Cucujoidea is a

superfamily SUPERFAMILY is a database and search platform of structural and functional annotation for all proteins and genomes. It classifies amino acid sequences into known structural domains, especially into SCOP superfamilies. Domains are functional, str ...

beetles Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 describ ...

. This group formerly included all of the families now included in the superfamily Coccinelloidea. They include some fungus beetles and a diversity of lineages of "bark beetles" unrelated to the "true" bark beetles (

Scolytinae A bark beetle is the common name for the subfamily of beetles Scolytinae. Previously, this was considered a distinct family (Scolytidae), but is now understood to be a specialized clade of the "true weevil" family (Curculionidae). Although the ...

), which are weevils (superfamily Curculionoidea).

Morphology

The morphology of Cucujoidea is varied and there are no features uniting all members of the superfamily. Adults can be recognised by the procoxal cavities being internally open in most taxa, females having tarsal formula 5-5-5 and males 5-5-5 or 5-5-4 (rarely 4-4-4), females with tergite VIII concealed dorsally by tergite VII, and males with tergite X completely membraneous. Larvae have frontal arms usually lyriform, the mandible mesal surface usually with well-developed mola, a maxillary articulating area usually present, a hypopharyngeal sclerome usually present, and two pretarsal setae.

Taxonomy

According to a 2015 revision, the following 25 families make up superfamily Cucujoidea: *

Agapythidae ''Agapytho'' is the only genus of beetles in the family Agapythidae. The genus contains a single species, ''Agapytho foveicollis'', which is endemic to New Zealand. Larvae and adults are found on sooty mold Sooty mold (also spelled sooty mould ...

Sen Gupta and Crowson, 1969 * Boganiidae Sen Gupta & Crowson, 1966 *

Cavognathidae Cavognathidae is a family of beetles, in the superfamily Cucujoidea Cucujoidea is a superfamily of beetles. This group formerly included all of the families now included in the superfamily Coccinelloidea. They include some fungus beetles and ...

Sen Gupta & Crowson, 1966 * Cryptophagidae Kirby, 1826 – silken fungus beetles *

Cucujidae The Cucujidae, "flat bark beetles," are a family of distinctively flat beetles found worldwide (except Africa and Antarctica) under the bark of dead trees. The family has received considerable taxonomic attention in recent years and now consists ...

Latreille, 1802 –

flat bark beetle The Cucujidae, "flat bark beetles," are a family (biology), family of distinctively flat beetles found worldwide (except Africa and Antarctica) under the bark of dead trees. The family has received considerable taxonomic attention in recent years ...

s *

Cybocephalidae Cybocephalidae is a family of sap, bark and fungus beetles in the order Coleoptera with a wide global distribution. The type genus ''Cybocephalus'' has more than 200 species in it and the entire family has about 220 species in all. Many species ar ...

Jacquelin du Val, 1858 * Cyclaxyridae Gimmel, Leschen & Ślipiński, 2009 – sooty mould beetles * Erotylidae Latreille, 1802 – pleasing fungus beetles *

Helotidae Helotidae is a family of beetles, in the suborder Polyphaga. The family includes about five extant genera, ''Helota'' MacLeay, ''Neohelota'' Ohta, ''Afrohelotina'' Kirejtshuk, ''Metahelotella'' Kirejtshuk, and ''Strophohelota'' Kirejtshuk. Hel ...

Reitter, 1876 *

Hobartiidae Hobartiidae is a family of beetles, in the superfamily Cucujoidea. There are only two known genera '' Hobartius'' (Sen Gupta & Crowson, 1966) and '' Hydnobioides'' (Sen Gupta & Crowson, 1966), with six species, five of which are native to Austral ...

Sen Gupta & Crowson, 1966 * Kateretidae Erichson in Agassiz, 1846 – short-winged flower beetles (= Brachypteridae) * Laemophloeidae Ganglbauer 1899 –

lined flat bark beetle Laemophloeidae, "lined flat bark beetles," is a family in the superfamily Cucujoidea characterized by predominantly dorso-ventrally compressed bodies, head and pronotal discs bordered by ridges or grooves, and inverted male genitalia. Size range ...

s * Lamingtoniidae Sen Gupta & Crowson, 1966 * Monotomidae Laporte, 1840 –

root-eating beetle Monotomidae is a family of beetles in the superfamily Cucujoidea. The family is found worldwide, with approximately 240 species in 33 genera. The ecological habits of the family are diverse, with different members of the group being found under t ...

s * Myraboliidae Lawrence and Britton, 1991 * Nitidulidae Latreille, 1802 – sap beetles *

Passandridae Passandridae, the "parasitic flat bark beetles," are a family of beetles notable for being one of the very few beetle families with larvae that are, as far as known, exclusively ectoparasitic on the immature stages of other beetles and Hymenoptera ...

Erichson, 1845 –

parasitic flat bark beetle Parasitism is a close relationship between species, where one organism, the parasite, lives on or inside another organism, the host, causing it some harm, and is adapted structurally to this way of life. The entomologist E. O. Wilson has c ...

s * Phalacridae Leach, 1815 –

shining flower beetle The Phalacridae are a family of beetles commonly called the shining flower beetles, They are often found in composite flowers. They are oval-shaped, usually tan, and aboout 2 mm in length. Most species feed on fungus, although a number feed ...

s *

Phloeostichidae Phloeostichidae is a family of beetles in the superfamily Cucujoidea. They are typically found under the bark of dead trees. Larvae have been found to consume plant tissue and some fungi, while the adults appear to be exclusively fungivores. The ...

Reitter, 1911 *

Priasilphidae Priasilphidae is a family of beetles in the superfamily Cucujoidea. They have a Gondwanan distribution, with the three known genera '' Chileosilpha, Priasilpha'' and '' Priastichus'' being native to Chile, New Zealand and Tasmania respectively. ...

Crowson, 1973 *

Protocucujidae Protocucujidae is a family of beetles, in the superfamily Cucujoidea Cucujoidea is a superfamily of beetles. This group formerly included all of the families now included in the superfamily Coccinelloidea. They include some fungus beetles an ...

Crowson, 1954 * Silvanidae Kirby, 1937 – silvanid flat bark beetles *

Smicripidae Smicripidae is a family of beetles, in the superfamily Cucujoidea. The common name for this family is palmetto beetles. The family only has one extant genus, '' Smicrips,'' with six extant species native to tropical and subtropical regions of th ...

Horn, 1879 –

palmetto beetle Smicripidae is a family of beetles, in the superfamily Cucujoidea. The common name for this family is palmetto beetles. The family only has one extant genus, ''Smicrips,'' with six extant species native to tropical and subtropical regions of the ...

s *

Sphindidae Sphindidae is a family of beetles, in the suborder Polyphaga. They are also known as slime mold beetles due to their exclusive feeding on slime molds during adult and larval stages, other aspects of their life history are obscure. Palaeontologic ...

Jacquelin du Val, 1860 –

dry-fungus beetle Sphindidae is a family of beetles, in the suborder Polyphaga. They are also known as slime mold beetles due to their exclusive feeding on Slime mold, slime molds during adult and larval stages, other aspects of their life history are obscure. Pa ...

s *

Tasmosalpingidae ''Tasmosalpingus'' is the only genus in the beetle family Tasmosalpingidae (superfamily Cucujoidea). There are two species in ''Tasmosalpingus'', found in Australia in Tasmania, Victoria and New South Wales. Adults have been collected using mala ...

Lawrence and Britton, 1991 * Parandrexidae Kirejtshuk, 1994 (Middle Jurassic-Early Cretaceous)

Extinct genera

* ''

Alloterocucus ''Alloterocucus'' is an extinct genus of beetle. It is known from a single species, ''Alloterocucus atratus'', known from the mid Cretaceous (latest Albian-earliest Cenomanian) Burmese amber of Myanmar. It belongs to the clade Cucujiformia, pr ...

'' Li et al., 2022 Burmese amber, Myanmar, mid- Cretaceous (latest Albian- earliest

Cenomanian The Cenomanian is, in the ICS' geological timescale, the oldest or earliest age of the Late Cretaceous Epoch or the lowest stage of the Upper Cretaceous Series. An age is a unit of geochronology; it is a unit of time; the stage is a unit in the s ...

)

References

External links

* * Beetle superfamilies Taxa named by Pierre André Latreille {{Cucujoidea-stub