Common Fallow Deer
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The European fallow deer (''Dama dama''), also known as the common fallow deer or simply fallow deer, is a
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
of
deer A deer (: deer) or true deer is a hoofed ruminant ungulate of the family Cervidae (informally the deer family). Cervidae is divided into subfamilies Cervinae (which includes, among others, muntjac, elk (wapiti), red deer, and fallow deer) ...
native to
Eurasia Eurasia ( , ) is a continental area on Earth, comprising all of Europe and Asia. According to some geographers, Physical geography, physiographically, Eurasia is a single supercontinent. The concept of Europe and Asia as distinct continents d ...
. It is one of two living species of
fallow deer Fallow deer is the common name for species of deer in the genus ''Dama'' of subfamily Cervinae. There are two living species, the European fallow deer (''Dama dama''), native to Europe and Anatolia, and the Persian fallow deer (''Dama mesopotamic ...
alongside the
Persian fallow deer The Persian fallow deer (''Dama mesopotamica'') is a deer species once native to all of the Middle East, but currently only living in Iran and Israel. It was reintroduced in Israel. It has been listed as endangered on the IUCN Red List since 2008 ...
(''Dama mesopotamica''). It is historically native to
Turkey Turkey, officially the Republic of Türkiye, is a country mainly located in Anatolia in West Asia, with a relatively small part called East Thrace in Southeast Europe. It borders the Black Sea to the north; Georgia (country), Georgia, Armen ...
and possibly the Italian Peninsula,
Balkan Peninsula The Balkans ( , ), corresponding partially with the Balkan Peninsula, is a geographical area in southeastern Europe with various geographical and historical definitions. The region takes its name from the Balkan Mountains that stretch throug ...
, and the island of
Rhodes Rhodes (; ) is the largest of the Dodecanese islands of Greece and is their historical capital; it is the List of islands in the Mediterranean#By area, ninth largest island in the Mediterranean Sea. Administratively, the island forms a separ ...
near
Anatolia Anatolia (), also known as Asia Minor, is a peninsula in West Asia that makes up the majority of the land area of Turkey. It is the westernmost protrusion of Asia and is geographically bounded by the Mediterranean Sea to the south, the Aegean ...
. During the
Pleistocene The Pleistocene ( ; referred to colloquially as the ''ice age, Ice Age'') is the geological epoch (geology), epoch that lasted from to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fin ...
it inhabited much of Europe, and has been reintroduced to its prehistoric distribution by humans. It has also been introduced to other regions in the world.


Taxonomy

Some taxonomists include the rarer
Persian fallow deer The Persian fallow deer (''Dama mesopotamica'') is a deer species once native to all of the Middle East, but currently only living in Iran and Israel. It was reintroduced in Israel. It has been listed as endangered on the IUCN Red List since 2008 ...
as a
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
(''D. d. mesopotamica''), with both species being grouped together as the fallow deer, while others treat it as a different species (''D. mesopotamica''). The
white-tailed deer The white-tailed deer (''Odocoileus virginianus''), also known Common name, commonly as the whitetail and the Virginia deer, is a medium-sized species of deer native to North America, North, Central America, Central and South America. It is the ...
(''Odocoileus virginianus'') was once classified as ''Dama virginiana'' and the
mule deer The mule deer (''Odocoileus hemionus'') is a deer indigenous to western North America; it is named for its ears, which are large like those of the mule. Two subspecies of mule deer are grouped into the black-tailed deer. Unlike the related whit ...
or
black-tailed deer Black-tailed deer or blacktail deer occupy coastal regions of western North America. There are two subspecies, the Columbian black-tailed deer (''Odocoileus hemionus columbianus'') which ranges from the Pacific Northwest of the United States and ...
(''Odocoileus hemionus'') as ''Dama hemionus''; they were given a separate genus in the 19th century.


Description

The male fallow deer is known as a buck, the female is a doe, and the young a fawn. Adult bucks are long, in shoulder height, and typically in weight; does are long, in shoulder height, and in weight. The largest bucks may measure long and weigh .Burnie D and Wilson DE (Eds.), ''Animal: The Definitive Visual Guide to the World's Wildlife''. DK Adult (2005), Fawns are born in spring around and weigh around . Their lifespan is around 12–16 years. Much variation occurs in the coat colour of the species, with four main variants: common, menil,
melanistic Melanism is the congenital excess of melanin in an organism resulting in dark pigment. Pseudomelanism, also called abundism, is another variant of pigmentation, identifiable by dark spots or enlarged stripes, which cover a large part of the bod ...
, and
leucistic Leucism () is a wide variety of conditions that result in partial loss of pigmentation in an animal—causing white, pale, or patchy coloration of the skin, hair, feathers, scales, or cuticles, but not the eyes. It is occasionally spelled ''leu ...
– a genuine colour variety, not albinistic. White is the lightest coloured, almost white; common and menil are darker, and melanistic is very dark, sometimes even black (and is easily confused with the
sika deer The sika deer (''Cervus nippon''), also known as the northern spotted deer or the Japanese deer, is a species of deer native to much of East Asia and introduced to other parts of the world. Previously found from northern Vietnam in the south t ...
). * Common: Chestnut coat with white mottles, it is most pronounced in summer with a much darker, unspotted coat in the winter. The light-coloured area around the tail is edged with black. The tail is light with a black stripe. * Menil: Spots are more distinct than common in summer and no black is seen around the rump patch or on the tail. In winter, spots are still clear on a darker brown coat. *
Melanistic Melanism is the congenital excess of melanin in an organism resulting in dark pigment. Pseudomelanism, also called abundism, is another variant of pigmentation, identifiable by dark spots or enlarged stripes, which cover a large part of the bod ...
(black): All year the coat is black, shading to greyish-brown. No light-coloured tail patch or spots are seen. *
Leucistic Leucism () is a wide variety of conditions that result in partial loss of pigmentation in an animal—causing white, pale, or patchy coloration of the skin, hair, feathers, scales, or cuticles, but not the eyes. It is occasionally spelled ''leu ...
(white, but not albino): Fawns are cream-coloured; adults become pure white, especially in winter. Dark eyes and nose are seen. The coat has no spots. Most herds consist of the common coat variation, yet animals of the menil coat variation are not rare. The melanistic coat variation is generally rarer, and the white coat variation is very much rarer still, although wild New Zealand herds often have a high melanistic percentage. Only bucks have
antlers Antlers are extensions of an animal's skull found in members of the Cervidae (deer) Family (biology), family. Antlers are a single structure composed of bone, cartilage, fibrous tissue, skin, nerves, and blood vessels. They are generally fo ...
, which are broad and shovel-shaped (palmate) from three years. In the first two years the antler is a single spike. Their preferred habitat is mixed woodland and open grassland. During the rut, bucks spread out and females move between them; at that time of year fallow deer are relatively ungrouped compared with the rest of the year, when they try to stay together in groups of up to 150. Agile and fast in case of danger, fallow deer can run at a maximum speed of over short distances. Being naturally less muscular than other cervids such as the roe deer, they are not as fast. Fallow deer can also jump up to high and up to in length. The diet of the European fallow deer has been described as highly flexible, and able to adapt to local conditions. A 1977 study of European fallow deer in the
New Forest The New Forest is one of the largest remaining tracts of unenclosed pasture land, heathland and forest in Southern England, covering southwest Hampshire and southeast Wiltshire. It was proclaimed a royal forest by William the Conqueror, featu ...
of Britain found that European fallow deer were selective mixed feeders, feeding primarily on grass (and to a less extend on
herbs Herbs are a widely distributed and widespread group of plants, excluding vegetables, with savory or aromatic properties that are used for flavoring and garnish (food), garnishing food, for medicinal purposes, or for fragrances. Culinary use typi ...
and
browse Browsing is a kind of orienting strategy. It is supposed to identify something of relevance for the browsing organism. In context of humans, it is a metaphor taken from the animal kingdom. It is used, for example, about people browsing open sh ...
) during the spring and summer (March–September), while primarily feeding on acorns and other
mast Mast, MAST or MASt may refer to: Engineering * Mast (sailing), a vertical spar on a sailing ship * Flagmast, a pole for flying a flag * Guyed mast, a structure supported by guy-wires * Mooring mast, a structure for docking an airship * Radio mas ...
during autumn (from September) until late December, with winter foods including grass as well as shrubs like
brambles ''Rubus'' is a large and diverse genus of flowering plants in the rose family, Rosaceae, subfamily Rosoideae, most commonly known as brambles. Fruits of various species are known as raspberries, blackberries, dewberries, and bristleberries. ...
,
bilberry Bilberries () are Eurasian low-growing shrubs in the genus ''Vaccinium'' in the flowering plant family Ericaceae that bear edible, dark blue berries. They resemble but are distinct from North American blueberries. The species most often referre ...
, heather,
holly ''Ilex'' () or holly is a genus of over 570 species of flowering plants in the family Aquifoliaceae, and the only living genus in that family. ''Ilex'' has the most species of any woody dioecious angiosperm genus. The species are evergreen o ...
, as well as
ivy ''Hedera'', commonly called ivy (plural ivies), is a genus of 12–15 species of evergreen climbing or ground-creeping woody plants in the family Araliaceae, native to Western Europe, Central Europe, Southern Europe, Macaronesia, northwestern ...
and
conifer Conifers () are a group of conifer cone, cone-bearing Spermatophyte, seed plants, a subset of gymnosperms. Scientifically, they make up the phylum, division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a sin ...
ous material.


Distribution

During the
Last Interglacial The Last Interglacial, also known as the Eemian, was the interglacial period which began about 130,000 years ago at the end of the Penultimate Glacial Period and ended about 115,000 years ago at the beginning of the Last Glacial Period. It cor ...
(also known as the Eemian) around 130-115,000 years ago and prior, European fallow deer were widely distributed over Europe, occurring as far north as the
British Isles The British Isles are an archipelago in the Atlantic Ocean, North Atlantic Ocean off the north-western coast of continental Europe, consisting of the islands of Great Britain, Ireland, the Isle of Man, the Inner Hebrides, Inner and Outer Hebr ...
. During the Last Glacial Period (115,000-11,700 years ago) the range of the species collapsed due to unfavourable climate conditions, surviving in refugia in Anatolia and probably the Balkans and possibly elsewhere, though the fossil record of their distribution during this time is sparse.Fernández-García, J. L. (2012
The endangered ''Dama dama mesopotamica'': genetic variability, allelic loss and hybridization signals
''Contributions to Zoology'', 81.4, 223-233.


Turkey

Turkey is the only country known to have definitively natural populations of European fallow deer since the
Last Glacial Maximum The Last Glacial Maximum (LGM), also referred to as the Last Glacial Coldest Period, was the most recent time during the Last Glacial Period where ice sheets were at their greatest extent between 26,000 and 20,000 years ago. Ice sheets covered m ...
, but populations there (alongside those of the
Persian fallow deer The Persian fallow deer (''Dama mesopotamica'') is a deer species once native to all of the Middle East, but currently only living in Iran and Israel. It was reintroduced in Israel. It has been listed as endangered on the IUCN Red List since 2008 ...
, which also formerly occurred in Turkey) have since become endangered and almost fully extirpated. European fallow deer in
Anatolia Anatolia (), also known as Asia Minor, is a peninsula in West Asia that makes up the majority of the land area of Turkey. It is the westernmost protrusion of Asia and is geographically bounded by the Mediterranean Sea to the south, the Aegean ...
underwent a major population decline due to the spread of agriculture (leading to the deforestation of lowland forests) and hunting, and populations in the Marmara and Aegean regions went extinct by the turn of the 20th century. Other wild populations of Turkish fallow deer survived for longer on islands at Ayvalık Islands Nature Park, Gökova, and Adaköy near
Marmaris Marmaris () is a municipality and Districts of Turkey, district of Muğla Province, Turkey. Its area is 906 km2, and its population is 97,818 (2022). It is a port city and tourist resort on the Aegean Sea, Mediterranean coast, along the shorel ...
, but also appear to have died out in recent years. Currently, the only extant wild population of the species known to be undoubtedly natural lives in Düzlerçami Game Reserve in the
Mount Güllük-Termessos National Park Mount Güllük-Termessos National Park (), established in 1970, is a national park located in Antalya Province, southwestern Turkey. Overview The national park is located in Korkuteli district of Antalya Province, northwest of Antalya, in the ...
in southern Turkey, although the area has been largely fenced since, making the population only semi-wild. This population is very few in number and is genetically distinct from other European fallow deer. A 2024 study suggests that the Turkish populations of fallow deer are ancestral to most fallow deer found throughout Europe as well as introduced populations worldwide. The translocations of fallow deer out of Turkey were facilitated by
Roman Roman or Romans most often refers to: *Rome, the capital city of Italy *Ancient Rome, Roman civilization from 8th century BC to 5th century AD *Roman people, the people of Roman civilization *Epistle to the Romans, shortened to Romans, a letter w ...
-era trade networks. The only other refugium found was one in the Balkans, whose surviving descendants are significantly fewer in number.


Native but originally extinct


Southern Balkans

On mainland
Greece Greece, officially the Hellenic Republic, is a country in Southeast Europe. Located on the southern tip of the Balkan peninsula, it shares land borders with Albania to the northwest, North Macedonia and Bulgaria to the north, and Turkey to th ...
and some Greek islands, such as
Corfu Corfu ( , ) or Kerkyra (, ) is a Greece, Greek island in the Ionian Sea, of the Ionian Islands; including its Greek islands, small satellite islands, it forms the margin of Greece's northwestern frontier. The island is part of the Corfu (regio ...
,
Kythira Kythira ( ; ), also transliterated as Cythera, Kythera and Kithira, is an island in Greece lying opposite the south-eastern tip of the Peloponnese peninsula. It is traditionally listed as one of the seven main Ionian Islands, although it is dist ...
, and
Thasos Thasos or Thassos (, ''Thásos'') is a Greek island in the North Aegean Sea. It is the northernmost major Greek island, and 12th largest by area. The island has an area of 380 km2 and a population of about 13,000. It forms a separate regiona ...
, that were connected to mainland due to lower sea level or proximity to land, fallow deer were present during the last ice age.Yiannouli E. & Trantalidou K. 1999: The fallow deer (''Dama dama'' Linnaeus, 1758): Archaeological presence and representation in Greece. The Holocene History of the European Vertebrate Fauna. Modern Aspects of Research. Workshop, 6th to 9nth April 1998, Berlin: 247-281. A belief arose that the species was almost extinct in Greece, returning during the Neolithic. Contrary to that, remains indicate that reduced numbers survived in several parts of the country like in
Thessaly Thessaly ( ; ; ancient Aeolic Greek#Thessalian, Thessalian: , ) is a traditional geographic regions of Greece, geographic and modern administrative regions of Greece, administrative region of Greece, comprising most of the ancient Thessaly, a ...
,
Peloponnese The Peloponnese ( ), Peloponnesus ( ; , ) or Morea (; ) is a peninsula and geographic region in Southern Greece, and the southernmost region of the Balkans. It is connected to the central part of the country by the Isthmus of Corinth land bridg ...
and Central Greece, increasing and becoming common during mid Neolithic, but mostly east of Pindus mountain range and especially in
Macedonia Macedonia (, , , ), most commonly refers to: * North Macedonia, a country in southeastern Europe, known until 2019 as the Republic of Macedonia * Macedonia (ancient kingdom), a kingdom in Greek antiquity * Macedonia (Greece), a former administr ...
and
Thrace Thrace (, ; ; ; ) is a geographical and historical region in Southeast Europe roughly corresponding to the province of Thrace in the Roman Empire. Bounded by the Balkan Mountains to the north, the Aegean Sea to the south, and the Black Se ...
.Trantalidou K. 2002: The Rhodian fallow deer. Game and trophy since prehistoric times, M. Μasseti (ed.), Island of deer. Natural history of the fallow deer of Rhodes and the vertebrates of the Dodecanese (Greece), Rhodes, Prefecture of Rhodes, 159-164. During the Neolithic period and the Bronze Age, the species survived on the islands of Corfu and Thasos, appeared on Euboea, and began to be introduced by man to other islands, including Crete, some of the Cyclades, Rhodes, Chios, Lesbos, Samos and Sporades. Early-historic-period remains have been found in eastern Greece and on the islands of Thasos, Chios, Rhodes and Crete. A few surviving individuals were observed on
Samos Samos (, also ; , ) is a Greek island in the eastern Aegean Sea, south of Chios, north of Patmos and the Dodecanese archipelago, and off the coast of western Turkey, from which it is separated by the Mycale Strait. It is also a separate reg ...
in 1700, while the species became extinct on
Lesbos Lesbos or Lesvos ( ) is a Greek island located in the northeastern Aegean Sea. It has an area of , with approximately of coastline, making it the third largest island in Greece and the List of islands in the Mediterranean#By area, eighth largest ...
late in the Ottoman period. On the Greek mainland, wild fallow deer survived until the 16th century in northeastern
Chalkidiki Chalkidiki (; , alternatively Halkidiki), also known as Chalcidice, is a peninsula and regional unit of Greece, part of the region of Central Macedonia, in the geographic region of Macedonia in Northern Greece. The autonomous Mount Athos reg ...
, until the 19th century in the forests of
Mount Olympus Mount Olympus (, , ) is an extensive massif near the Thermaic Gulf of the Aegean Sea, located on the border between Thessaly and Macedonia (Greece), Macedonia, between the regional units of Larissa (regional unit), Larissa and Pieria (regional ...
,
Vermio Mountains The Vermio Mountains (), known in antiquity as the Bermion (), is a mountain range in northern Greece. It lies between the Imathia Regional Unit of the Central Macedonia Region and the Kozani Regional Unit of the Western Macedonia Region. The r ...
,
Arakynthos Arakynthos (Greek: Αράκυνθος) is a former municipality in Aetolia-Acarnania, West Greece, Greece. Since the 2011 local government reform it is part of the municipality Agrinio Agrinio (Greek language, Greek: Αγρίνιο, ; Latin: '' ...
,
Evrytania Evrytania ( ; ) is one of the regional units of Greece. It is part of the modern regions of Greece, region of Central Greece (administrative region), Central Greece. Its capital is Karpenisi (approx. 8,000 inhabitants). Geography Evrytania is ...
and
Boeotia Boeotia ( ), sometimes Latinisation of names, Latinized as Boiotia or Beotia (; modern Greek, modern: ; ancient Greek, ancient: ), is one of the regional units of Greece. It is part of the modern regions of Greece, region of Central Greece (adm ...
and until the 1910s in
Thesprotia Thesprotia (; , ) is one of the regional units of Greece. It is part of the Epirus region. Its capital and largest town is Igoumenitsa. Thesprotia is named after the Thesprotians, an ancient Greek tribe that inhabited the region in antiquity. His ...
. The last individuals were hunted in
Acarnania Acarnania () is a region of west-central Greece that lies along the Ionian Sea, west of Aetolia, with the Achelous River for a boundary, and north of the gulf of Calydon, which is the entrance to the Gulf of Corinth. Today it forms the western part ...
during the 1930s. In
Bulgaria Bulgaria, officially the Republic of Bulgaria, is a country in Southeast Europe. It is situated on the eastern portion of the Balkans directly south of the Danube river and west of the Black Sea. Bulgaria is bordered by Greece and Turkey t ...
, the autochthonous population of fallow deer is believed to have declined and disappeared after the 9th or 10th centuries, and the species was reintroduced there much more recently. The species remained in European Turkey into the 19th century. A male fallow deer was captured in Thrace in 1977 and translocated to Düzlerçamı, suggesting that a small population existed there at that time. In
Albania Albania ( ; or ), officially the Republic of Albania (), is a country in Southeast Europe. It is located in the Balkans, on the Adriatic Sea, Adriatic and Ionian Seas within the Mediterranean Sea, and shares land borders with Montenegro to ...
(possibly in
Butrint Butrint (, , ) was an ancient Greek polis and later Roman city and the seat of an early Christian bishopric in Epirus. Originally a settlement of the Greek tribe of the Chaonians, it later became part of the state of Epirus and later a Roman ...
), the fallow deer seemed to be plentiful during the first half of the 19th century. A 2024 genetic study suggests that the Balkans served as one of two refugia for fallow deer during the glacial periods, alongside Anatolia. Members of this population were also translocated around Europe during the
Iron Age The Iron Age () is the final epoch of the three historical Metal Ages, after the Chalcolithic and Bronze Age. It has also been considered as the final age of the three-age division starting with prehistory (before recorded history) and progre ...
and Roman Empire, but have largely been replaced by Turkish-origin fallow deer (including in their native Balkans) aside from parts of southern Europe. The Balkan fallow deer are thought to represent the ancestors of modern Iberian, Italian, and Rhodes fallow deer, with the Rhodes population dating back to Neolithic translocations.


Possible native populations

Aside from Turkey, other areas of Europe that could have potentially served as refuges for the species during the last ice age include parts of the eastern
Mediterranean The Mediterranean Sea ( ) is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean basin and almost completely enclosed by land: on the east by the Levant in West Asia, on the north by Anatolia in West Asia and Southern ...
, including most of the Italian Peninsula, parts of the
Balkans The Balkans ( , ), corresponding partially with the Balkan Peninsula, is a geographical area in southeastern Europe with various geographical and historical definitions. The region takes its name from the Balkan Mountains that stretch throug ...
, and the
Greek Greek may refer to: Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group *Greek language, a branch of the Indo-European language family **Proto-Greek language, the assumed last common ancestor of all kno ...
island of
Rhodes Rhodes (; ) is the largest of the Dodecanese islands of Greece and is their historical capital; it is the List of islands in the Mediterranean#By area, ninth largest island in the Mediterranean Sea. Administratively, the island forms a separ ...
, all of which still host populations of this species. However,
palaeontological Paleontology, also spelled as palaeontology or palæontology, is the scientific study of the life of the past, mainly but not exclusively through the study of fossils. Paleontologists use fossils as a means to classify organisms, measure geolo ...
and archaeozoological evidence of the species' diffusion into these areas during the ice age is very fragmentary, thus whether the present populations in these areas are truly native descendants of
relict A relict is a surviving remnant of a natural phenomenon. Biology A relict (or relic) is an organism that at an earlier time was abundant in a large area but now occurs at only one or a few small areas. Geology and geomorphology In geology, a r ...
populations or were introduced by humans is unknown. Presently, the
IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is an inventory of the global conservation status and extinction risk of biological ...
's range map lists European fallow deer as being native to Italy, Turkey, Rhodes, and most of the Balkans, as having a population of uncertain origin in central
Bosnia and Herzegovina Bosnia and Herzegovina, sometimes known as Bosnia-Herzegovina and informally as Bosnia, is a country in Southeast Europe. Situated on the Balkans, Balkan Peninsula, it borders Serbia to the east, Montenegro to the southeast, and Croatia to th ...
, and being introduced to the rest of Europe. In the text, though, all the eastern Mediterranean populations aside from Turkey are listed as having an uncertain origin. A 2024 study suggests that Italian and Iberian populations descend from a now-extinct Balkan population that was translocated early on.


Rhodes, Greece

The Rhodian population of European fallow deer is smaller on average than those of central and northern Europe, though they are similarly coloured. European fallow deer are said to have been introduced to Rhodes in
Neolithic The Neolithic or New Stone Age (from Ancient Greek, Greek 'new' and 'stone') is an archaeological period, the final division of the Stone Age in Mesopotamia, Asia, Europe and Africa (c. 10,000 BCE to c. 2,000 BCE). It saw the Neolithic Revo ...
times; although fossils of the species on Rhodes do indeed go back to Neolithic times, no major evidence has been found of
domestication Domestication is a multi-generational Mutualism (biology), mutualistic relationship in which an animal species, such as humans or leafcutter ants, takes over control and care of another species, such as sheep or fungi, to obtain from them a st ...
, so they could be considered native. In 2005, the Rhodian fallow deer was found to be genetically distinct from all other populations and to be of urgent conservation concern. At the entrance to the harbour of
Rhodes Rhodes (; ) is the largest of the Dodecanese islands of Greece and is their historical capital; it is the List of islands in the Mediterranean#By area, ninth largest island in the Mediterranean Sea. Administratively, the island forms a separ ...
city, statues of a fallow deer buck and doe now grace the location where the
Colossus of Rhodes The Colossus of Rhodes (; ) was a statue of the Greek sun god Helios, erected in the city of Rhodes, on the Greek island of the same name, by Chares of Lindos in 280 BC. One of the Seven Wonders of the Ancient World, it was constructed to ...
once stood. A 2024 study suggests them to be a basal lineage of the Balkan fallow deer, originating from a very early translocation.


Introduced

Outside of Europe, this species has been introduced to
Algeria Algeria, officially the People's Democratic Republic of Algeria, is a country in the Maghreb region of North Africa. It is bordered to Algeria–Tunisia border, the northeast by Tunisia; to Algeria–Libya border, the east by Libya; to Alger ...
,
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,
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,
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,
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,
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,
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, the
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,
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, the
Falkland Islands The Falkland Islands (; ), commonly referred to as The Falklands, is an archipelago in the South Atlantic Ocean on the Patagonian Shelf. The principal islands are about east of South America's southern Patagonian coast and from Cape Dub ...
, Fernando Pó,
Israel Israel, officially the State of Israel, is a country in West Asia. It Borders of Israel, shares borders with Lebanon to the north, Syria to the north-east, Jordan to the east, Egypt to the south-west, and the Mediterranean Sea to the west. Isr ...
,
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,
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,
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,
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,
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,
Peru Peru, officially the Republic of Peru, is a country in western South America. It is bordered in the north by Ecuador and Colombia, in the east by Brazil, in the southeast by Bolivia, in the south by Chile, and in the south and west by the Pac ...
,
Réunion Réunion (; ; ; known as before 1848) is an island in the Indian Ocean that is an overseas departments and regions of France, overseas department and region of France. Part of the Mascarene Islands, it is located approximately east of the isl ...
,
São Tomé São Tomé is the capital and largest city of the Central African island country of São Tomé and Príncipe. Its name is Portuguese for " Saint Thomas". Founded in the 15th century, it is one of Africa's oldest colonial cities. History Álv ...
, the
Seychelles Seychelles (, ; ), officially the Republic of Seychelles (; Seychellois Creole: ), is an island country and archipelagic state consisting of 155 islands (as per the Constitution) in the Indian Ocean. Its capital and largest city, Victoria, ...
,
South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. Its Provinces of South Africa, nine provinces are bounded to the south by of coastline that stretches along the Atlantic O ...
,
Tunisia Tunisia, officially the Republic of Tunisia, is a country in the Maghreb region of North Africa. It is bordered by Algeria to the west and southwest, Libya to the southeast, and the Mediterranean Sea to the north and east. Tunisia also shares m ...
, and the
United States The United States of America (USA), also known as the United States (U.S.) or America, is a country primarily located in North America. It is a federal republic of 50 U.S. state, states and a federal capital district, Washington, D.C. The 48 ...
.


Australia

European fallow deer were introduced to
Tasmania Tasmania (; palawa kani: ''Lutruwita'') is an island States and territories of Australia, state of Australia. It is located to the south of the Mainland Australia, Australian mainland, and is separated from it by the Bass Strait. The sta ...
in 1830 and to mainland Australia in the 1880s. The deer can now be found in all Australian jurisdictions, except Western Australia and the Northern Territory. The European fallow deer is the most widespread and numerous of introduced deer species in Australia. Proper control of deer populations in
New South Wales New South Wales (commonly abbreviated as NSW) is a States and territories of Australia, state on the Eastern states of Australia, east coast of :Australia. It borders Queensland to the north, Victoria (state), Victoria to the south, and South ...
(NSW) was precluded for some years by the classification of these deer as "game animals", as well as being a feral pest species. This led to an explosion in numbers, a vast increase in range in that state, impacts on agricultural production, increased environmental damage, and a dramatic increase in vehicle accidents involving deer. This policy has since been reversed on privately held land only, and on such land the deer is once again only classified as a feral pest species; they remain game animals on public land. The NSW government now asks the public to assist by ''not'' transporting or releasing feral deer onto any land, implying that intentional release of deer has been a factor in the vast increase in range in NSW in recent years.


Argentina

The European fallow deer was introduced to
Victoria Island Victoria Island () is a large island in the Arctic Archipelago that straddles the boundary between Nunavut and the Northwest Territories of Canada. It is the eighth-largest island in the world, and at in area, it is Canada's second-largest i ...
in
Neuquén Province Neuquén () is a Provinces of Argentina, province of Argentina, located in the west of the country, at the northern end of Patagonia. It borders Mendoza Province to the north, Rio Negro Province to the southeast, and Chile to the west. It also me ...
by billionaire Aaron Anchorena, who intended to increase hunting opportunities. He freed wildlife of European and Asian origin, making them common inhabitants of the island.


Canada

The European fallow deer is listed as an
invasive species An invasive species is an introduced species that harms its new environment. Invasive species adversely affect habitats and bioregions, causing ecological, environmental, and/or economic damage. The term can also be used for native spec ...
in the province of
British Columbia British Columbia is the westernmost Provinces and territories of Canada, province of Canada. Situated in the Pacific Northwest between the Pacific Ocean and the Rocky Mountains, the province has a diverse geography, with rugged landscapes that ...
. In 2021, the
Canadian federal government The Government of Canada (), formally His Majesty's Government (), is the body responsible for the federal administration of Canada. The term ''Government of Canada'' refers specifically to the executive, which includes ministers of the Crown ...
, local First Nations, and local residents put forward a plan to eradicate the fallow deer population on Sidney Island, a small island located off the southwest coast of British Columbia.


Great Britain and Ireland

The European fallow deer was spread across
Central Europe Central Europe is a geographical region of Europe between Eastern Europe, Eastern, Southern Europe, Southern, Western Europe, Western and Northern Europe, Northern Europe. Central Europe is known for its cultural diversity; however, countries in ...
by the Romans. Recent finds at
Fishbourne Roman Palace Fishbourne Roman Palace or Fishbourne Villa is in the village of Fishbourne, West Sussex, Fishbourne, near Chichester in West Sussex. The palace is the largest known Roman residence north of the Alps, and has an unusually early date of 75 A ...
show that European fallow deer were introduced into southern England in the first century AD. Fallow deer were established in Britain by the fourth century AD. Genetic studies have shown that this population became extinct and the fallow deer was re-introduced from
Anatolia Anatolia (), also known as Asia Minor, is a peninsula in West Asia that makes up the majority of the land area of Turkey. It is the westernmost protrusion of Asia and is geographically bounded by the Mediterranean Sea to the south, the Aegean ...
prior to the Norman conquest, not introduced from Scilly by the Normans as had previously been believed. Deer from England are a likely source of their re-introduction elsewhere in northern Europe. European fallow deer are now widespread on the UK mainland and are present in most of England and Wales south of a line drawn from
the Wash The Wash is a shallow natural rectangular bay and multiple estuary on the east coast of England in the United Kingdom. It is an inlet of the North Sea and is the largest multiple estuary system in the UK, as well as being the largest natural ba ...
to the
Mersey The River Mersey () is a major river in North West England. Its name derives from Old English and means "boundary river", possibly referring to its having been a border between the ancient kingdoms of Mercia and Northumbria. For centuries it ...
. Populations in the
New Forest The New Forest is one of the largest remaining tracts of unenclosed pasture land, heathland and forest in Southern England, covering southwest Hampshire and southeast Wiltshire. It was proclaimed a royal forest by William the Conqueror, featu ...
and the
Forest of Dean The Forest of Dean is a geographical, historical and cultural region in the western part of the Counties of England, county of Gloucestershire, England. It forms a roughly triangle, triangular plateau bounded by the River Wye to the west and no ...
are long-standing, and many of the other populations originated from park escapees. They are not quite so widespread in the northern parts of England, but are present in most lowland areas and also in parts of Scotland, principally in Strathtay and around Loch Lomond. According to the British Deer Society distribution survey 2007, they have increased in
range Range may refer to: Geography * Range (geographic), a chain of hills or mountains; a somewhat linear, complex mountainous or hilly area (cordillera, sierra) ** Mountain range, a group of mountains bordered by lowlands * Range, a term used to i ...
since the previous survey in 2000, although the increase in range is not as spectacular as for some of the other deer species. A significant number of the European fallow deer in the
Forest of Dean The Forest of Dean is a geographical, historical and cultural region in the western part of the Counties of England, county of Gloucestershire, England. It forms a roughly triangle, triangular plateau bounded by the River Wye to the west and no ...
and in
Epping Forest Epping Forest is a area of ancient woodland, and other established habitats, which straddles the border between Greater London and Essex. The main body of the forest stretches from Epping in the north, to Chingford on the edge of the Lond ...
are of the black variety. One particularly interesting population, known as "long-haired fallow deer", inhabit
Mortimer Forest Mortimer Forest is a forest on the Shropshire/ Herefordshire border in England, near the town of Ludlow. It covers hilly terrain, including the marilyn of High Vinnalls, rising to . History Mortimer Forest was an ancient hunting forest, si ...
on the England/Wales border; a significant part of the population has long body hair with distinct ear tufts. A historical herd is at
Phoenix Park The Phoenix Park () is a large urban park in Dublin, Ireland, lying west of the city centre, north of the River Liffey. Its perimeter wall encloses of recreational space. It includes large areas of grassland and tree-lined avenues, and since ...
in Ireland, where a herd of 400–450 European fallow deer descends from the original herd introduced in the 1660s. In a 2023 study, this herd was shown to comprise the first wild deer outside of North America to have contracted SARS-CoV-2, raising concerns about a potential
natural reservoir In Infection, infectious disease ecology and epidemiology, a natural reservoir, also known as a disease reservoir or a reservoir of infection, is the population of organisms or the specific environment in which an infectious pathogen naturally li ...
arising within European deer herds.


New Zealand

From 1860, European fallow deer were introduced into New Zealand. Significant herds exist in a number of low-altitude forests.


South Africa

European fallow deer are popular in the rural areas of
KwaZulu-Natal KwaZulu-Natal (, also referred to as KZN) is a Provinces of South Africa, province of South Africa that was created in 1994 when the government merged the Zulu people, Zulu bantustan of KwaZulu ("Place of the Zulu" in Zulu language, Zulu) and ...
for hunting purposes, in parts of the
Gauteng Gauteng ( , ; Sotho-Tswana languages, Sotho-Tswana for 'place of gold'; or ) is one of the nine provinces of South Africa. Situated on the Highveld, Gauteng is the smallest province by land area in South Africa. Although Gauteng accounts f ...
Province to beautify ranches, and in the
Eastern Cape The Eastern Cape ( ; ) is one of the nine provinces of South Africa. Its capital is Bhisho, and its largest city is Gqeberha (Port Elizabeth). Due to its climate and nineteenth-century towns, it is a common location for tourists. It is also kno ...
where they were introduced on game farms for the hunting industry because of their exotic qualities. European fallow deer adapted extremely well to the South African environment with access to savanna grasslands, particularly in the cooler climate ranges such as the
highveld The Highveld (Afrikaans: ''Hoëveld,'' , ) is the portion of the South African inland plateau which has an altitude above roughly , but below , thus excluding the Lesotho mountain regions to the south-east of the Highveld. It is home to some of t ...
. They also occur in the western cape.


Sweden

One noted historical herd of European fallow deer is located in the Ottenby nature reserve in
Öland Öland (, ; ; sometimes written ''Oland'' internationally) is the second-largest Swedish island and the smallest of the traditional provinces of Sweden. Öland has an area of and is located in the Baltic Sea just off the coast of Småland. ...
, where
Charles X Gustav of Sweden Charles X Gustav, also Carl X Gustav (; 8 November 1622 – 13 February 1660), was King of Sweden from 1654 until his death. He was the son of John Casimir, Count Palatine of Zweibrücken-Kleeburg and Catherine of Sweden. After his father's ...
erected a
dry stone Dry stone, sometimes called drystack or, in Scotland, drystane, is a building method by which structures are constructed from stones without any mortar to bind them together. A certain amount of binding is obtained through the use of carefully ...
wall some 4 km long to enclose a royal fallow deer herd in the mid-17th century; the herd still exists as of 2006.


United States

In recent times, European fallow deer have been introduced in parts of the United States. A small feral population exists on one barrier island in
Georgia Georgia most commonly refers to: * Georgia (country), a country in the South Caucasus * Georgia (U.S. state), a state in the southeastern United States Georgia may also refer to: People and fictional characters * Georgia (name), a list of pe ...
. Fallow deer have also been introduced in
Texas Texas ( , ; or ) is the most populous U.S. state, state in the South Central United States, South Central region of the United States. It borders Louisiana to the east, Arkansas to the northeast, Oklahoma to the north, New Mexico to the we ...
, along with many other exotic deer species, where they are often hunted on large game ranches. In
Pennsylvania Pennsylvania, officially the Commonwealth of Pennsylvania, is a U.S. state, state spanning the Mid-Atlantic (United States), Mid-Atlantic, Northeastern United States, Northeastern, Appalachian, and Great Lakes region, Great Lakes regions o ...
, European fallow deer are considered livestock, since no feral animals are breeding in the wild. Occasional reports of wild European fallow deer in Pennsylvania and Indiana are generally attributed to escapes from preserves or farms. A herd of white European fallow deer is located near Argonne National Laboratories in northeastern Illinois. A small herd of 15 mostly white European fallow deer resides at the Belle Isle Nature Zoo on Belle Isle in
Detroit, Michigan Detroit ( , ) is the List of municipalities in Michigan, most populous city in the U.S. state of Michigan. It is situated on the bank of the Detroit River across from Windsor, Ontario. It had a population of 639,111 at the 2020 United State ...
. Until the turn of the 21st century, this herd had the run of the island; the herd was thereafter confined, with
extirpation Local extinction, also extirpation, is the termination of a species (or other taxon) in a chosen geographic area of study, though it still exists elsewhere. Local extinctions are contrasted with extinction, global extinctions. Local extinctions ...
being the initial goal. A small herd, believed to be the oldest in the United States, exists in the Land Between the Lakes National Recreation Area (LBL) in far western Kentucky and Tennessee. The European fallow deer herd in the LBL "was brought to LBL by the Hillman Land Company in 1918. LBL's herd is believed to be the oldest population of fallow deer in the country, and at one time was the largest. Today, the herd numbers fewer than 150 and hunting of fallow deer is not permitted. Although LBL's wildlife management activities focus on native species, the fallow herd is maintained for wildlife viewing and because of its historical significance." European fallow deer are present in the
Point Reyes National Seashore Point Reyes National Seashore is a park preserve located on the Point Reyes, Point Reyes Peninsula in Marin County, California. As a national seashore, it is maintained by the US United States National Park Service, National Park Service as an ...
, California, and
Mendocino County Mendocino County (; ''Mendocino'', Spanish language, Spanish for "of Antonio de Mendoza, Mendoza") is a County (United States), county located on the North Coast (California), North Coast of the U.S. state of California. As of the 2020 United S ...
near Ridgewood Ranch, west of
Redwood Valley, California Redwood Valley (formerly Basil) is a census-designated place (CDP) in Mendocino County, California, United States. It is located north of Ukiah, the county seat, at an elevation of , and comprises the northern portion of the Ukiah Valley. It ...
; some of them are leucistic.


Mating system

European fallow deer are highly dimorphic,
polygynous Polygyny () is a form of polygamy entailing the marriage of a man to several women. The term polygyny is from Neoclassical Greek πολυγυνία (); . Incidence Polygyny is more widespread in Africa than in any other continent. Some scholar ...
breeders;Feldhamer, G. A., Farris-Renner, K. C., & Barker, C. M. (1988). ''Dama dama''. ''Mammalian Species'', ''97''(317), 1–8. .McElligot, A. G., Mattiangeli, V., Mattiello, S., Verga, M., Reynolds, C. A., & Hayden, T. (1998). Fighting tactics of fallows bucks (''Dama dama'', Cervidae): Reducing the Risks of Serious Conflict. ''Ethology'', ''104''(9), 789–803. . the breeding season or rut lasts about 135 days. In the Northern Hemisphere, the breeding season tends to occur in the second half of October, while it occurs in April in the Southern Hemisphere, some matings can still occur before and after.Thirgood, S. J. (1991). Alternative Mating Strategies and Reproductive Success in Fallow Deer. ''Behaviour'', ''116''(1/2), 1–10. . . This mating behaviour within the rut most often occurs in leks, where males congregate in small groups on mating territories in which the females’ only purpose for visiting these territories is for copulation. Variation within European fallow deer
mating system A mating system is a way in which a group is structured in relation to sexual behaviour. The precise meaning depends upon the context. With respect to animals, the term describes which males and females mating, mate under which circumstances. Reco ...
s occurs; other than the traditional behaviour of lekking, different types of mating behaviours can include
harem A harem is a domestic space that is reserved for the women of the house in a Muslim family. A harem may house a man's wife or wives, their pre-pubescent male children, unmarried daughters, female domestic Domestic worker, servants, and other un ...
s, dominance groups, stands, temporary stands, and multiple stands.Langbein, J. & Thirgood, S. J. (1989). Variation in Mating Systems of Fallow Deer (''Dama dama'') in Relation to Ecology. ''Ethology'', ''83''(3), 195–214. . Different populations, environmental variation, size, and even age can determine the type of variation within a European fallow deer mating system, but lekking behaviour is the most commonly found and studied in nature; variation can be explained by three characteristics (1) the optimal strategy under specific environmental or social conditions, (2) the strategy of an individual may be dependent on the strategies of other individual males within the same population, and (3) individual males may be less capable at gaining access to females, since they can be outcompeted by other males that are more capable. Female European fallow deer are Estrous cycle, polyestrous; they are receptive to males during multiple periods of estrus throughout the mating season while not gestating. Male rut behaviour includes licking and sniffing around the anus and vulva to determine whether a female is fertile. Males produce high-pitched whines repeatedly to initiate mating; following this display, a female may allow the male to mount; copulation (zoology), copulation can last as long as 5 minutes.


Ecology and mating system characteristics

Many deer species—including European fallow deer—have a social organization that can be tremendously plastic depending on their environment, meaning that group size and habitat type are closely linked to herd size.Putman, R. J. (1986).
Grazing in temperate ecosystems: Large herbivores and the ecology of the New Forest
'. Croom Helm: Beckenham.
Most of the detailed research on the ecological characteristics and behaviour of European fallow deer occurs in large blocks of woodland, which means some bias may be present. European fallow deer can be found in a variety of habitats, which can range from cool and wet to hot and dry. European fallow deer seem to have a preference for older forests with dispersed areas of grass, trees, and a variety of other vegetation. The largest herd occurs right before the rutting season, while the smallest groups are females with fawns. Throughout a large portion of the year, the sexes remain separated and only congregate during the mating months, but other patterns may be described, such as bachelor groups and even mixed groups. Male European fallow deer produce low-frequency vocalizations called groans; the sound of these groans results from the consistent and complex shape of the vocal tract involving the oral and nasal cavities. Ruts are characterized by males gaining the best territory possible to increase their odds for mating, and are often characterized by the presence of females on stands. During this time, males stop feeding to defend their ruts from subordinate males. Males defending this territory often lose an average of 17% of their body weight, and the liver exhibits steatosis, which is reversible. Throughout breeding seasons, the males may obtain the same rut; in some cases, ruts can be held by more than one individual; some possibilities for this include high population density and less rut space, or more suitable habitats, which can be shared.


Parental care

After a female is impregnated, gestation lasts up to 245 days. Usually one fawn is born; twins are rare. The females can conceive when they are 16 months old, whereas the males can successfully breed at 16 months, but most do not breed until they are 48 months old. The females can become very cagey just before they give birth to their fawn and find secluded areas such as a bush or cave; sometimes females give birth near the herd. As soon as the female gives birth, the she then licks the fawn to clean it; this helps initiate the maternal bond between the two, and only females provide parental investment; males do not participate in rearing the fawn. After the birth of the fawn occurs, the females do not return to the herd for at least 10 days and for most of the days the mother is separated from the fawn, returning only to feed the fawn. The nursing period lasts about 4 months and happens every 4 hours each day. Ruminant, Rumination is a critical part of development in the fawn's life, and this develops about 2 to 3 weeks into the fawn's life. Females initiate the weaning periods for the fawn, which lasts about 20 days; 3 to 4 weeks; later, the fawn will start to follow its mother, and they will finally rejoin the herd together. The mother frequently licks the fawn's anal area to stimulate suckling, urination, and defecation, which is a critical part of the development of the fawn. Weaning is completed at around 7 months, and at around 12 months, the fawn is independent; after the 135 days of reproduction, the rut comes to an end, which can be characterized by the changes in group size and behaviour.


Contests and weaponry

Since European fallow deer are polygynous species that congregate once every year, males must fight to obtain access to estrous females.Bergeron, P., Festa-Bianchet, M., von Hardenberg, A., & Bassano, B. (2008). Heterogeneity in Male Horn Growth and Longevity in a Highly Sexually Dimorphic Ungulate. ''Oikos'', ''117''(1), 77–82. . . The relationship between antler size and body condition can be treated as indicators to reflect body condition within a given year.Ciuti, S., & Apollonio, M. (2011). Do Antlers Honestly Advertise the Phenotypic Quality of Fallow Buck (''Dama dama'') in a Lekking Population? ''Ethology'', ''117''(2), 133–144. . These secondary sexual characteristics can have dual functions, which include the attractiveness of males, which females can ultimately choose, and fighting ability of the male.Darwin, C. (1859). ''On the Origin of Species by Means of Natural Selection.'' (Murray, London).Darwin, C. (1871) ''The Descent of Man, and Selection in Relation to Sex.''Andersson M. (1994).
Sexual selection
'. Princeton University Press, Princeton, New Jersey
It was found that males with larger antlers had higher mating success, where males with asymmetrical antlers did not. When males develop their antlers, trade-offs are made between reproduction and survival. Genetic variations exist within fallow deer populations with variable antler growth, males that exhibited faster-growing antlers early in life are able to grow longer antlers without any significant cost; this shows that there is phenotypic variation among fallow deer populations. Aggressive behaviour is often observed when individuals are seeking out mates, scarce resources, and even territories.Jennings, D. J., Boys, R. J., & Gammell, M. P. (2017). Weapon damage is associated with contest dynamics but not mating success in fallow deer (''Dama dama''). ''Biology Letters'', ''13''(11), 20170565. . Species that compete using their weapons usually engage with mutual agreement, but if any noticeable asymmetries are seen, such as a broken or lost weapon, this may alter the behaviour of an individual to engage in a fight. Likelihood and severity of antler damage were looked at in fallow deer, to test whether antler damage was associated with contest tactics and duration, and if an association existed with the tendency for individuals to engage in fighting. Individuals with undamaged antlers were more likely to attack, using high-risk tactics that included jumping, clashing, or backward-pushing behaviour, this was exhibited by both contestants; dominant males were more likely to have damaged antlers. Dominance ranks exist within fallow deer populations, which can be linked to aggression level and body size; when competing for a male, however, how ranks are obtained is not studied extensively.Jennings, D. J., Carlin, C. M., Hayden, T. J., & Gammell, M. P. (2010). Investment in fighting in relation to body condition, age and dominance rank in the male fallow deer, ''Dama dama''. ''Animal Behaviour'', ''79''(6), 1293–1300. .


Endurance rivalry

Male fallow deer are highly competitive during the rutting season; successful mating depends mainly on body size and dominance rank.McElligott, A. G., Gammell, M. P., Harty, H. C., Paini, D. R., Murphy, D. T., Walsh, J. T., & Hayden, T. J. (2001). Sexual size dimorphism in fallow deer (''Dama dama''): Do larger, heavier males gain greater mating success? ''Behavioral Ecology and Sociobiology'', ''49''(4), 266–272. . . Many factors can determine the seasonal reproductive success of an individual male fallow deer; these factors include body size, which can affect reproduction and survival. The amount of time spent in a lek can be an important factor in determining male reproductive success; energy can play an important role for the duration of competitive leks.Craig, A. S., Herman, L. M., Gabriele, C. M., & Pack, A. A. (2003). Migratory timing of humpback whales (''Megaptera novaengliae'') in the central North Pacific varies with age, sex and reproductive status. ''Behaviour'', ''140''(8), 981–1001. . . Among ungulates, European fallow deer exhibit one of the most outstanding examples of sexual dimorphism, as males are much larger than females. For sexual selection to lead to the evolution of sexual dimorphism, where males are bigger than females, advantages must be present: (1) Advantages during combat, (2) Endurance rivalry advantage, (3) Female preference for larger males, and (4) Advantages during sperm competition. Sexual selection has chosen bigger males over an evolutionary time scale and conferred advantages during competition of mates by a variety of mechanisms, which are intrasexual competition, access to females, and resource accessibility, which affects attractiveness to females. Body size is important during male-male agonistic interactions and endurance rivalry, while females tend to have a preference for larger males.Alonso, J. C., Magaña, M., Palacín, C., & Carlos, M. A. (2010). Correlates of male mating success in great bustard leks: the effects of age, weight, and display effort. ''Behavioral Ecology and Sociobiology'', ''64''(10), 1589–1600. . . Dominance rank is a good indicator of body size and body mass, but age was not an important factor. In a study done by McElligott et al. (2001), it was found that mating success was related to body size, pre-rut and rut rank. Similarly, in another study, researchers found that age, weight, and display effort were all significant factors in determining mating success; in both studies, mating success was measured by the frequency of copulations, which means that a variety of factors in different fallow deer populations can affect the overall energy allocation which will ultimately affect mating success. Maternal investment early in life can be critical to the development of body size, since it can be quite variable at that stage depending on resources and habitat type. Mature male body size can be a better indicator of overall male quality rather than body mass, since body mass depends on a variety of resources each year and is not a static trait; body mass can be a complex trait to measure. File:Daniel wisniok.jpg, Male fallow deer File:Fallow deer fawn in uruguayan zoo..JPG, Fallow deer fawn in Uruguay


Threats

Since the 20th century, a serious decline in the populations of European fallow deer has been seen in Turkey, the only region where it is definitely thought to be native, and it has disappeared from almost all regions where it was formerly found aside from Düzlerçami Game Reserve in the
Mount Güllük-Termessos National Park Mount Güllük-Termessos National Park (), established in 1970, is a national park located in Antalya Province, southwestern Turkey. Overview The national park is located in Korkuteli district of Antalya Province, northwest of Antalya, in the ...
, where a semiwild, genetically distinct population exists. The Turkish government undertook a breeding program at Düzlerçami starting in 1966, with the population growing from 7 to 500 animals, but it underwent a massive collapse until 2000 due to reasons not fully understood, but thought to be linked to urbanization, recreational activities, and poaching, and numbered less than 30 (with only 10 individuals roaming outside the fenced areas) individuals by 2007 and less than 130 individuals by 2010. This population remains at risk from inbreeding and poaching. Species reintroduction, Reintroduction to other areas of Turkey has not been successful but should still be considered to increase the species' population. The population on Rhodes, which is of uncertain origin, but is known to be very genetically distinct from others, is also of major conservation concern. It numbers around 400-500 animals and is at risk from poaching and wildfires. The population is also at risk of outbreeding depression, as in some parts of Rhodes, mainland European fallow deer are kept in fenced areas; these deer could escape and breed with the Rhodian fallow deer. Rhodian fallow deer also damage summer crops and due to a lack of a compensation system, persecution against the population could happen. A reduction of water resources on the island due to climate change could also affect the animals. Despite this, there are signs of population recovery on Rhodes as of 2008 due to conservation measures. Despite these threats, the European fallow deer is common across the other areas where it could potentially be native, as well as the areas throughout Europe that it was introduced to early on, thus it is considered to be of Least-concern species, least concern by the IUCN Red List.


See also

* Deer of Great Britain


References


Further reading

* Clutton-Brock, J. (1978). ''A Natural History of Domesticated Animals''. London: British Museum. * de Vos, Antoon (1982). ''Deer farming guidelines on practical aspects''. FAO ANIMAL PRODUCTION AND HEALTH PAPER 27. . . Retrieved on 4 January 2008. * Lyneborg, L. (1971). ''Mammals'' [of Europe]. . * ''Level 1 DSC Training Manual''. https://web.archive.org/web/20090212222910/http://www.eskdalewildlife.com/training.html *


External links


Fallow deer in New Zealand



Fallow Deer Fighting
{{Authority control Cervines Fauna of the Falkland Islands Introduced mammals of Australia National symbols of Antigua and Barbuda Mammals described in 1758 Mammals of Africa Mammals of Asia Mammals of Europe Mammals of Madagascar Mammals of Mauritius Mammals of New Zealand Mammals of North Africa Mammals of Peru Mammals of Réunion Mammals of South Africa Mammals of Sub-Saharan Africa Mammals of the Comoros Mammals of the Middle East, Deer, Fallow Mammals of Turkey Taxa named by Carl Linnaeus