") is a clade of placental mammals that includes the modern order Carnivora and its extinct stem-relatives.Bryant, H.N., and M. Wolson (2004“Phylogenetic Nomenclature of Carnivoran Mammals.”''First Inte ...
. The miacoids were small, genet-like carnivoramorphs that occupy a variety of niches such as terrestrial and arboreal habitats. Recent studies have shown a supporting amount of evidence that Miacoidea is an evolutionary grade of carnivoramorphs that, while viverravids are monophyletic basal group, the miacids are paraphyletic in respect to Carnivora (as shown in the phylogeny below).Bryant, H.N., and M. Wolson (2004
“Phylogenetic Nomenclature of Carnivoran Mammals.”
''First International Phylogenetic Nomenclature Meeting''. Paris, Museum National d’Histoire Naturelle
Carnivoramorpha as a whole first appeared in the
Paleocene The Paleocene, ( ) or Palaeocene, is a geological epoch that lasted from about 66 to 56 million years ago (mya). It is the first epoch of the Paleogene Period in the modern Cenozoic Era. The name is a combination of the Ancient Greek ''pala ...
of North America about 60 million years ago. Crown carnivorans first appeared around 42 million years ago in the
Middle Eocene The Eocene ( ) Epoch is a geological epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period in the modern Cenozoic Era. The name ''Eocene'' comes from the Ancient Greek (''ēṓs'', "da ...
. Their molecular phylogeny shows the extant Carnivora are a
monophyletic In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic gr ...
group, the
crown group In phylogenetics, the crown group or crown assemblage is a collection of species composed of the living representatives of the collection, the most recent common ancestor of the collection, and all descendants of the most recent common ancestor ...
of the
Carnivoramorpha Carnivoramorpha ("carnivoran-like forms") is a clade of placental mammals that includes the modern order Carnivora and its extinct stem-relatives.Bryant, H.N., and M. Wolson (2004“Phylogenetic Nomenclature of Carnivoran Mammals.”''First Inte ...
. From there carnivorans have split into two clades based on the composition of the bony structures that surround the middle ear of the skull, the cat-like feliforms and the dog-like caniforms. In feliforms, the auditory bullae are double-chambered, composed of two bones joined by a
septum In biology, a septum (Latin for ''something that encloses''; plural septa) is a wall, dividing a cavity or structure into smaller ones. A cavity or structure divided in this way may be referred to as septate. Examples Human anatomy * Interat ...
. Caniforms have single-chambered or partially divided auditory bullae, composed of a single bone. Initially the early representatives of carnivorans were small as the creodonts (specifically, the oxyaenids) and mesonychians dominated the apex predator niches during the Eocene, but in the Oligocene carnivorans became a dominant group of apex predators with the nimravids, and by the
Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...
most of the extant carnivoran families have diversified and become the primary terrestrial predators in the Northern Hemisphere. The phylogenetic relationships of the carnivorans are shown in the following cladogram:


Classification of the extant carnivorans

In 1758 the Swedish
botanist Botany, also called , plant biology or phytology, is the science of plant life and a branch of biology. A botanist, plant scientist or phytologist is a scientist who specialises in this field. The term "botany" comes from the Ancient Greek wo ...
Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, ...
placed all carnivorans known at the time into the group Ferae (not to be confused with the modern concept of Ferae which also includes pangolins) in the tenth edition of his book ''
Systema Naturae ' (originally in Latin written ' with the ligature æ) is one of the major works of the Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the system, now known as binomial ...
''. He recognized six genera: ''
Canis ''Canis'' is a genus of the Caninae which includes multiple extant species, such as wolves, dogs, coyotes, and golden jackals. Species of this genus are distinguished by their moderate to large size, their massive, well-developed skulls and de ...
'' (canids and hyaenids), ''
Phoca ''Phoca'' ( ) is a genus of the earless seals, within the family Phocidae, source of the French name for seal, 'phoque'. It now contains just two species, the common seal (or harbour seal) and the spotted seal (or largha seal). Several specie ...
'' (pinnipeds), ''
Felis ''Felis'' is a genus of small and medium-sized cat species native to most of Africa and south of 60° latitude in Europe and Asia to Indochina. The genus includes the domestic cat. The smallest ''Felis'' species is the black-footed cat with a h ...
'' (felids), ''
Viverra ''Viverra'' is a mammalian genus that was first nominated and described by Carl Linnaeus in 1758 as comprising several species including the large Indian civet (''V. zibetha''). The genus was subordinated to the viverrid family by John Edward ...
'' (viverrids, herpestids, and mephitids), '' Mustela'' (non-badger mustelids), '' Ursus'' (ursids, large species of mustelids, and procyonids). It wasn't until 1821 that the English writer and traveler
Thomas Edward Bowdich Thomas Edward Bowdich (20 June 179110 January 1824) was an English traveller and author. Life Bowdich was born at Bristol and educated at Bristol Grammar School. In 1813, he married Sarah Wallis, who shared his subsequent career. In 1814, thro ...
gave the group its modern and accepted name.Bowditch, T. E. 1821. An analysis of the natural classifications of Mammalia for the use of students and travelers J. Smith Paris. 115. (refer pages 24, 33) Initially the modern concept of Carnivora was divided into two suborders: the terrestrial Fissipedia and the marine
Pinnipedia Pinnipeds (pronounced ), commonly known as seals, are a widely distributed and diverse clade of carnivorous, fin-footed, semiaquatic, mostly marine mammals. They comprise the extant families Odobenidae (whose only living member is the walrus ...
. Below is the classification of how the extant families were related to each other after American paleontologist
George Gaylord Simpson George Gaylord Simpson (June 16, 1902 – October 6, 1984) was an American paleontologist. Simpson was perhaps the most influential paleontologist of the twentieth century, and a major participant in the modern synthesis, contributing '' Tempo ...
in 1945: * Order Carnivora Bowdich, 1821 ** Suborder
Fissipedia Fissipedia is a former biological suborder comprising the largely land-based families of the order Carnivora. By and large, members of this suborder are meat-eaters, with the giant panda and red panda (each exclusively herbivorous) being t ...
Blumenbach, 1791 *** Superfamily
Canoidea Caniformia is a suborder within the order Carnivora consisting of "dog-like" carnivorans. They include dogs ( wolves, foxes, etc.), bears, raccoons, and mustelids. The Pinnipedia (seals, walruses and sea lions) are also assigned to this group ...
G. Fischer de Waldheim, 1817 **** Family
Canidae Canidae (; from Latin, '' canis'', " dog") is a biological family of dog-like carnivorans, colloquially referred to as dogs, and constitutes a clade. A member of this family is also called a canid (). There are three subfamilies found withi ...
G. Fischer de Waldheim, 1817 – dogs **** Family
Ursidae Bears are carnivoran mammals of the family Ursidae. They are classified as caniforms, or doglike carnivorans. Although only eight species of bears are extant, they are widespread, appearing in a wide variety of habitats throughout the Nort ...
G. Fischer de Waldheim, 1817 – bears **** Family
Procyonidae Procyonidae is a New World family of the order Carnivora. It comprises the raccoons, ringtails, cacomistles, coatis, kinkajous, olingos, and olinguitos. Procyonids inhabit a wide range of environments and are generally omnivorous. Charac ...
Bonaparte, 1850 – raccoons and pandas **** Family
Mustelidae The Mustelidae (; from Latin ''mustela'', weasel) are a family of carnivorous mammals, including weasels, badgers, otters, ferrets, martens, minks and wolverines, among others. Mustelids () are a diverse group and form the largest family in th ...
G. Fischer de Waldheim, 1817 – skunks, badgers, otters and weasels *** Superfamily Feloidea G. Fischer de Waldheim, 1817 **** Family
Viverridae Viverridae is a family (biology), family of small to medium-sized, feliform mammals. The viverrids () comprise 33 species placed in 14 genera. This family was named and first described by John Edward Gray in 1821. Viverrids occur all over Africa, ...
J. E. Gray, 1821 – civets and mongooses **** Family Hyaenidae J. E. Gray, 1821 – hyenas **** Family
Felidae Felidae () is the family of mammals in the order Carnivora colloquially referred to as cats, and constitutes a clade. A member of this family is also called a felid (). The term "cat" refers both to felids in general and specifically to the dom ...
G. Fischer de Waldheim, 1817 – cats ** Suborder
Pinnipedia Pinnipeds (pronounced ), commonly known as seals, are a widely distributed and diverse clade of carnivorous, fin-footed, semiaquatic, mostly marine mammals. They comprise the extant families Odobenidae (whose only living member is the walrus ...
Iliger, 1811 *** Family Otariidae J. E. Gray, 1825 – eared seals *** Family Odobenidae J. A. Allen, 1880 – walrus *** Family Phocidae J. E. Gray, 1821 – earless seals Since then, however, the methods in which mammalogists use to assess the phylogenetic relationships among the carnivoran families has been improved with using more complicated and intensive incorporation of genetics, morphology and the fossil record. Research into Carnivora phylogeny since 1945 has found Fisspedia to be paraphlyetic in respect to Pinnipedia, with pinnipeds being either more closely related to bears or to weasels. The small carnivoran families Viverridae, Procyonidae, and Mustelidae have been found to be
polyphyletic A polyphyletic group is an assemblage of organisms or other evolving elements that is of mixed evolutionary origin. The term is often applied to groups that share similar features known as homoplasies, which are explained as a result of conver ...
: * Mongooses and a handful of Malagasy endemic species are found to be in a clade with hyenas, with the Malagasy species being in their own family
Eupleridae Eupleridae is a family of carnivorans endemic to Madagascar and comprising 10 known living species in seven genera, commonly known as euplerids, Malagasy mongooses or Malagasy carnivorans. The best known species is the fossa (''Cryptoproct ...
.Anne D. Yoder and John J. Flynn 2003
Origin of Malagasy Carnivora
/ref>Yoder, A., M. Burns, S. Zehr, T. Delefosse, G. Veron, S. Goodman, J. Flynn. 2003
Single origin of Malagasy Carnivora from an African ancestor – Letters to Nature
/ref>Philippe Gaubert, W. Chris Wozencraft, Pedro Cordeiro-Estrela and Géraldine Veron. 2005 – Mosaics of Convergences and Noise in Morphological Phylogenies: What's in a Viverrid-Like Carnivoran? * The African palm civet is a basal cat-like carnivoran. * The linsang is more closely related to cats. * Pandas are not procyonids nor are they a natural grouping. The giant panda is a true bear while the red panda is a distinct family. * Skunks and stink badgers are placed in their own family, and are the sister group to a clade containing Ailuridae, Procyonidae and Mustelidae ''sensu stricto''. Below is a table chart of the extant carnivoran families and number of extant species recognized by various authors of the first and fourth volumes of ''Handbook of the Mammals of the World'' published in 2009 and 2014 respectively:


Anatomy


Skull

The canine teeth are usually large and conical. The canines are thick and stress resistant. All of the terrestrial species of carnivorans have three
incisor Incisors (from Latin ''incidere'', "to cut") are the front teeth present in most mammals. They are located in the premaxilla above and on the mandible below. Humans have a total of eight (two on each side, top and bottom). Opossums have 18, wher ...
s on each side of each jaw (the exception is the
sea otter The sea otter (''Enhydra lutris'') is a marine mammal native to the coasts of the northern and eastern North Pacific Ocean. Adult sea otters typically weigh between , making them the heaviest members of the weasel family, but among the smal ...
(''Enhydra lutris'') which only has two lower incisor teeth). The third molar has been lost. The carnassial pair is made up of the fourth upper premolar and the first lower molar teeth. Like most mammals, the dentition is
heterodont In anatomy, a heterodont (from Greek, meaning 'different teeth') is an animal which possesses more than a single tooth morphology. In vertebrates, heterodont pertains to animals where teeth are differentiated into different forms. For exampl ...
, though in some species, such as the
aardwolf The aardwolf (''Proteles cristata'') is an insectivorous species of hyena, native to East and Southern Africa. Its name means "earth-wolf" in Afrikaans and Dutch. It is also called maanhaar-jackal (Afrikaans for " mane-jackal"), termite-eat ...
(''Proteles cristata''), the teeth have been greatly reduced and the cheek teeth are specialised for eating insects. In pinnipeds, the teeth are
homodont In anatomy, a heterodont (from Greek, meaning 'different teeth') is an animal which possesses more than a single tooth morphology. In vertebrates, heterodont pertains to animals where teeth are differentiated into different forms. For example, ...
as they have evolved to grasp or catch fish, and the cheek teeth are often lost. In bears and raccoons the carnassial pair is secondarily reduced. The skulls are heavily built with a strong
zygomatic arch In anatomy, the zygomatic arch, or cheek bone, is a part of the skull formed by the zygomatic process of the temporal bone (a bone extending forward from the side of the skull, over the opening of the ear) and the temporal process of the zygo ...
. Often a
sagittal crest A sagittal crest is a ridge of bone running lengthwise along the midline of the top of the skull (at the sagittal suture) of many mammalian and reptilian skulls, among others. The presence of this ridge of bone indicates that there are exception ...
is present, sometimes more evident in sexually dimorphic species such as
sea lion Sea lions are pinnipeds characterized by external ear flaps, long foreflippers, the ability to walk on all fours, short and thick hair, and a big chest and belly. Together with the fur seals, they make up the family Otariidae, eared seals. ...
s and
fur seal Fur seals are any of nine species of pinnipeds belonging to the subfamily Arctocephalinae in the family '' Otariidae''. They are much more closely related to sea lions than true seals, and share with them external ears (pinnae), relatively l ...
s, though it has also been greatly reduced in some small carnivorans. The
braincase In human anatomy, the neurocranium, also known as the braincase, brainpan, or brain-pan is the upper and back part of the skull, which forms a protective case around the brain. In the human skull, the neurocranium includes the calvaria or skul ...
is enlarged with the frontoparietal bone at the front. In most species, the eyes are at the front of the face. In caniforms, the rostrum is usually long with many teeth, while in feliforms it is shorter with fewer teeth. The carnassial teeth of feliforms are generally more sectional than those of caniforms. The turbinates are large and complex in comparison to other mammals, providing a large surface area for olfactory receptors.


Postcranial region

Aside from an accumulation of characteristics in the dental and cranial features, not much of their overall anatomy unites carnivorans as a group. All species of carnivorans are
quadrupedal Quadrupedalism is a form of locomotion where four limbs are used to bear weight and move around. An animal or machine that usually maintains a four-legged posture and moves using all four limbs is said to be a quadruped (from Latin ''quattuor ...
and most have five digits on the front feet and four digits on the back feet. In terrestrial carnivorans, the feet have soft pads. The feet can either be
digitigrade In terrestrial vertebrates, digitigrade () locomotion is walking or running on the toes (from the Latin ''digitus'', 'finger', and ''gradior'', 'walk'). A digitigrade animal is one that stands or walks with its toes (metatarsals) touching the groun ...
as seen in cats, hyenas and dogs or
plantigrade 151px, Portion of a human skeleton, showing plantigrade habit In terrestrial animals, plantigrade locomotion means walking with the toes and metatarsals flat on the ground. It is one of three forms of locomotion adopted by terrestrial mammals. ...
as seen in bears, skunks, raccoons, weasels, civets and mongooses. In pinnipeds, the limbs have been modified into
flippers Flipper may refer to: Common meanings *Flipper (anatomy), a forelimb of an aquatic animal, useful for steering and/or propulsion in water *Alternate name for a swimfin, footwear that boosts human swimming efficiency * Flipper (cricket), a type of ...
. Unlike
cetacean Cetacea (; , ) is an infraorder of aquatic mammals that includes whales, dolphins, and porpoises. Key characteristics are their fully aquatic lifestyle, streamlined body shape, often large size and exclusively carnivorous diet. They propel th ...
s and sirenians, which have fully functional
tail The tail is the section at the rear end of certain kinds of animals’ bodies; in general, the term refers to a distinct, flexible appendage to the torso. It is the part of the body that corresponds roughly to the sacrum and coccyx in mammal ...
s to help them swim, pinnipeds use their limbs underwater to swim. Earless seals use their back flippers; sea lions and fur seals use their front flippers, and the
walrus The walrus (''Odobenus rosmarus'') is a large flippered marine mammal with a discontinuous distribution about the North Pole in the Arctic Ocean and subarctic seas of the Northern Hemisphere. The walrus is the only living species in the fami ...
use all of their limbs. As a result, pinnipeds have significantly shorter tails than other carnivorans. Aside from the pinnipeds, dogs, bears, hyenas, and cats all have distinct and recognizable appearances. Dogs are usually
cursorial A cursorial organism is one that is adapted specifically to run. An animal can be considered cursorial if it has the ability to run fast (e.g. cheetah) or if it can keep a constant speed for a long distance (high endurance). "Cursorial" is often u ...
mammals and are gracile in appearance, often relying on their teeth to hold prey; bears are much larger and rely on their physical strength to forage for food. Compared to dogs and bears, cats have longer and stronger forelimbs armed with retractable claws to hold on to prey. Hyenas are dog-like feliforms that have sloping backs due to their front legs being longer than their hind legs. The raccoon family and
red panda The red panda (''Ailurus fulgens''), also known as the lesser panda, is a small mammal native to the eastern Himalayas and southwestern China. It has dense reddish-brown fur with a black belly and legs, white-lined ears, a mostly white muzzle ...
are small, bear-like carnivorans with long tails. The other small carnivoran families Nandiniidae, Prionodontidae,
Viverridae Viverridae is a family (biology), family of small to medium-sized, feliform mammals. The viverrids () comprise 33 species placed in 14 genera. This family was named and first described by John Edward Gray in 1821. Viverrids occur all over Africa, ...
, Herpestidae, Eupleridae, Mephitidae and Mustelidae have through
convergent evolution Convergent evolution is the independent evolution of similar features in species of different periods or epochs in time. Convergent evolution creates analogous structures that have similar form or function but were not present in the last com ...
maintained the small, ancestral appearance of the miacoids, though there is some variation seen such as the robust and stout physicality of
badger Badgers are short-legged omnivores in the family Mustelidae (which also includes the otters, wolverines, martens, minks, polecats, weasels, and ferrets). Badgers are a polyphyletic rather than a natural taxonomic grouping, being united by ...
s and the
wolverine The wolverine (), (''Gulo gulo''; ''Gulo'' is Latin for " glutton"), also referred to as the glutton, carcajou, or quickhatch (from East Cree, ''kwiihkwahaacheew''), is the largest land-dwelling species of the family Mustelidae. It is a musc ...
(''Gulo gulo''). Male carnivorans usually have
bacula Bacula is an open-source, enterprise-level computer backup system for heterogeneous networks. It is designed to automate backup tasks that had often required intervention from a systems administrator or computer operator. Bacula supports Li ...
, though they are absent in
hyena Hyenas, or hyaenas (from Ancient Greek , ), are feliform carnivoran mammals of the family Hyaenidae . With only four extant species (each in its own genus), it is the fifth-smallest family in the Carnivora and one of the smallest in the cl ...
s and
binturong The binturong (''Arctictis binturong'') (, ), also known as the bearcat, is a viverrid native to South and Southeast Asia. It is uncommon in much of its range, and has been assessed as Vulnerable on the IUCN Red List because of a declining pop ...
s. The length and density of the fur vary depending on the environment that the species inhabits. In warm climate species, the fur is often short in length and lighter. In cold climate species, the fur is either dense or long, often with an oily substance that helps to retain heat. The pelage coloration differs between species, often including black, white, orange, yellow, red, and many shades of grey and brown. Some are striped, spotted, blotched, banded, or otherwise boldly patterned. There seems to be a correlation between habitat and color pattern; for example spotted or banded species tend to be found in heavily forested environments. Some species like the grey wolf are polymorphic with different individual having different coat colors. The
arctic fox The Arctic fox (''Vulpes lagopus''), also known as the white fox, polar fox, or snow fox, is a small fox native to the Arctic regions of the Northern Hemisphere and common throughout the Arctic tundra biome. It is well adapted to living in ...
(''Vulpes lagopus'') and the
stoat The stoat (''Mustela erminea''), also known as the Eurasian ermine, Beringian ermine and ermine, is a mustelid native to Eurasia and the northern portions of North America. Because of its wide circumpolar distribution, it is listed as Least Conc ...
(''Mustela erminea'') have fur that changes from white and dense in the winter to brown and sparse in the summer. In pinnipeds,
polar bear The polar bear (''Ursus maritimus'') is a hypercarnivorous bear whose native range lies largely within the Arctic Circle, encompassing the Arctic Ocean, its surrounding seas and surrounding land masses. It is the largest extant bear spec ...
s, and sea otters a thick insulating layer of blubber helps maintain their body temperature.


Relationship with humans

Carnivorans are arguably the group of mammals of most interest to humans. The dog is noteworthy for not only being the first species of carnivoran to be domesticated, but also the first species of any taxon. In the last 10,000 to 12,000 years humans have selectively bred dogs for a variety of different tasks and today there are well over 400 breeds. The
cat The cat (''Felis catus'') is a domestic species of small carnivorous mammal. It is the only domesticated species in the family Felidae and is commonly referred to as the domestic cat or house cat to distinguish it from the wild members of ...
is another domesticated carnivoran and it is today considered one of the most successful species on the planet, due to their close proximity to humans and the popularity of cats as pets. Many other species are popular, and they are often
charismatic megafauna Charismatic megafauna are animal species that are large—in the relevant category that they represent—with symbolic value or widespread popular appeal, and are often used by environmental activists to gain public support for environmentalist go ...
. Many civilizations have incorporated a species of carnivoran into their culture such as the
lion The lion (''Panthera leo'') is a large cat of the genus '' Panthera'' native to Africa and India. It has a muscular, broad-chested body; short, rounded head; round ears; and a hairy tuft at the end of its tail. It is sexually dimorphic; adu ...
, viewed as royalty. Yet many species such as
wolves The wolf (''Canis lupus''; : wolves), also known as the gray wolf or grey wolf, is a large canine native to Eurasia and North America. More than thirty subspecies of ''Canis lupus'' have been recognized, and gray wolves, as popularly un ...
and the
big cat The term "big cat" is typically used to refer to any of the five living members of the genus ''Panthera'', namely the tiger, lion, jaguar, leopard, and snow leopard. Despite enormous differences in size, various cat species are quite similar ...
s have been broadly hunted, resulting in extirpation in some areas. Habitat loss and human encroachment as well as
climate change In common usage, climate change describes global warming—the ongoing increase in global average temperature—and its effects on Earth's climate system. Climate change in a broader sense also includes previous long-term changes to ...
have been the primary cause of many species going into decline. Four species of carnivorans have gone extinct since the 1600s:
Falkland Island Wolf Falkland may refer to: * Falkland, British Columbia, a community in Canada * Falkland, Nova Scotia, a community in Canada * Falkland Islands, an archipelago in the south Atlantic Ocean * Falkland, Fife, a former burgh in Fife, Scotland ** Falkla ...
(''Dusicyon australis'') in 1876; the Sea Mink (''Neogale macrodon'') in 1894; the Japanese Sea Lion (''Zalophus japonicus'') in 1951 and the Caribbean Monk Seal (''Neomonachus tropicalis'') in 1952. Some species such as the
red fox The red fox (''Vulpes vulpes'') is the largest of the true foxes and one of the most widely distributed members of the Order (biology), order Carnivora, being present across the entire Northern Hemisphere including most of North America, Europe ...
(''Vulpes vulpes'') and
stoat The stoat (''Mustela erminea''), also known as the Eurasian ermine, Beringian ermine and ermine, is a mustelid native to Eurasia and the northern portions of North America. Because of its wide circumpolar distribution, it is listed as Least Conc ...
(''Mustela erminea'') have been introduced to
Australasia Australasia is a region that comprises Australia, New Zealand and some neighbouring islands in the Pacific Ocean. The term is used in a number of different contexts, including geopolitically, physiogeographically, philologically, and ecologic ...
and have caused many native species to become endangered or even extinct.


See also

*
List of carnivorans File:Carnivora_portraits.jpg, 300px, Various carnivorans, with feliforms to the left, and caniforms to the right rect 580 485 0 0 Cheetah rect 582 0 1191 485 Grizzly bear rect 0 486 577 972 Spotted hyena rect 581 487 1190 971 Wolf rect 0 975 577 ...
*
List of carnivorans by population This is a list of estimated global populations of Carnivora species. This list is not comprehensive, as not all carnivorans have had their numbers quantified. See also *Lists of organisms by population *Lists of mammals by population Refer ...


References


External links


High-Resolution Images of Carnivore Brains
{{Authority control Mammal orders Extant Lutetian first appearances Taxa named by Thomas Edward Bowdich