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''Camelops''Being occasionally called ''Western Camel'' or ''Yesterday's Camel''. is an extinct
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
of
camel A camel (from: la, camelus and grc-gre, κάμηλος (''kamēlos'') from Hebrew or Phoenician: גָמָל ''gāmāl''.) is an even-toed ungulate in the genus ''Camelus'' that bears distinctive fatty deposits known as "humps" on its back. ...
s that lived in
North North is one of the four compass points or cardinal directions. It is the opposite of south and is perpendicular to east and west. ''North'' is a noun, adjective, or adverb indicating direction or geography. Etymology The word ''north ...
and
Central America Central America ( es, América Central or ) is a subregion of the Americas. Its boundaries are defined as bordering the United States to the north, Colombia to the south, the Caribbean Sea to the east, and the Pacific Ocean to the west. ...
, ranging from Alaska to Guatemala, from the middle
Pliocene The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58Pleistocene The Pleistocene ( , often referred to as the ''Ice age'') is the geological Epoch (geology), epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fina ...
. It is more closely related to the
Old World The "Old World" is a term for Afro-Eurasia that originated in Europe , after Europeans became aware of the existence of the Americas. It is used to contrast the continents of Africa, Europe, and Asia, which were previously thought of by thei ...
dromedary The dromedary (''Camelus dromedarius'' or ;), also known as the dromedary camel, Arabian camel, or one-humped camel, is a large even-toed ungulate, of the genus '' Camelus'', with one hump on its back. It is the tallest of the three species o ...
and bactrian and
wild bactrian camel The wild Bactrian camel (''Camelus ferus'') is a critically endangered species of camel living in parts of northwestern China and southwestern Mongolia. It is closely related to the Bactrian camel (''Camelus bactrianus''). Both are large, doubl ...
s than the
New World The term ''New World'' is often used to mean the majority of Earth's Western Hemisphere, specifically the Americas."America." ''The Oxford Companion to the English Language'' (). McArthur, Tom, ed., 1992. New York: Oxford University Press, p. ...
guanaco, vicuña,
alpaca The alpaca (''Lama pacos'') is a species of South American camelid mammal. It is similar to, and often confused with, the llama. However, alpacas are often noticeably smaller than llamas. The two animals are closely related and can success ...
and
llama The llama (; ) (''Lama glama'') is a domesticated South American camelid, widely used as a meat and pack animal by Andean cultures since the Pre-Columbian era. Llamas are social animals and live with others as a herd. Their wool is soft ...
; making it a true camel of the Camelini
tribe The term tribe is used in many different contexts to refer to a category of human social group. The predominant worldwide usage of the term in English is in the discipline of anthropology. This definition is contested, in part due to confl ...
. Its name is derived from the
Ancient Greek Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic pe ...
(, "camel") and (, "face"), i.e. "camel-face".


Taxonomy and evolution

The genus ''Camelops'' first appeared during the middle
Pliocene The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58Pleistocene The Pleistocene ( , often referred to as the ''Ice age'') is the geological Epoch (geology), epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fina ...
(around 11,000 years ago). Despite the fact that camels are popularly associated with the deserts of Asia and Africa, the family
Camelidae Camelids are members of the biological family Camelidae, the only currently living family in the suborder Tylopoda. The seven extant members of this group are: dromedary camels, Bactrian camels, wild Bactrian camels, llamas, alpacas, vicuñas ...
, which comprises camels and llamas, originated in North America during the middle
Eocene The Eocene ( ) Epoch is a geological epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period in the modern Cenozoic Era. The name ''Eocene'' comes from the Ancient Greek (''ēṓs'', ...
period, at least 44 Mya. Both the camel and
horse The horse (''Equus ferus caballus'') is a domesticated, one-toed, hoofed mammal. It belongs to the taxonomic family Equidae and is one of two extant subspecies of ''Equus ferus''. The horse has evolved over the past 45 to 55 million yea ...
families originated in the Americas and migrated into
Eurasia Eurasia (, ) is the largest continental area on Earth, comprising all of Europe and Asia. Primarily in the Northern and Eastern Hemispheres, it spans from the British Isles and the Iberian Peninsula in the west to the Japanese archipelag ...
via the Bering Strait. Modern camels are descended from the extinct genus '' Paracamelus'', which probably crossed the Bering land bridge into Asia between 7.5 and 6.5 Mya. The divergence between ''Paracamelus'' and ''Camelops'' occurred about 11–10 Mya. ''Paracamelus'' would continue to live in North America as the High Arctic camel until the middle Pleistocene. During Pleistocene warm periods, a smaller morph of ''Camelops'' inhabited
Alaska Alaska ( ; russian: Аляска, Alyaska; ale, Alax̂sxax̂; ; ems, Alas'kaaq; Yup'ik: ''Alaskaq''; tli, Anáaski) is a state located in the Western United States on the northwest extremity of North America. A semi-exclave of the U ...
and northern
Yukon Yukon (; ; formerly called Yukon Territory and also referred to as the Yukon) is the smallest and westernmost of Canada's three territories. It also is the second-least populated province or territory in Canada, with a population of 43,964 as ...
. These specimens date to around 50–45 thousand years ago, and seem to have been extirpated from the area after this time, similar to the contemporaneous
mastodon A mastodon ( 'breast' + 'tooth') is any proboscidean belonging to the extinct genus ''Mammut'' (family Mammutidae). Mastodons inhabited North and Central America during the late Miocene or late Pliocene up to their extinction at the end of the ...
, the
ground sloth Ground sloths are a diverse group of extinct sloths in the mammalian superorder Xenarthra. The term is used to refer to all extinct sloths because of the large size of the earliest forms discovered, compared to existing tree sloths. The Caribb ...
''
Megalonyx ''Megalonyx'' ( Greek, "large claw") is an extinct genus of ground sloths of the family Megalonychidae, native to North America during the Pliocene and Pleistocene epochs. It became extinct during the Quaternary extinction event at the end ...
'', and the giant beaver '' Castoroides''. The skull of a ''Camelops'' specimen was found above the Glenns Ferry Formation in present-day
Idaho Idaho ( ) is a U.S. state, state in the Pacific Northwest region of the Western United States. To the north, it shares a small portion of the Canada–United States border with the province of British Columbia. It borders the states of Monta ...
in a thick layer of coarse gravel known as the Tauna Gravels. Above this layer of gravel is another layer of fine river channel sands, where the skull was found. The age of this fossil is as young as 2 million years old and perhaps even younger, which can be inferred because it is younger than the other fossils found at the Hagerman Fossil Beds National Monument. During the late
Oligocene The Oligocene ( ) is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that define the epoch are well identified but t ...
and early
Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...
periods, camels apparently underwent swift evolutionary change, resulting in several genera with different anatomical structures, ranging from those with short limbs, those with gazelle-like bodies, and giraffe-like camels with long legs and long necks. This rich diversity decreased until only a few species, such as ''Camelops hesternus'', remained in North America, before going extinct entirely around 11,000 years ago. By the end of the Pleistocene, with the extinction of '' Paracamelus'' and '' Titanotylopus'', ''Camelops'' was the only true camel remaining in North America and possibly both Americas. ''Camelops'' extinction was part of a larger North American extinction in which native
horses The horse (''Equus ferus caballus'') is a domesticated, one-toed, hoofed mammal. It belongs to the taxonomic family Equidae and is one of two extant subspecies of ''Equus ferus''. The horse has evolved over the past 45 to 55 million ...
,
mastodon A mastodon ( 'breast' + 'tooth') is any proboscidean belonging to the extinct genus ''Mammut'' (family Mammutidae). Mastodons inhabited North and Central America during the late Miocene or late Pliocene up to their extinction at the end of the ...
s, and other
camelid Camelids are members of the biological family Camelidae, the only currently living family in the suborder Tylopoda. The seven extant members of this group are: dromedary camels, Bactrian camels, wild Bactrian camels, llamas, alpacas, vicuñas, ...
s also died out. Possible causal factors for this megafaunal extinction include global climate change and hunting pressure from human beings. The mass extinction coincided roughly with the appearance of people belonging to the big game-hunting
Clovis culture The Clovis culture is a prehistoric Paleoamerican culture, named for distinct stone and bone tools found in close association with Pleistocene fauna, particularly two mammoths, at Blackwater Locality No. 1 near Clovis, New Mexico, in 1936 a ...
, who were prolific hunters with distinct fluted stone tools, which allowed for a spear shaft to be attached to the stone tool. Biochemical analyses have shown that Clovis tools were used in butchering camels.


Description

Because soft tissues are generally not preserved in the
fossil record A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
, it is not certain if ''Camelops'' possessed hump(s), like modern camels, or lacked ones, like modern Camelidaes of South America ( guanacos and vicuñas). One-humped camels are now known to have evolved from two-humped camels, but two-humps, as an evolutionary outcome, likely associated with arctic climates and two-humped camels presumably evolved into one-humped camels in warmer regions in Eurasia, while ''Camelops'' first appeared in southern North America and lived among both warmer and colder regions of the continent until early
Holocene The Holocene ( ) is the current geological epoch. It began approximately 11,650 cal years Before Present (), after the Last Glacial Period, which concluded with the Holocene glacial retreat. The Holocene and the preceding Pleistocene togeth ...
. ''C. hesternus'' had legs 20% longer than that of the
dromedary The dromedary (''Camelus dromedarius'' or ;), also known as the dromedary camel, Arabian camel, or one-humped camel, is a large even-toed ungulate, of the genus '' Camelus'', with one hump on its back. It is the tallest of the three species o ...
, and was about tall at the shoulder and weighed about .Anthony J. Stuart, 2021, ''Vanished Giants: The Lost World of the Ice Age'', "6.17 Yesterday's Camel: Camelops Hesternus", p.99,
University of Chicago Press The University of Chicago Press is the largest and one of the oldest university presses in the United States. It is operated by the University of Chicago and publishes a wide variety of academic titles, including '' The Chicago Manual of Style' ...


Palaeobiology

The species ranged widely from Alaska in the north to
Oregon Oregon () is a U.S. state, state in the Pacific Northwest region of the Western United States. The Columbia River delineates much of Oregon's northern boundary with Washington (state), Washington, while the Snake River delineates much of it ...
and
California California is a state in the Western United States, located along the Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the most populous U.S. state and the 3rd largest by area. It is also the m ...
in the west, southernmost Baja California Peninsula and
Mexico Mexico (Spanish language, Spanish: México), officially the United Mexican States, is a List of sovereign states, country in the southern portion of North America. It is borders of Mexico, bordered to the north by the United States; to the so ...
and
Guatemala Guatemala ( ; ), officially the Republic of Guatemala ( es, República de Guatemala, links=no), is a country in Central America. It is bordered to the north and west by Mexico; to the northeast by Belize and the Caribbean; to the east by Hon ...
in the south, and
Tennessee Tennessee ( , ), officially the State of Tennessee, is a landlocked U.S. state, state in the Southeastern United States, Southeastern region of the United States. Tennessee is the List of U.S. states and territories by area, 36th-largest by ...
in the east, and a notable number of fossils have been excavated among central North America such as at
Colorado Colorado (, other variants) is a state in the Mountain West subregion of the Western United States. It encompasses most of the Southern Rocky Mountains, as well as the northeastern portion of the Colorado Plateau and the western edge of the ...
and
California California is a state in the Western United States, located along the Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the most populous U.S. state and the 3rd largest by area. It is also the m ...
. Plant remains found in the teeth of the Rancho La Brea ''C. hesternus'' fossils further reveal that rather than being limited to grazing, this species likely ate mixed species of plants, including coarse shrubs growing in coastal southern California. ''Camelops'' probably could travel long distances, similar to modern camel species. Whether or not ''Camelops'' could survive for long periods without water, as with extant camels, is still unknown; this may have been an adaptation that occurred much later, after camelids migrated to Asia and Africa. The
creosote bush ''Larrea tridentata'', called creosote bush and greasewood as a plant, chaparral as a medicinal herb, and ''gobernadora'' (Spanish for "governess") in Mexico, due to its ability to secure more water by inhibiting the growth of nearby plants. In S ...
from the US-Mexican border has been speculated to be part of ''Camelopss diet, based on the fact that no local living ungulate eats it, yet it was readily consumed by the Arabian camels of the experimental United States Camel Corps in the mid-19th century.


Extinction

The last species of ''Camelops'' are hypothesized to have disappeared as a result of the Blitzkrieg model. This model presents the hypothesis that ''Camelops'', along with other North American
megafauna In terrestrial zoology, the megafauna (from Greek μέγας ''megas'' "large" and New Latin ''fauna'' "animal life") comprises the large or giant animals of an area, habitat, or geological period, extinct and/or extant. The most common thresho ...
, disappeared as new cultures of experienced and efficient hunters moved southeastward across the continent. The result of this migration and expansion of human populations was a significant reduction in range for the megafauna. Of the many ''Camelops'' specimens recovered in North America, only a small number demonstrate modification through human actions. Some specimens have been interpreted as having been killed by humans based on the presence of spirally fractured bone fragments. None of the reported ''Camelops'' sites has been associated with stone tools, however, which would be an indicator of possible human use. At many of these ''Camelops'' sites, no fossils have been found of carcasses that were evidently processed, but rather small fragments and pieces of remains. Researchers originally thought that ''Camelops'' species were in fact hunted and butchered by early humans in North America because of these reasons: the fragmenting of bones into shapes that look like tools, damage or weathering of the “working” edge of said tools, having attributes that were similar to the making of chopping tools, and scarred fragments from possible chopping tools. Further examination showed, though, that these assumptions were misguided, and that while humans did coexist and associate with ''Camelops'', human use has yet to be completely proven as the sole cause of extinction.


See also

*'' Aepycamelus'' *''
Eulamaops ''Eulamaops'' is an extinct genus of camelid, endemic to South America during the Pleistocene (Lujanian, 781,000—12,000 years ago), existing about . Fossil remains of ''Eulamaops'' have been found in the Luján Formation in Argentina ...
'' *''
Oxydactylus ''Oxydactylus'' is an extinct genus of camelid endemic to North America. It lived from the Late Oligocene to the Middle Miocene (28.4–13.7 mya), existing for approximately . The name is from the Ancient Greek οξύς (oxys, "sharp")and δ ...
'' *''
Poebrotherium ''Poebrotherium'' ( ) is an extinct genus of camelid, endemic to North America. They lived from the Eocene to Miocene epochs, 46.3—13.6 mya, existing for approximately . Discovery and history ''Poebrotherium'' was first named by scientist ...
'' *'' Procamelus'' *''
Protylopus ''Protylopus'' is an extinct genus of camel that lived during middle to late Eocene some 50-40 million years ago in North America. Along with being the oldest camel known, it was also the smallest, reaching a length of , and probably weighing a ...
'' *''
Stenomylus ''Stenomylus'' is an extinct genus of miniature camelid native to North America that died out around 30 million years ago. Its name is derived from the Greek (, "narrow") and (, "molar"). ''Stenomylus'' was extremely diminutive compared to ...
'' * Syrian camel, an extinct species that reached at least tall at the shoulder * Pleistocene megafauna * Snowmastodon Project


References

{{Taxonbar, from=Q133439 Prehistoric camelids Prehistoric even-toed ungulate genera Pliocene even-toed ungulates Pleistocene even-toed ungulates Piacenzian first appearances Holocene extinctions Pliocene mammals of North America Pleistocene mammals of North America Taxa named by Joseph Leidy Fossil taxa described in 1854