Colaspis Melaina

''Colaspis'' is a genus of leaf beetles in the subfamily Eumolpinae. It is one of the largest genera in the subfamily, containing over 200 species, and it is known from both North and South America. A number of species from this genus are considered to be pests, such as the grape colaspis (''Colaspis brunnea''). Some species are known from the fossil record from the Eocene of Colorado in the United States.

Taxonomy

In some publications for the Neotropical realm, ''Colaspis'' is also known as ''Maecolaspis''. This alternative name was created by the Czech entomologist Jan Bechyné in 1950, due to an error relating to the type species of ''Colaspis'': Bechyné incorrectly considered ''Colaspis testacea'' to be the type species of ''Colaspis'', rather than ''Chrysomela flavicornis''. As a result, ''Metaxyonycha'', which shared the same type species, was synonymised with ''Colaspis''. The species of ''Colaspis'' in the Junk-Schenkling catalog were then placed under Bechyné's new name, "''Maecolaspis''", with ''Chrysomela flavicornis'' as the type species. These errors were corrected by W. J. Brown in 1961, who synonymized ''Maecolaspis'' with ''Colaspis''. Despite this, Bechyné continued to use ''Maecolaspis'' as valid, while Doris Holmes Blake published several papers on ''Colaspis'' without mentioning Bechyné's ''Maecolaspis''. This effectively created two systems for the taxonomy of the Neotropical leaf beetles, which has frequently confused those studying the Neotropical Eumolpinae.

Many species originally placed in ''Colaspis'' were split by Bechyné into several smaller genera, such as ''Allocolaspis'', ''Callicolaspis'', ''Percolaspis'', ''Nodocolaspis'' and ''Zenocolaspis''. The genus requires further subdivision.

Species

These extant species belong to the genus ''Colaspis'' (= ''Maecolaspis''):

* ''Colaspis abdominalis'' Lefèvre, 1877
* ''Colaspis aberrans'' ( Bechyné, 1953)
* ''Colaspis achardi'' Bechyné, 1949
** ''Colaspis achardi achardi'' Bechyné, 1949
** ''Colaspis achardi pseudoachardi'' ( Bechyné, 1950)
* ''Colaspis adducta'' Clavareau, 1914
* ''Colaspis adusta'' Lefèvre, 1885
* ''Colaspis aenea'' Fabricius, 1801
* ''Colaspis aeneicollis'' Bowditch, 1921
* '' Colaspis aeneola'' Weise, 1921 ^g
** ''Colaspis aeneola aeneola'' Weise, 1921
** ''Colaspis aeneola barticensis'' ( Bechyné, 1951)
* ''Colaspis aerea'' Lefèvre, 1884 ^g
** ''Colaspis aerea aerea'' Lefèvre, 1884
** ''Colaspis aerea odetta'' ( Bechyné, 1953)
* ''Colaspis aeruginosa'' ( Germar, 1824)
** ''Colaspis aeruginosa aeruginosa'' ( Germar, 1824)
** ''Colaspis aeruginosa tenuesculpta'' ( Bechyné, 1950)
* ''Colaspis affinis'' Weise, 1921
* '' Colaspis alcyonea'' Suffrian, 1866
* ''Colaspis amabilis'' Jacoby, 1900
* ''Colaspis amazonae'' Jacoby, 1900
* ''Colaspis amplicollis'' Blake, 1971
* '' Colaspis anceps'' Lefèvre, 1878
* ''Colaspis ansa'' Riley, 2020
* ''Colaspis apurensis'' ( Bechyné, 1951)
* ''Colaspis araguensis'' ( Bechyné, 1958)
* ''Colaspis arizonensis'' Schaeffer, 1933 ^{i c g b}
* '' Colaspis assimilis'' Klug, 1829
* ''Colaspis balyi'' Jacoby, 1881
* ''Colaspis barberi'' Blake, 1967
* ''Colaspis basipennis'' Bowditch, 1921
* ''Colaspis batesi'' Jacoby, 1900
* ''Colaspis bicolor'' Germar, 1824
* ''Colaspis bidenticollis'' Bowditch, 1921
* ''Colaspis blakeae'' Ostmark, 1975
* '' Colaspis bohumilae'' ( Bechyné, 1950)
* '' Colaspis braxatibiae'' Blake, 1978
* '' Colaspis bridarollii'' Bechyné, 1949
* ''Colaspis brownsvillensis'' Blake, 1976 ^{i c g b} (Brownsville milkvine leaf beetle)
* ''Colaspis brunnea'' (Fabricius, 1798) ^{i c g b} (grape colaspis)
* ''Colaspis brunneipennis'' Bowditch, 1921
* ''Colaspis buckleyi'' ( Bechyné, 1953)
** ''Colaspis buckleyi buckleyi'' ( Bechyné, 1953)
** ''Colaspis buckleyi incomparabilis'' ( Bechyné, 1950)
* ''Colaspis calcarifera'' ( Bechyné, 1954) (note: pages 196–199 are missing in the PDF)
* '' Colaspis caligula'' Agrain, Cabrera, Holgado & Vicchi, 2016
* ''Colaspis callichloris'' Lefèvre, 1878
* ''Colaspis carolinensis'' Blake, 1974 ^{i c g}
* ''Colaspis castanea'' Boheman, 1858
* '' Colaspis championi'' Jacoby, 1881 ^{i c g}
* ''Colaspis chapalensis'' Blake, 1976
* '' Colaspis chapuisi'' Jacoby, 1884
* ''Colaspis chlorana'' Lefèvre, 1891
* ''Colaspis chlorites'' Erichson, 1847
* ''Colaspis chloropsis'' Blake, 1976
* ''Colaspis coelestina'' Erichson, 1847
* ''Colaspis concolor'' Bowditch, 1921 ^g
* ''Colaspis confusa'' Bowditch, 1921
* '' Colaspis corrugata'' Lefèvre, 1885
* ''Colaspis corumbensis'' Blake, 1978
* ''Colaspis costipennis'' Crotch, 1873 ^{i c g b}
* ''Colaspis cribellata'' Lefèvre, 1888
* ''Colaspis cribricollis'' Lefèvre, 1884
** ''Colaspis cribricollis cribricollis'' Lefèvre, 1884
** ''Colaspis cribricollis machacalisa'' ( Bechyné, 1958)
* ''Colaspis crinicornis'' Schaeffer, 1933 ^{i c g b}
* ''Colaspis cruriflava'' Blake, 1977 ^{i c g b}
* '' Colaspis cubensis'' Blake, 1967
* ''Colaspis darlingtoni'' Blake, 1967
* '' Colaspis dejeani'' Lefèvre, 1884 ^g
* '' Colaspis deleta'' Suffrian, 1867
* ''Colaspis delphina'' ( Bechyné, 1951)
* ''Colaspis demersa'' ( Bechyné, 1950)
* '' Colaspis dentifera'' ( Bechyné, 1951)
* '' Colaspis dentipyga'' ( Bechyné, 1958)
* '' Colaspis diduma'' Blake, 1976
* ''Colaspis dilatipes'' Bowditch, 1921
* ''Colaspis dionysea'' Bechyné, 1949
* ''Colaspis discolor'' Weise, 1921
* ''Colaspis dispar'' Bowditch, 1921
* '' Colaspis diversa'' Lefèvre, 1878
* '' Colaspis dugesi'' Lefèvre, 1885
* ''Colaspis ekraspedona'' Blake, 1978
* ''Colaspis exarata'' Lefèvre, 1884
* ''Colaspis farri'' Blake, 1967
* ''Colaspis favosa'' Say, 1824 ^{i c g b}
* ''Colaspis femoralis'' Olivier, 1808
* ''Colaspis fervida'' (Suffrian, 1866)
* ''Colaspis flavantenna'' Blake, 1978
* ''Colaspis flavicornis'' (Fabricius, 1787)
** ''Colaspis flavicornis flavicornis'' (Fabricius, 1787)
** ''Colaspis flavicornis pedator'' ( Bechyné, 1950)
** ''Colaspis flavicornis urbana'' ( Bechyné, 1950)
* ''Colaspis flavipes'' Olivier, 1808
* ''Colaspis flavocostata'' Schaeffer, 1933 ^{i c g b}
* '' Colaspis floridana'' Schaeffer, 1933 ^{i c g}
* ''Colaspis foersteri'' ( Bechyné, 1958)
* ''Colaspis foveicollis'' Jacoby, 1880
* ''Colaspis freyi'' Bechyné, 1950
** ''Colaspis freyi adscripta'' ( Bechyné, 1953)
** ''Colaspis freyi freyi'' Bechyné, 1950
* ''Colaspis fulvilabris'' Jacoby, 1880
* '' Colaspis fulvotestacea'' Lefèvre, 1878
* ''Colaspis fuscipes'' Bowditch, 1921
* ''Colaspis gemellata'' Lefèvre, 1885
* ''Colaspis geminata'' Boheman, 1858
* ''Colaspis gemmingeri'' Harold, 1874
** ''Colaspis gemmingeri chaparensis'' ( Bechyné, 1951)
** ''Colaspis gemmingeri gemmingeri'' Harold, 1874
* ''Colaspis gemmula'' Erichson, 1847
* '' Colaspis geniculata'' Lefèvre, 1891
* ''Colaspis goyazensis'' Bowditch, 1921
* '' Colaspis guatemalensis'' Blake, 1976
* ''Colaspis gwendolina'' ( Bechyné, 1953)
* ''Colaspis hesperia'' Blake, 1974 ^{i c g b}
* ''Colaspis heteroclita'' Jacoby, 1900
* ''Colaspis hirticornis'' ( Bechyné, 1955)
* ''Colaspis hypochlora'' Lefèvre, 1878
** ''Colaspis hypochlora anteposita'' ( Bechyné, 1950)
** ''Colaspis hypochlora hypochlora'' Lefèvre, 1878
* ''Colaspis impressipennis'' Bowditch, 1921
* '' Colaspis inconspicua'' Jacoby, 1890
* '' Colaspis interstitialis'' Lefèvre, 1877
* ''Colaspis intricata'' Lefèvre, 1888
* ''Colaspis jalapae'' ( Bechyné, 1950)
* ''Colaspis jansoni'' Jacoby, 1881
* ''Colaspis janssensi'' ( Bechyné, 1950)
* ''Colaspis jocosa'' Bowditch, 1921
* ''Colaspis joliveti'' ( Bechyné, 1950)
* ''Colaspis juxaoculus'' Blake, 1978
* '' Colaspis kaszabi'' ( Bechyné, 1953)
* ''Colaspis keyensis'' Blake, 1974 ^{i c g}
* ''Colaspis kirra'' Blake, 1976
* '' Colaspis klagii'' Bowditch, 1921
* ''Colaspis labilis'' ( Bechyné, 1953)
* ''Colaspis lampomela'' Blake, 1978
* ''Colaspis laplatensis'' Bechyné, 1949
* ''Colaspis laeta'' Germar, 1821
* ''Colaspis lata'' Schaeffer, 1933 ^{i c g}
* ''Colaspis lauei'' ( Bechyné, 1950)
* ''Colaspis lebasi'' Lefèvre, 1878
** ''Colaspis lebasi falconensis'' ( Bechyné, 1997)
** ''Colaspis lebasi gwendolina'' ( Bechyné, 1953)
** ''Colaspis lebasi lebasi'' Lefèvre, 1878
** ''Colaspis lebasi praecipitata'' ( Bechyné, 1997)
** ''Colaspis lebasi ruderalis'' ( Bechyné, 1997)
* ''Colaspis lebasiformis'' ( Bechyné, 1953)
* '' Colaspis lebasoides'' Bowditch, 1921
* ''Colaspis legionalis'' ( Bechyné, 1953) ^g
** ''Colaspis legionalis bubonica'' ( Bechyné, 1997)
** ''Colaspis legionalis dispersa'' ( Bechyné, 1997)
** ''Colaspis legionalis legionalis'' ( Bechyné, 1953)
* ''Colaspis leopoldina'' ( Bechyné, 1954)
* ''Colaspis leucopus'' Harold, 1875
* ''Colaspis levicostata'' Blake, 1976
* '' Colaspis libatrix'' ( Bechyné, 1953)
* ''Colaspis longipennis'' Blake, 1976
* '' Colaspis lophodes'' Blake, 1974
* ''Colaspis louisianae'' Blake, 1974 ^{i c g b}
* ''Colaspis luciae'' Blake, 1967
* ''Colaspis luridula'' Lefèvre, 1878
* ''Colaspis luteicornis'' (Fabricius, 1792)
* ''Colaspis lutescens'' Lefèvre, 1886
* ''Colaspis macroptera'' Blake, 1976
* '' Colaspis maida'' ( Bechyné, 1954)
* ''Colaspis manausa'' Blake, 1978
* '' Colaspis melaina'' Blake, 1974 ^{i c g}
* ''Colaspis melancholica'' Jacoby, 1881
* ''Colaspis meriamae'' ( Bechyné, 1951)
* '' Colaspis metallica'' Lefèvre, 1891 ^g
* ''Colaspis mexicana'' Jacoby, 1881
* ''Colaspis micans'' Weise, 1921
* '' Colaspis minuta'' Lefèvre, 1891
* ''Colaspis missionea'' Bechyné, 1949
* ''Colaspis mixticolor'' ( Bechyné, 1951)
* ''Colaspis moesta'' Horn, 1895
* ''Colaspis monomorpha'' ( Bechyné, 1950)
* ''Colaspis monrosi'' ( Bechyné, 1950)
* '' Colaspis montana'' Jacoby, 1891
* ''Colaspis musae'' Bechyné, 1950 ^g
* ''Colaspis nigricornis'' Fabricius, 1801
* '' Colaspis nigripennis'' Jacoby, 1880
* ''Colaspis nigrocyanea'' Crotch, 1873 ^{i c g b}
* '' Colaspis notaticornis'' Lefèvre, 1877
* ''Colaspis obliqua'' Bowditch, 1921
* ''Colaspis obscura'' Fabricius, 1801
* ''Colaspis occidentalis'' (Linnaeus, 1758) ^g
* '' Colaspis orientalis'' Blake, 1967
* ''Colaspis otileensis'' Bowditch, 1921
* '' Colaspis pallipes'' Lefèvre, 1877
* '' Colaspis pantaria'' ( Bechyné, 1951)
** ''Colaspis pantaria emarginator'' ( Bechyné, 1951)
** ''Colaspis pantaria guayanensis'' ( Bechyné, 1997)
** ''Colaspis pantaria pantaria'' ( Bechyné, 1951)
** ''Colaspis pantaria vaticina'' ( Bechyné, 1997)
* ''Colaspis panzoensis'' ( Bechyné, 1951)
* ''Colaspis paracostata'' Blake, 1978
* '' Colaspis perfidia'' ( Bechyné, 1954)
* ''Colaspis perichrysea'' ( Bechyné, 1951)
* ''Colaspis perturbata'' Weise, 1921 ^g
** ''Colaspis perturbata antanossa'' ( Bechyné, 1950)
** ''Colaspis perturbata coloresignata'' ( Bechyné, 1951)
** ''Colaspis perturbata ostrina'' ( Bechyné, 1950)
** ''Colaspis perturbata perturbata'' Weise, 1921
** ''Colaspis perturbata phylis'' ( Bechyné, 1953)
* ''Colaspis pini'' Barber, 1937 ^{i c g b} (pine colaspis)
* ''Colaspis planicostata'' Blake, 1974 ^{i c g b}
* ''Colaspis pleuralis'' Weise, 1921
* ''Colaspis plicatula'' Jacoby, 1882
* ''Colaspis pohli'' ( Bechyné, 1955)
** ''Colaspis pohli major'' ( Bechyné, 1958)
** ''Colaspis pohli pohli'' ( Bechyné, 1955)
* '' Colaspis propinqua'' Lefèvre, 1885
* ''Colaspis prospectans'' ( Bechyné, 1958)
* ''Colaspis proteus'' Bechyné, 1949
* ''Colaspis pseudofavosa'' E. Riley, 1978 ^{i c g b}
* ''Colaspis pseudogeminata'' ( Bechyné, 1950)
* ''Colaspis pseudopruinosa'' Bechyné, 1949
* ''Colaspis pubiceps'' ( Bechyné, 1952)
* ''Colaspis punctigera'' Weise, 1921
* '' Colaspis punctipennis'' Bowditch, 1921
* ''Colaspis purpurala'' Blake, 1978
* ''Colaspis purpurea'' Blake, 1971
* ''Colaspis quattuordecimcostata'' Lefèvre, 1877
* ''Colaspis recurva'' Blake, 1974 ^{i c g b}
* ''Colaspis reflexomicans'' ( Bechyné, 1953)
* ''Colaspis romani'' Weise, 1921
* ''Colaspis rufipes'' Jacoby, 1900
* ''Colaspis rugifera'' Weise, 1921
* ''Colaspis rugulosa'' Lefèvre, 1891
* ''Colaspis rustica'' Boheman, 1859
** ''Colaspis rustica distortella'' ( Bechyné, 1952)
** ''Colaspis rustica rustica'' Boheman, 1859
** ''Colaspis rustica santoamarensis'' ( Bechyné, 1950)
** ''Colaspis rustica thoracophora'' ( Bechyné, 1952)
* ''Colaspis sanguinea'' Blake, 1977 ^{i c g}
* ''Colaspis sanjoseana'' ( Bechyné, 1950)
* ''Colaspis scintillifera'' Bechyné, 1949
** ''Colaspis scintillifera ascintillans'' ( Bechyné, 1950)
** ''Colaspis scintillifera scintillifera'' Bechyné, 1949
* ''Colaspis shuteae'' Blake, 1976
* ''Colaspis similaris'' Blake, 1976
* ''Colaspis similis'' Blake, 1977 ^{i c g}
* ''Colaspis simplex'' Weise, 1921
* ''Colaspis simplicipes'' Bechyné, 1950
* ''Colaspis skelleyi'' Riley, 2020
* ''Colaspis smaragdula'' Olivier, 1808
* ''Colaspis soror'' Weise, 1921
* '' Colaspis speciosa'' Lefèvre, 1885
* ''Colaspis spinigera'' Blake, 1976
* ''Colaspis stenorachis'' Blake, 1976
* ''Colaspis strigata'' Lefèvre, 1878
* '' Colaspis subacuta'' ( Bechyné, 1950)
* ''Colaspis submersa'' Bechyné, 1949
* '' Colaspis suffriani'' Weise, 1914
* ''Colaspis suggona'' Blake, 1977 ^{i c g b}
* ''Colaspis suilla'' Fabricius, 1801 ^{i c g b}
* '' Colaspis sulphuripes'' Lefèvre, 1877
** ''Colaspis sulphuripes matura'' ( Bechyné, 1951)
** ''Colaspis sulphuripes sulphuripes'' Lefèvre, 1877
* '' Colaspis surrubrensis'' ( Bechyné, 1950)
* ''Colaspis tabacilla'' ( Bechyné, 1951)
** ''Colaspis tabacilla berriensis'' ( Bechyné, 1951)
** ''Colaspis tabacilla puertocabellensis'' ( Bechyné, 1953)
** ''Colaspis tabacilla sedula'' ( Bechyné, 1951)
** ''Colaspis tabacilla tabacilla'' ( Bechyné, 1951)
* ''Colaspis taylori'' Bechyné, 1949
* ''Colaspis thomasi'' Riley, 2020
* ''Colaspis tibialis'' Lefèvre, 1885
* ''Colaspis townsendi'' Bowditch, 1921
* ''Colaspis trichopyga'' ( Bechyné, 1954)
** ''Colaspis trichopyga gravida'' ( Bechyné, 1955)
** ''Colaspis trichopyga trichopyga'' ( Bechyné, 1954)
* ''Colaspis tucumanensis'' Bowditch, 1921
* ''Colaspis unicolor'' Olivier, 1808
* ''Colaspis utinga'' ( Bechyné & Bechyné, 1961)
* '' Colaspis variabilis'' Blake, 1976
* '' Colaspis vieta'' ( Bechyné, 1950)
* ''Colaspis violetta'' Bechyné, 1949
* ''Colaspis viridiceps'' Schaeffer, 1933 ^{i c g b}
* '' Colaspis viridicollis'' Bowditch, 1921
* '' Colaspis viridipunctata'' Jacoby, 1900
* ''Colaspis viridissima'' Lefèvre, 1877
* ''Colaspis viriditincta'' Schaeffer, 1919 ^{i c g b}
* '' Colaspis virulenta'' ( Bechyné, 1953)
* '' Colaspis willincki'' ( Bechyné, 1950)
* ''Colaspis yucatana'' Jacoby, 1890
* ''Colaspis yungarum'' ( Bechyné, 1951)
* ''Colaspis zanthophaia'' Blake, 1976
* ''Colaspis zilchi'' ( Bechyné, 1954)
* ''Colaspis zischkai'' ( Bechyné, 1951)

These four extinct species belong to the genus ''Colaspis'':
* †''Colaspis aetatis'' Wickham, 1911
* †''Colaspis diluvialis'' Wickham, 1914
* †''Colaspis luti'' Scudder, 1900
* †''Colaspis proserpina'' Wickham, 1914

Synonyms:
* ''Colaspis aemula'' Weise, 1921: synonym of ''Colaspis flavipes'' Olivier, 1808
* ''Colaspis cartwrighti'' Blake, 1977: synonym of ''Tymnes tricolor'' (Fabricius, 1792)
* ''Colaspis coneja'' Kolbe, 1901: synonym of '' Nodocolaspis columbica'' ( Jacoby, 1900)
* ''Colaspis costipennis'' Lefèvre, 1877: renamed to '' Colaspis lophodes'' Blake, 1974
* ''Colaspis elegans'' Jacoby, 1900: synonym of ''Nodocolaspis costipennis'' (Lefèvre, 1877)
* ''Colaspis flavipes'' (Fabricius, 1781): synonym of ''Colaspis occidentalis'' (Linnaeus, 1758)
* ''Colaspis pectoralis'' Lefèvre, 1885: synonym of '' Coytiera (Campylochira) zikani'' Bechyné, 1950
* ''Colaspis prasina'' Lefèvre, 1878: synonym of ''Colaspis flavicornis'' (Fabricius, 1787)

Data sources: i = ITIS, c = Catalogue of Life, g = GBIF, b = Bugguide.net

References

External links

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''Colaspis''
at insectoid.info

{{Taxonbar, from=Q2331275

Chrysomelidae genera
Eumolpinae
Beetles of North America
Beetles of South America
Taxa named by Johan Christian Fabricius
Eocene insects
Prehistoric insects of North America