''Clinocottus'' is a

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

of marine

ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fish or actinopterygians, is a class of bony fish that comprise over 50% of living vertebrate species. They are so called because of their lightly built fins made of webbings of sk ...

es belonging to the

family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...

Cottidae The Cottidae are a family of fish in the superfamily Cottoidea, the sculpins. It is the largest sculpin family, with about 275 species in 70 genera.Kane, E. A. and T. E. Higham. (2012)Life in the flow lane: differences in pectoral fin morphology ...

, the typical sculpins. They are nearshore benthic fishes native to the northeastern

Pacific Ocean The Pacific Ocean is the largest and deepest of Earth's five Borders of the oceans, oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean, or, depending on the definition, to Antarctica in the south, and is ...

. They are mentioned as sharpnose sculpins.''Clinocottus'' Gill, 1861

ITIS The Integrated Taxonomic Information System (ITIS) is an American partnership of federal agencies designed to provide consistent and reliable information on the taxonomy of biological species. ITIS was originally formed in 1996 as an interagenc ...

Taxonomy

''Clinocottus'' was first proposed as a

monospecific genus In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispe ...

in 1861 by the American

zoologist Zoology ( , ) is the scientific study of animals. Its studies include the structure, embryology, classification, habits, and distribution of all animals, both living and extinct, and how they interact with their ecosystems. Zoology is one ...

Theodore Gill Theodore Nicholas Gill (March 21, 1837 – September 25, 1914) was an American ichthyologist, mammalogist, malacologist, and librarian. Career Born and educated in New York City under private tutors, Gill early showed interest in natural hist ...

with ''Oligocottus analis'', which had been described in 1858 by Charles Frédéric Girard from

Monterey, California Monterey ( ; ) is a city situated on the southern edge of Monterey Bay, on the Central Coast (California), Central Coast of California. Located in Monterey County, California, Monterey County, the city occupies a land area of and recorded a popu ...

, as its only species. Molecular data indicate that this genus is

polyphyletic A polyphyletic group is an assemblage that includes organisms with mixed evolutionary origin but does not include their most recent common ancestor. The term is often applied to groups that share similar features known as Homoplasy, homoplasies ...

paraphyletic Paraphyly is a taxonomic term describing a grouping that consists of the grouping's last common ancestor and some but not all of its descendant lineages. The grouping is said to be paraphyletic ''with respect to'' the excluded subgroups. In co ...

, it does not constitute a natural assemblage within Cottidae. Workers have found that the lavender sculpin (''Leiocottus hirundo'') is the

sister taxon In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and ...

to ''C. analis'', a species which is not closely related to ''C. acuticeps''. In turn, ''C. acuticeps'' is determined to be basal to '' Artedius'', or to the

clade In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach t ...

referred to as Oligocottinae. The 5th edition of ''

Fishes of the World ''Fishes of the World'' is a standard reference for the systematics of fishes. It was first written in 1976 by the American ichthyologist Joseph S. Nelson (1937–2011). Now in its fifth edition (2016), the work is a comprehensive overview of t ...

'' classifies the genus ''Clinocottus'' within the

subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end botanical subfamily names with "-oideae", and zo ...

Cottinae Cottinae is a subfamily of ray-finned fishes belonging to the Family (biology), family Cottidae, the typical sculpins. The subfamily has species throughout the northern hemisphere in both marine and freshwater habitats. Genera The following gener ...

of the family Cottidae, however, other authors classify the genus within the subfamily Oligocottinae of the family

Psychrolutidae The fish family Psychrolutidae (commonly known as blobfishes, flathead sculpins, or tadpole sculpins) contains over 35 recognized species in 8 genera. This family consists of bottom-dwelling marine sculpins shaped like tadpoles, with large heads ...

Etymology

''Clinocottus'' combines '' Clinus'' a genus of blennies with '' Cottus'', probably because Girard though the rounded head of the mosshead sculpin was reminiscent of the ehad shape of some

gobies Gobiidae or gobies is a family of bony fish in the order Gobiiformes, one of the largest fish families comprising over 2,000 species in more than 200 genera. Most of gobiid fish are relatively small, typically less than in length, and the fam ...

and blennies.

Species

There are currently five recognized species in this genus: * '' Clinocottus acuticeps'' ( C. H. Gilbert, 1896) (Sharpnose sculpin) * '' Clinocottus analis'' ( Girard, 1858) (Woolly sculpin) * '' Clinocottus embryum'' ( D. S. Jordan & Starks, 1895) (Calico sculpin) * '' Clinocottus globiceps'' (Girard, 1858) (Mosshead sculpin) * '' Clinocottus recalvus'' ( Greeley, 1899) (Bald sculpin)

Characteristics

''Clinocottus'' sculpins are characterised by having the

anus In mammals, invertebrates and most fish, the anus (: anuses or ani; from Latin, 'ring' or 'circle') is the external body orifice at the ''exit'' end of the digestive tract (bowel), i.e. the opposite end from the mouth. Its function is to facil ...

in a forward position between the

anal fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish swim. Apart from the tail or caudal fin, fish fins have no direct connection with the back bone and are supported o ...

and the

pelvic fin Pelvic fins or ventral fins are paired fins located on the ventral (belly) surface of fish, and are the lower of the only two sets of paired fins (the other being the laterally positioned pectoral fins). The pelvic fins are homologous to the hi ...

s, they have a heavy and blunt penis and in both sexes the anal fin is unmodified. ''C. analis'' and ''C. acuticeps'' have pointed heads, which is regarded as a basal feature and the remaining 3 species have sizeable, rounded heads. This is suggestive of the paraphyly of the group. These are small sculpins, the largest species is the mosshead sculpin with a maximum published

total length Fish measurement is the measuring of individual fish and various parts of their anatomies, for data used in many areas of ichthyology, including taxonomy and fishery biology. Overall length Standard length (SL) is the length of a fish measured f ...

of and the smallest is the calico sculpin with a maximum published

of .

Distribution

''Clinocottus'' sculpins are found in the eastern Pacific Ocean where they are found in the intertidal zone.

References

{{Taxonbar, from=Q2047619 Cottinae Taxa named by Theodore Gill