Charaxes Zoolina
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''Charaxes zoolina'', the club-tailed emperor or club-tailed charaxes, is a
butterfly Butterflies are winged insects from the lepidopteran superfamily Papilionoidea, characterized by large, often brightly coloured wings that often fold together when at rest, and a conspicuous, fluttering flight. The oldest butterfly fossi ...
of the family
Nymphalidae The Nymphalidae are the largest family of butterflies, with more than 6,000 species distributed throughout most of the world. Belonging to the superfamily Papilionoidea, they are usually medium-sized to large butterflies. Most species ha ...
. It is found in southern Africa."''Charaxes'' Ochsenheimer, 1816"
at Markku Savela's ''Lepidoptera and Some Other Life Forms''


Description

The
wingspan The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the opposite wingtip. For example, the Boeing 777–200 has a wingspan of , and a wandering albatross (''Diomedea exulans'') caught in 1965 had a wingsp ...
is 40–45 mm in males and 50–58 mm in females. The species of this group 'zoolina'' groupdiffer from all others in the hindwing having in the male only one long tail, at vein 2, but in the female two, at veins 2 and 4. ''Ch. zoolina'' occurs in three local races, each with a greenish and a brown or brown-yellow form. The greenish white forms differ from '' kahldeni'' in the broad dark marginal band on the upperside of the wing and the yellow-brown forms from ''homeyeri'' [= ''kahldeni'' in the absence of the longitudinal stripe in cellule 1 c on the hindwing beneath.- ''zoolina'' Westw. wings with the ground-colour greenish white; forewing above with the costal margin, a transverse spot at the end of the cell and the apical area black, the latter more or less spotted with white; hindwing with a black marginal band, 4-5mm. in breadth, with small white dots. Wings beneath almost as above, but with larger white spots in the marginal bands, common black median band and black longitudinal stripe in cellule 1 c of the hindwing; distal margin of the hindwing scarcely appreciably angled at vein 4. The female is larger and has the light spots in the marginal bands much larger and joined together; hindwing above with yellowish marginal streaks. Natal to Uganda and Abyssinia, f. ''neanthes'' Hew. The ground-colour is lighter or darker ochre-yellow and the black markings of the type-form are replaced by ferruginous or dark brown; occasionally they are black, ab. ''obscuratus'' Suff. Under surface irrorated with dark, without the longitudinal stripe in cellule 1 c of the hindwing. ab. ''ehmckei'' Dew. Distal margin of the hindwing distinctly angled at vein 4. Otherwise only differs from neanthes in having both wings above broadly whitish at the base and beneath without distinct median band. Angola, f. ''phanera'' Jord. has in the male a distinct angle at vein 4 of the hindwing and less extended black markings on the forewing, but otherwise agrees with ''zoolina''. Angola. ab. ''betsimisaraka'' Luc. occurs on Madagascar and only differs from ''zoolina'' in the hindwing of the male having a distinct though small tooth at the end of vein 4 and in the transverse streak at the end of the cell of the forewing only reaching the base of vein 3. f. ''betanimena'' Luc. corresponds to f. ''neanthes'', but is darker and with the marginal band darker rust-brown and more continuous. The hindwing in the male distinctly toothed at vein 4. Wings beneath without median band, or with this sharply pronounced and margined with white [is] ab. ''andriba'' Ward. Madagascar.


Biology

Flight period is from October to June. The habitat is Afrotropics, forest. Larvae feed on ''Entada'' - '' Entada spicata'', ''
Entada abyssinica ''Entada'' is a genus of flowering plants in the family Fabaceae, in the mimosoid clade of the subfamily Caesalpinioideae. It consists of some 30 species of trees, shrubs and tropical lianas. About 21 species are known from Africa, six from A ...
'', '' Entada natalensis'', '' Acacia natalitia'', ''
Acacia schweinfurthi ''Acacia'', commonly known as wattles or acacias, is a genus of about of shrubs and trees in the subfamily Mimosoideae of the pea family Fabaceae. Initially, it comprised a group of plant species native to Africa, South America, and Australa ...
'', '' Acacia brevispica'', and '' Acacia kraussiana''.


Subspecies

Listed alphabetically. *''C. z. betsimisaraka'' Lucas, 1872 (Madagascar) *''C. z. ehmckei'' Homeyer and Dewitz, 1882 (Angola) *''C. z. mafugensis'' Jackson, 1956Jackson, T.H.E. 1956. Notes on the Rhopalocera of the Kigezi district of Uganda with descriptions of new species and subspecies. ''Journal of the East Africa Natural History Society'' 23: 63-102. (south-western Uganda, Rwanda, Burundi, Democratic Republic of the Congo, north-western Tanzania) *''C. z. zoolina'' (Westwood,
850 __NOTOC__ Year 850 ( DCCCL) was a common year starting on Wednesday of the Julian calendar. Events By place Europe * February 1 – King Ramiro I dies in his palace at Santa María del Naranco (near Oviedo), after an 8-year reign ...
(southern Sudan, southern Ethiopia, Somalia, Uganda, Kenya, Tanzania, Malawi, Zambia, Mozambique, Zimbabwe, northern Botswana, north-eastern Namibia, South Africa, Eswatini)


Taxonomy

''Charaxes zoolina'' is a member of the
species group In biology, a species complex is a group of closely related organisms that are so similar in appearance and other features that the boundaries between them are often unclear. The taxa in the complex may be able to hybridize readily with each oth ...
''Charaxes zoolina''. The
clade In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach t ...
members are *''Charaxes zoolina'' Nominate *'' Charaxes kahldeni'' The group differs from all the others in ''Charaxes'' in the male having a tail on vein 2, but the female having two long tails on veins 2, 4. There are two different forms, one with black markings on a greenish white base colour (wet-season form) and one with reddish-brown markings on red-yellow whitish colour (dry-season form). These forms can be from eggs of the same female.


References


Seitz, A. ''Die Gross-Schmetterlinge der Erde'' 13: Die Afrikanischen Tagfalter. Plate XIII 32
''b'' *
Victor Gurney Logan Van Someren Victor Gurney Logan Van Someren (1886 in Melbourne – 24 July 1976) was a zoologist and entomologist. Van Someren was born in Australia. He attended George Watson's College and studied zoology at University of Edinburgh. He was also a dentis ...
, 1974 Revisional notes on African ''Charaxes'' (Lepidoptera: Nymphalidae). Part IX. ''Bulletin of the British Museum of Natural History'' (Entomology) 29 (8):415-487


External links


Images of ''C. zoolina zoolina''
Royal Museum for Central Africa The Royal Museum for Central Africa (RMCA) (; ; ), communicating under the name AfricaMuseum since 2018, is an ethnography and natural history museum situated in Tervuren in Flemish Brabant, Belgium, just outside Brussels. It was originally b ...
(Albertine Rift Project)
Images of ''C. zoolina mafugensis''
(Albertine Rift Project)

Images
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''Charaxes zoolina zoolina''
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''Charaxes zoolina betsimisaraka''
''Charaxes mafugensis'' images
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''Charaxes ehmckei'' images
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''Charaxes betsimisaraka'' images
at BOLD zoolina Butterflies described in 1850 Butterflies of Africa Taxa named by John O. Westwood {{Charaxinae-stub