The Cetopsidae are a small
family
Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
of
catfish
Catfish (or catfishes; order Siluriformes or Nematognathi) are a diverse group of ray-finned fish. Named for their prominent barbels, which resemble a cat's whiskers, catfish range in size and behavior from the three largest species alive, ...
es (
order
Order, ORDER or Orders may refer to:
* Categorization, the process in which ideas and objects are recognized, differentiated, and understood
* Heterarchy, a system of organization wherein the elements have the potential to be ranked a number of d ...
Siluriformes), commonly called the whale catfishes.
Taxonomy
This family contains five
genera
Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial ...
divided into two subfamilies, Cetopsinae and Helogeneinae.
Helogeneinae was previously a family-level group, but now it has been reclassified as a subfamily of Cetopsidae. This subfamily contains four species in the genus ''
Helogenes''.
The subfamily Cetopsinae contains four genera. ''
Cetopsidium'' contains six species, ''
Cetopsis'' contains 21 species, ''
Denticetopsis'' contains seven species, and ''
Paracetopsis'' contains three species; this makes a total of 37 cetopsines.
The genera have been changed as recently as 2005 with the genera ''Bathycetopsis'', ''Hemicetopsis'', and ''Pseudocetopsis'' set in
synonymy with ''Cetopsis'' and the description of the new genus ''Cetopsidium''.
''Cetopsidium'' is the sister group to the rest of Cetopsinae. ''Denticetopsis'' forms the next sister group to the remaining cetopsine genera. The tribes Cetopsidiini, Denticetopsini, and Cetopsini are erected for the cetopsine genera.
Distribution
Cetopsids have a wide distribution in
South America
South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the souther ...
.
Latitudinally, cetopsines extend from northern
Colombia to central
Argentina
Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, t ...
. They inhabit major habitats such as the
Orinoco River
The Orinoco () is one of the longest rivers in South America at . Its drainage basin, sometimes known as the Orinoquia, covers , with 76.3 percent of it in Venezuela and the remainder in Colombia. It is the fourth largest river in the wo ...
,
Amazon River, and the
Guyanas.
In trans-
Andean
The Andes, Andes Mountains or Andean Mountains (; ) are the longest continental mountain range in the world, forming a continuous highland along the western edge of South America. The range is long, wide (widest between 18°S – 20°S l ...
South America, cetopsines are found along the
Pacific
The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the continen ...
slope from the
Jurubidá River of Colombia south to the
Tumbes River of northern
Peru
, image_flag = Flag of Peru.svg
, image_coat = Escudo nacional del Perú.svg
, other_symbol = Great Seal of the State
, other_symbol_type = Seal (emblem), National seal
, national_motto = "Fi ...
. Along the
Caribbean trans-Andean versant, species of cetopsines occur from the
Sinú River of northwestern Colombia east to the
Lake Maracaibo basin of northwestern
Venezuela
Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivariana de Venezuela), is a country on the northern coast of South America, consisting of a continental landmass and many islands and islets in ...
. East of the Andean
Cordilleras, the Cetopsinae occur in the
Aroa and
Yaracuy River basins along the Caribbean versant of northern Venezuela, through the Orinoco River system and the coastal rivers of the
Guianas, south through the Amazon basin to the southern portions of the
Río de la Plata
The Río de la Plata (, "river of silver"), also called the River Plate or La Plata River in English, is the estuary formed by the confluence of the Uruguay River and the Paraná River at Punta Gorda. It empties into the Atlantic Ocean and f ...
basin. Cetopsines also occur in the
Juquiá River
The Juquiá River is a river of São Paulo state in southeastern Brazil.
See also
*List of rivers of São Paulo
List of rivers in São Paulo (Brazilian State).
The list is arranged by drainage basin from north to south, with respective tributar ...
basin of the state of
São Paulo
São Paulo (, ; Portuguese for 'Saint Paul') is the most populous city in Brazil, and is the capital of the state of São Paulo, the most populous and wealthiest Brazilian state, located in the country's Southeast Region. Listed by the Ga ...
and the
São Francisco River
The São Francisco River (, ) is a large river in Brazil. With a length of , it is the longest river that runs entirely in Brazilian territory, and the fourth longest in South America and overall in Brazil (after the Amazon, the Paraná and t ...
basin of eastern
Brazil
Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
.
''Helogenes'', the single genus of Helogeneinae, occurs through much of the Amazon basin, the southern portions of the Orinoco River basin, the coastal rivers of the Guianas, and at least the lower portions of the
Tocantins River.
Description
The family Cetopsidae includes species of small- to medium-sized catfishes which share an anal fin with a long base, the lack of nasal
barbels, and, usually, a lack of
dorsal and
pectoral fin
Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as ...
spines. In Cetopsinae, the
swim bladder is highly reduced and is enclosed in a bony capsule.
Cetopsines lack an
adipose fin
Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as s ...
, while it may be present (though small) in Helogeneinae.
Many species are characterized by small eyes obscured by a thick, overlying
integument
In biology, an integument is the tissue surrounding an organism's body or an organ within, such as skin, a husk, shell, germ or rind.
Etymology
The term is derived from ''integumentum'', which is Latin for "a covering". In a transferred, or ...
that make them appear blind.
The dorsal and pectoral fins usually lack spines, except in ''Cetopsidium'' (both present) and ''Denticetopsis'' (only pectoral spines present).
In most species of ''Cetopsis'' and ''Cetopsidium'', mature males have a convex margin to their anal fin and elongated distal filaments of the dorsal and pectoral fin spines.
The maximum length of this family is about
SL in Cetopsinae.
''Cetopsidium'' species are generally smaller in body size than ''Cetopsis''.
The maximum length in ''Helogenes'' species is .
Ecology
Most cetopsids feed on
insect
Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (head, thorax and abdomen), three pairs ...
s.
''C. candiru'' and ''C. coecutiens'' are well known to have extremely voracious appetites. These fish will attack
carrion
Carrion () is the decaying flesh of dead animals, including human flesh.
Overview
Carrion is an important food source for large carnivores and omnivores in most ecosystems. Examples of carrion-eaters (or scavengers) include crows, vultures ...
, other living fish while in
gillnet
Gillnetting is a fishing method that uses gillnets: vertical panels of netting that hang from a line with regularly spaced floaters that hold the line on the surface of the water. The floats are sometimes called "corks" and the line with corks is ...
s, and even people.
Some people erroneously believe these fish are
parasitic
Parasitism is a close relationship between species, where one organism, the parasite, lives on or inside another organism, the host, causing it some harm, and is adapted structurally to this way of life. The entomologist E. O. Wilson ha ...
like
parasitic catfishes.
References
{{Taxonbar, from=Q761337
Fish of South America
Catfish families
Taxa named by Pieter Bleeker