Conchoraptor Gracilis
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''Conchoraptor'' (meaning "
conch Conch ( , , ) is a common name of a number of different medium-to-large-sized sea snails. Conch shells typically have a high Spire (mollusc), spire and a noticeable siphonal canal (in other words, the shell comes to a noticeable point on both ...
plunderer") is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of
oviraptorid Oviraptoridae is a group of bird-like, herbivorous and omnivorous maniraptoran dinosaurs. Oviraptorids are characterized by their toothless, parrot-like beaks and, in some cases, elaborate crests. They were generally small, measuring between one ...
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutio ...
s that lived in
Asia Asia ( , ) is the largest continent in the world by both land area and population. It covers an area of more than 44 million square kilometres, about 30% of Earth's total land area and 8% of Earth's total surface area. The continent, which ...
during the
Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
epoch, about 70 million years ago. It is known from the Barun Goyot and Nemegt formations of
Mongolia Mongolia is a landlocked country in East Asia, bordered by Russia to the north and China to the south and southeast. It covers an area of , with a population of 3.5 million, making it the world's List of countries and dependencies by po ...
.


Discovery

When first discovered in the Red Beds of Hermiin Tsav of the Early
Maastrichtian The Maastrichtian ( ) is, in the International Commission on Stratigraphy (ICS) geologic timescale, the latest age (geology), age (uppermost stage (stratigraphy), stage) of the Late Cretaceous epoch (geology), Epoch or Upper Cretaceous series (s ...
Nemegt Formation The Nemegt Formation (also known as Nemegtskaya Svita) is a geological formation in the Gobi Desert of Mongolia, dating to the Late Cretaceous. The formation consists of river channel sediments and contains fossils of fish, turtles, crocodilians ...
by a Polish-Mongolian expedition in 1971, scientists believed that ''Conchoraptor'' was a juvenile ''
Oviraptor ''Oviraptor'' (; ) is a genus of oviraptorid dinosaur that lived in Asia during the Late Cretaceous period. The first remains were collected from the Djadokhta Formation of Mongolia in 1923 during a paleontological expedition led by Roy Chapman ...
'' and that the animal's missing crest would have begun to grow when the animal reached
sexual maturity Sexual maturity is the capability of an organism to reproduce. In humans, it is related to both puberty and adulthood. ''Puberty'' is the biological process of sexual maturation, while ''adulthood'', the condition of being socially recognized ...
. Further study of multiple skeletons showed that ''Conchoraptor'' belonged to a new genus. The hands of ''Conchoraptor'' were a major reason scientists decided to split it off from ''Oviraptor''. Anatomically the hands seemed to be an
evolution Evolution is the change in the heritable Phenotypic trait, characteristics of biological populations over successive generations. It occurs when evolutionary processes such as natural selection and genetic drift act on genetic variation, re ...
ary intermediate between those of ''
Ajancingenia ''Heyuannia'' ("from Heyuan") is a genus of oviraptorid dinosaur that lived in Asia during the Late Cretaceous epoch, in what is now China and Mongolia. It was the first oviraptorid found in China; most others were found in neighbouring Mongolia ...
'' and ''Oviraptor'', making it obvious that this animal was not a member of a known species. The
type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
of this new genus, ''Conchoraptor gracilis'', was described and named by Barsbold, in 1986. The generic name is derived from Greek ''konkhè'', "mussel", and
Latin Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...
''raptor'', "plunderer". This name reflects Barsbold's hypothesis that oviraptorids, rather than preying primarily upon eggs as had been traditionally thought, may have been specialized to feed on
mollusk Mollusca is a phylum of protostomic invertebrate animals, whose members are known as molluscs or mollusks (). Around 76,000  extant species of molluscs are recognized, making it the second-largest animal phylum after Arthropoda. The ...
s. The
specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
means "gracile" in Latin. The
holotype A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...
is MPC-D 100/20, a partial skull. Numerous specimens were formally referred to the species later, including some recovered from the
Barun Goyot Formation The Baruungoyot Formation (also known as Barun Goyot) is a geological formation dating to the Late Cretaceous Period. It is located within and is widely represented in the Gobi Desert Basin, in the Ömnögovi Province of Mongolia. Descriptio ...
.


Description

''Conchoraptor'' was a small dinosaur, only in length. Unlike many other
oviraptorid Oviraptoridae is a group of bird-like, herbivorous and omnivorous maniraptoran dinosaurs. Oviraptorids are characterized by their toothless, parrot-like beaks and, in some cases, elaborate crests. They were generally small, measuring between one ...
s, ''Conchoraptor'' lacked a head crest. Like its relatives, it lacked teeth. Instead of teeth, oviraptorids had powerful beaks, possibly adapted to crushing
mollusk Mollusca is a phylum of protostomic invertebrate animals, whose members are known as molluscs or mollusks (). Around 76,000  extant species of molluscs are recognized, making it the second-largest animal phylum after Arthropoda. The ...
shells. The skull of ''Conchoraptor'' was heavily pneumatised, with most of the bones hollow.


Classification

''Conchoraptor'' was by Barsbold assigned to the
Oviraptoridae Oviraptoridae is a group of bird-like, herbivorous and omnivorous maniraptoran dinosaurs. Oviraptorids are characterized by their toothless, parrot-like beaks and, in some cases, elaborate crests. They were generally small, measuring between one ...
in 1986. Recent analyses show it was a member of the oviraptorid subfamily Ingeniinae (now Heyuanninae). The
cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not s ...
below follows an analysis by Fanti ''et al.'', 2012.


Paleobiology


Senses

Internal connections between both ears functioned as resonance chambers and improved hearing.


Feeding

Originally it was suggested that Conchoraptor had a diet that primarily consisted of mollusks. A 2022 study of the bite force of ''Conchoraptor'' and comparisons with other oviraptorosaurs such as ''
Incisivosaurus ''Incisivosaurus'' ("incisor lizard") is a genus of small, probably herbivorous theropod dinosaurs from the early Cretaceous Period of what is now the People's Republic of China. The first specimen to be described (by Xu ''et al.'' in 2002), IVPP ...
'', ''
Citipati ''Citipati'' (; meaning "funeral pyre lord") is a genus of oviraptorid dinosaur that lived in Asia during the Late Cretaceous Period (geology), period, about 75 million to 71 million years ago. It is mainly known from the Ukhaa Tolgod locality ...
'', and ''
Khaan ''Khaan'' (; from Mongol 'lord') is an extinct genus of Oviraptoridae, oviraptorid dinosaur known from remains found in the Djadochta Formation of Mongolia, dating to the Late Cretaceous (Campanian, 75-71 million years ago). Description ' ...
'' instead suggests that ''Conchoraptor'' had a very strong bite force. The moderate jaw gape seen in oviraptorosaurs is indicative of herbivory in the majority of the group, but it is clear they were likely feeding on much tougher or more various types of vegetation than other herbivorous theropods in their environment, such as ornithomimosaurs and therizinosaurs were able to. The examinations suggest oviraptorosaurs may have been powerful-biting generalists or specialists that partook of niche partitioning both in body size and cranial function.


Paleoenvironment

The
Nemegt Formation The Nemegt Formation (also known as Nemegtskaya Svita) is a geological formation in the Gobi Desert of Mongolia, dating to the Late Cretaceous. The formation consists of river channel sediments and contains fossils of fish, turtles, crocodilians ...
of Mongolia, is estimated to date back to the
Maastrichtian The Maastrichtian ( ) is, in the International Commission on Stratigraphy (ICS) geologic timescale, the latest age (geology), age (uppermost stage (stratigraphy), stage) of the Late Cretaceous epoch (geology), Epoch or Upper Cretaceous series (s ...
stage Stage, stages, or staging may refer to: Arts and media Acting * Stage (theatre), a space for the performance of theatrical productions * Theatre, a branch of the performing arts, often referred to as "the stage" * ''The Stage'', a weekly Brit ...
, between 70 and 66 million years ago of the Late
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 143.1 to 66 mya (unit), million years ago (Mya). It is the third and final period of the Mesozoic Era (geology), Era, as well as the longest. At around 77.1 million years, it is the ...
period Period may refer to: Common uses * Period (punctuation) * Era, a length or span of time *Menstruation, commonly referred to as a "period" Arts, entertainment, and media * Period (music), a concept in musical composition * Periodic sentence (o ...
. During the Late Cretaceous period, the land that is now the Nemegt Formation had an arid environment with fields of sand
dune A dune is a landform composed of wind- or water-driven sand. It typically takes the form of a mound, ridge, or hill. An area with dunes is called a dune system or a dune complex. A large dune complex is called a dune field, while broad, flat ...
s and only intermittent
stream A stream is a continuous body of water, body of surface water Current (stream), flowing within the stream bed, bed and bank (geography), banks of a channel (geography), channel. Depending on its location or certain characteristics, a strea ...
s. It is slightly younger than the nearby
Djadochta Formation The Djadochta formation (sometimes transcribed and also known as Djadokhta, Djadokata, or Dzhadokhtskaya) is a highly fossiliferous geological formation in Central Asia, Gobi Desert, dating from the Late Cretaceous period, about 75 million ...
, and seems to have been slightly wetter. This formation is noteworthy for the exquisite preservation of small and fragile dinosaur skeletons, a rare occurrence considering that these fossils are typically broken up and dispersed when found in other rock formations.Novacek, M. J., 1996. "Dinosaurs of the Flaming. Cliffs" Anchor/Doubleday. 367 p. ''Conchoraptor'' shared its habitat with other oviraptorosaurs including ''
Avimimus ''Avimimus'' ( ), meaning "bird mimic" (Latin ''avis'' = bird + ''mimus'' = mimic), is a genus of oviraptorosaurian theropod dinosaur, named for its bird-like characteristics, that lived in the late Cretaceous in what is now Mongolia, around 85 t ...
'' and ''
Nomingia ''Nomingia'' is a genus of Oviraptorosauria, oviraptorosaur theropod dinosaur hailing from the Late Cretaceous Bugin Tsav Beds of Mongolia. Discovery and naming The remains, consisting of most of the vertebral column, pelvic girdle and left ...
'', other theropods include troodontids such as ''
Tochisaurus ''Tochisaurus'' (meaning "Ostrich lizard") is a genus of small troodontid theropod dinosaur from the Late Cretaceous Period of Mongolia. The type (and only named) species is ''Tochisaurus nemegtensis''. In 1948, a Soviet-Mongolian expedition fou ...
'' and ''
Zanabazar Öndör Gegeen Zanabazar (born Eshidorji) was the first '' Jebtsundamba Khutuktu'' and the first ''Bogd Gegeen'' or supreme spiritual authority, of the Gelugpa (Yellow Hat) lineage of Tibetan Mahayana Buddhism in Mongolia. The son of a Mongol ...
'', the tyrannosaurids ''
Alioramus ''Alioramus'' (; meaning 'different branch') is a genus of tyrannosaurid theropod dinosaurs from the Late Cretaceous period of Asia. It currently contains two species. The type species, ''A. remotus'' is known from a partial skull and three foo ...
'' and ''
Tarbosaurus ''Tarbosaurus'' ( ; meaning "alarming lizard") is a genus of large tyrannosaurid dinosaur that lived in Asia during the Late Cretaceous epoch, about 70 million years ago (Maastrichtian age). It contains the single type species: ''Tarbosaurus ...
''. Ornithischians include the pachycephalosaurs ''
Homalocephale ''Homalocephale'' (from Greek ὁμαλός, ''homalos'', "even", and κεφαλή, ''kephalē'', "head") is a genus of pachycephalosaurid dinosaur that lived during the Late Cretaceous period of what is now the Nemegt Formation, Mongolia, about ...
'' and ''
Prenocephale ''Prenocephale'' (meaning "sloping head") is a genus of small pachycephalosaurid dinosaur from the Late Cretaceous Nemegt Formation of Mongolia. It was similar in many ways to its close relative, ''Homalocephale''. Discovery The holotype specime ...
'', the hadrosaurid ''
Saurolophus ''Saurolophus'' (; meaning "lizard crest") is a genus of large hadrosaurid dinosaur from the Late Cretaceous period of Asia and North America, that lived in what is now the Horseshoe Canyon and Nemegt formations about 70 million to 66 million ...
'', the ankylosaurs ''
Saichania ''Saichania'' (Mongolian meaning "beautiful one") is a genus of herbivorous ankylosaurid dinosaur from the Late Cretaceous period of Mongolia. The first fossils of ''Saichania'' were found in the early 1970s in Mongolia. In 1977 the type speci ...
'' and ''
Tarchia ''Tarchia'' (meaning "brainy one") is a genus of herbivorous ankylosauridae, ankylosaurid dinosaur from the late Cretaceous of Mongolia. Discovery and naming In 1970, a Polish-Mongolian expedition discovered an ankylosaurian skull near Khulsan. ...
''.


See also

*
Timeline of oviraptorosaur research This timeline of oviraptorosaur research is a chronological listing of events in the history of paleontology focused on the oviraptorosaurs, a group of beaked, bird-like theropod dinosaurs. The early history of oviraptorosaur paleontology is cha ...


References


External links

* * {{Taxonbar, from=Q132558 Oviraptoridae Dinosaur genera Maastrichtian dinosaurs Nemegt Formation Fossil taxa described in 1986 Taxa named by Rinchen Barsbold Dinosaurs of Mongolia