Clymene Dolphin
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The Clymene dolphin (''Stenella clymene''), in older texts known as the short-snouted spinner dolphin, is a dolphin endemic to the
Atlantic Ocean The Atlantic Ocean is the second largest of the world's five borders of the oceans, oceanic divisions, with an area of about . It covers approximately 17% of Earth#Surface, Earth's surface and about 24% of its water surface area. During the ...
. It is the only confirmed case of hybrid speciation in marine mammals, descending from the spinner dolphin and the striped dolphin.


Taxonomy

The Clymene dolphin was first formally described by
John Edward Gray John Edward Gray (12 February 1800 – 7 March 1875) was a British zoologist. He was the elder brother of zoologist George Robert Gray and son of the pharmacologist and botanist Samuel Frederick Gray (1766–1828). The same is used for a z ...
in 1846, although, unusually, he did not assign it its current name until four years later, in 1850. From then on, until a reassessment in 1981, the Clymene dolphin was regarded as a subspecies of the spinner dolphin (''Stenella longirostris''). In 1981, Perrin ''et al.'' asserted the Clymene's existence as separate species. Until this time, because Clymenes are relatively remote and were regarded as very similar to the more accessible spinners, they were never heavily studied. Anatomical and behavioral traits suggested that this species is a hybrid of the spinner dolphin and striped dolphin (''Stenella coeruleoalba''), and DNA testing has shown that it is indeed a hybrid species. The common and scientific names are probably derived from the Greek Oceanid Clymene, although it has also been argued that it may instead come from the Greek word for "notorious".


Description

The Clymene dolphin looks very similar to the spinner dolphin. At close quarters, it is possible to observe that the beak of the Clymene is slightly shorter than that of its relative. The dorsal fin is also less erect and triangular. The basic color of the Clymene dolphin is "cetacean neapolitan"; it occurs in three shaded layers, the underside being white. Next, a strip of light grey runs from just above the beak, round either side of the eye back to the tail stock, where the band thickens. The top layer, from the forehead, along the back to the dorsal fin, and down to the top of the tail stock, is a dark grey. The beak, lips, and flippers are also dark grey in color. Clymene dolphins grow to about in length and in weight.


Behavior and biology

Clymene dolphins spend most of their lives in waters over in depth, but occasionally move into shallower, coastal regions. They feed on squid and small schooling fish, hunting either at night, or in mesopelagic waters where there is only limited light. Predators include cookie-cutter sharks, as evidenced by bite marks seen on a number of animals. Clymenes are fairly active dolphins. They do spin longitudinally when jumping clear of the water, but not with as much regularity and complexity as the spinner dolphin. They will also approach boats and ride bow waves. Group sizes vary from just four up to around 150 individuals, although about forty is typical. Many of these groups appear to be single-sex, and also to be segregated by the approximate age of the individuals. Clymene dolphins are also highly vocal, making short whistles in a range of 6–19 kHz. No figures are available for the size of animals at birth. Gestation, lactation, and maturation periods are all unknown, but are unlikely to vary greatly from others in the genus ''Stenella''. Their longevity is also unknown, although at least one sixteen-year-old individual has been reported from a stranding.


Population and distribution

The Clymene dolphin is endemic to the Atlantic Ocean. Its full range is still poorly understood, particularly at its southern end. The species certainly prefer
temperate In geography, the temperate climates of Earth occur in the middle latitudes (approximately 23.5° to 66.5° N/S of the Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ran ...
and
tropical The tropics are the regions of Earth surrounding the equator, where the sun may shine directly overhead. This contrasts with the temperate or polar regions of Earth, where the Sun can never be directly overhead. This is because of Earth's ax ...
waters. The northern end of the range runs roughly from
New Jersey New Jersey is a U.S. state, state located in both the Mid-Atlantic States, Mid-Atlantic and Northeastern United States, Northeastern regions of the United States. Located at the geographic hub of the urban area, heavily urbanized Northeas ...
east-southeast to southern
Morocco Morocco, officially the Kingdom of Morocco, is a country in the Maghreb region of North Africa. It has coastlines on the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to Algeria–Morocc ...
. The southern tip runs from somewhere around
Angola Angola, officially the Republic of Angola, is a country on the west-Central Africa, central coast of Southern Africa. It is the second-largest Portuguese-speaking world, Portuguese-speaking (Lusophone) country in both total area and List of c ...
to
Rio de Janeiro Rio de Janeiro, or simply Rio, is the capital of the Rio de Janeiro (state), state of Rio de Janeiro. It is the List of cities in Brazil by population, second-most-populous city in Brazil (after São Paulo) and the Largest cities in the America ...
. They appear to prefer deep water. Numerous sightings have been recorded in the
Gulf of Mexico The Gulf of Mexico () is an oceanic basin and a marginal sea of the Atlantic Ocean, mostly surrounded by the North American continent. It is bounded on the northeast, north, and northwest by the Gulf Coast of the United States; on the southw ...
. The species has not been sighted, however, in the
Mediterranean Sea The Mediterranean Sea ( ) is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean basin and almost completely enclosed by land: on the east by the Levant in West Asia, on the north by Anatolia in West Asia and Southern Eur ...
. The total population is unknown. The only population estimate available is for the north part of the Gulf of Mexico, where a count of 6,500 individuals was reported. However, it is suspected that there are three well-defined populations in the Atlantic Ocean, located in the North Atlantic, South Atlantic, and the Gulf of Mexico. As of more recent research, it is presumed that individuals from the South Atlantic and the Gulf of Mexico move between these two populations, while the North Atlantic population seems to be more isolated. The species may naturally be rare in comparison with others in the genus ''Stenella''.


Human interaction

Some individuals have been killed from directed fisheries in the
Caribbean The Caribbean ( , ; ; ; ) is a region in the middle of the Americas centered around the Caribbean Sea in the Atlantic Ocean, North Atlantic Ocean, mostly overlapping with the West Indies. Bordered by North America to the north, Central America ...
and others may have been caught in nets off
West Africa West Africa, also known as Western Africa, is the westernmost region of Africa. The United Nations geoscheme for Africa#Western Africa, United Nations defines Western Africa as the 16 countries of Benin, Burkina Faso, Cape Verde, The Gambia, Gha ...
.


Conservation

The West African population of the Clymene dolphin is listed in Appendix IIAppendix II
" of the Convention on the Conservation of Migratory Species of Wild Animals (CMS). As amended by the Conference of the Parties in 1985, 1988, 1991, 1994, 1997, 1999, 2002, 2005, and 2008. Effective: 5 March 2009.
of the convention on the Conservation of Migratory Species of Wild Animals ( CMS), since it has an unfavorable conservation status or would benefit significantly from international co-operation organized by tailored agreements. The Clymene dolphin is covered by the Memorandum of Understanding Concerning the Conservation of the Manatee and Small Cetaceans of Western Africa and Macaronesia.


See also

*
Marine mammal Marine mammals are mammals that rely on marine ecosystems for their existence. They include animals such as cetaceans, pinnipeds, sirenians, sea otters and polar bears. They are an informal group, unified only by their reliance on marine enviro ...
* List of cetaceans * List of genetic hybrids


References


Bibliography

* Carwardine, Mark. ''Whales Dolphins and Porpoises'', Dorling Kindersley Handbooks, . * Dee, Eileen Mary, and Mark McGinley. 2010
''Clymene dolphin''
Encyclopedia of Earth. topic ed. C. Michael Hogan. ed. Cutler J. Cleveland, NCSE, Washington DC * Jefferson, Thomas A. "Clymene Dolphin" in ''Encyclopedia of Marine Mammals'', 234–236. * Perrin and Mead. (1994). "Clymene Dolphin" in ''Handbook of Marine Mammals''. 5: 161–171. * Reeves, Stewart, Clapham, and Powell. ''National Audubon Society Guide to Marine Mammals of the World'', . {{Taxonbar, from=Q838940 Clymene dolphin Cetaceans of the Atlantic Ocean Clymene dolphin Taxa named by John Edward Gray Mammal hybrids Dolphins Oceanic dolphins