The Carolinian forest refers to a life zone in eastern

North America North America is a continent in the Northern Hemisphere, Northern and Western Hemisphere, Western hemispheres. North America is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South Ameri ...

characterized primarily by the predominance of

deciduous In the fields of horticulture and botany, the term deciduous () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed Leaf, leaves, usually in the autumn; to the shedding of petals, aft ...

(broad-leaf) forest. The term "Carolinian", which is most commonly used in Canada, refers to the deciduous forests which span across much of the eastern United States from

North Carolina North Carolina ( ) is a U.S. state, state in the Southeastern United States, Southeastern region of the United States. It is bordered by Virginia to the north, the Atlantic Ocean to the east, South Carolina to the south, Georgia (U.S. stat ...

northward into

southern Ontario Southern Ontario is a Region, primary region of the Canadian Provinces and territories of Canada, province of Ontario. It is the most densely populated and southernmost region in Canada, with approximately 13.5 million people, approximately 36% o ...

, Canada. These deciduous forests in the United States and southern Ontario share many similar characteristics and species hence their association. Today the term is often used to refer to the Canadian portion (northern limit) of the deciduous forest region while the portion in the United States is often referred to as the "Eastern deciduous forest".

Location and extent

The Carolinian zone spans across much of the eastern United States, with extensive coverage in the Virginias,

Kentucky Kentucky (, ), officially the Commonwealth of Kentucky, is a landlocked U.S. state, state in the Southeastern United States, Southeastern region of the United States. It borders Illinois, Indiana, and Ohio to the north, West Virginia to the ...

Tennessee Tennessee (, ), officially the State of Tennessee, is a landlocked U.S. state, state in the Southeastern United States, Southeastern region of the United States. It borders Kentucky to the north, Virginia to the northeast, North Carolina t ...

Maryland Maryland ( ) is a U.S. state, state in the Mid-Atlantic (United States), Mid-Atlantic region of the United States. It borders the states of Virginia to its south, West Virginia to its west, Pennsylvania to its north, and Delaware to its east ...

Delaware Delaware ( ) is a U.S. state, state in the Mid-Atlantic (United States), Mid-Atlantic and South Atlantic states, South Atlantic regions of the United States. It borders Maryland to its south and west, Pennsylvania to its north, New Jersey ...

Pennsylvania Pennsylvania, officially the Commonwealth of Pennsylvania, is a U.S. state, state spanning the Mid-Atlantic (United States), Mid-Atlantic, Northeastern United States, Northeastern, Appalachian, and Great Lakes region, Great Lakes regions o ...

, parts of southern

New York New York most commonly refers to: * New York (state), a state in the northeastern United States * New York City, the most populous city in the United States, located in the state of New York New York may also refer to: Places United Kingdom * ...

state,

Connecticut Connecticut ( ) is a U.S. state, state in the New England region of the Northeastern United States. It borders Rhode Island to the east, Massachusetts to the north, New York (state), New York to the west, and Long Island Sound to the south. ...

, and

Rhode Island Rhode Island ( ) is a state in the New England region of the Northeastern United States. It borders Connecticut to its west; Massachusetts to its north and east; and the Atlantic Ocean to its south via Rhode Island Sound and Block Is ...

, eastern

Ohio Ohio ( ) is a U.S. state, state in the Midwestern United States, Midwestern region of the United States. It borders Lake Erie to the north, Pennsylvania to the east, West Virginia to the southeast, Kentucky to the southwest, Indiana to the ...

, and small parts of southern

Michigan Michigan ( ) is a peninsular U.S. state, state in the Great Lakes region, Great Lakes region of the Upper Midwest, Upper Midwestern United States. It shares water and land boundaries with Minnesota to the northwest, Wisconsin to the west, ...

Indiana Indiana ( ) is a U.S. state, state in the Midwestern United States, Midwestern region of the United States. It borders Lake Michigan to the northwest, Michigan to the north and northeast, Ohio to the east, the Ohio River and Kentucky to the s ...

, and western

. It extends up into

Southern Ontario Southern Ontario is a Region, primary region of the Canadian Provinces and territories of Canada, province of Ontario. It is the most densely populated and southernmost region in Canada, with approximately 13.5 million people, approximately 36% o ...

, Canada which is located in the fertile

ecozone An ecozone may refer to: * Biogeographic realm, the broadest biogeographic division of Earth's land surface (referred to as ''ecozone'' by BBC) * Biome, a large collection of flora and fauna occupying a major habitat * Bioregion, an ecologically ...

of the Mixedwood Plains and includes ecodistricts 7E-1 to 7E-6. Trees found here include various species of ash,

birch A birch is a thin-leaved deciduous hardwood tree of the genus ''Betula'' (), in the family Betulaceae, which also includes alders, hazels, and hornbeams. It is closely related to the beech- oak family Fagaceae. The genus ''Betula'' contains 3 ...

chestnut The chestnuts are the deciduous trees and shrubs in the genus ''Castanea'', in the beech family Fagaceae. The name also refers to the edible nuts they produce. They are native to temperate regions of the Northern Hemisphere. Description ...

hickory Hickory is a common name for trees composing the genus ''Carya'', which includes 19 species accepted by ''Plants of the World Online''. Seven species are native to southeast Asia in China, Indochina, and northeastern India (Assam), and twelve ...

oak An oak is a hardwood tree or shrub in the genus ''Quercus'' of the beech family. They have spirally arranged leaves, often with lobed edges, and a nut called an acorn, borne within a cup. The genus is widely distributed in the Northern Hemisp ...

, and

walnut A walnut is the edible seed of any tree of the genus '' Juglans'' (family Juglandaceae), particularly the Persian or English walnut, '' Juglans regia''. They are accessory fruit because the outer covering of the fruit is technically an i ...

; tallest of all is the

tulip tree ''Liriodendron'' () is a genus of two species of characteristically large trees, deciduous over most of their populations, in the magnolia family (Magnoliaceae). These trees are widely known by the common name tulip tree or tuliptree for their ...

. Fruit trees native to this zone include the pawpaw. Animal life includes

raccoon The raccoon ( or , ''Procyon lotor''), sometimes called the North American, northern or common raccoon (also spelled racoon) to distinguish it from Procyonina, other species of raccoon, is a mammal native to North America. It is the largest ...

opossums Opossums () are members of the marsupial order Didelphimorphia () endemic to the Americas. The largest order of marsupials in the Western Hemisphere, it comprises 126 species in 18 genera. Opossums originated in South America and entered North A ...

squirrel Squirrels are members of the family Sciuridae (), a family that includes small or medium-sized rodents. The squirrel family includes tree squirrels, ground squirrels (including chipmunks and prairie dogs, among others), and flying squirrel ...

s (including the relatively rare

southern flying squirrel The southern flying squirrel (''Glaucomys volans''), also known commonly as the assapan, is a species of squirrel in the family Sciuridae. ''G. volans'' is one of three species of flying squirrels found in North America. It is found in deciduou ...

nuthatch The nuthatches () constitute a genus, ''Sitta'', of small passerine birds belonging to the family Sittidae. Characterised by large heads, short tails, and powerful bills and feet, nuthatches advertise their territory using loud, simple songs. Mo ...

es, and

chickadee The chickadees are a group of North American birds in the family Paridae included in the genus '' Poecile''. Species found in North America are referred to as chickadees; species found elsewhere in the world are called tits. They are small-si ...

Carolinian Canada

The Carolinian forest in Canada is located at the southern tip of Ontario between Lake Erie, Lake Huron and Lake Ontario. The region contains an extremely high

biodiversity Biodiversity is the variability of life, life on Earth. It can be measured on various levels. There is for example genetic variability, species diversity, ecosystem diversity and Phylogenetics, phylogenetic diversity. Diversity is not distribut ...

of species, over 500 of which are considered rare.

Climate

The reason for the high biodiversity in this region is its unique climate; ranging from

humid subtropical A humid subtropical climate is a subtropical -temperate climate type, characterized by long and hot summers, and cool to mild winters. These climates normally lie on the southeast side of all continents (except Antarctica), generally between la ...

to a mild

humid continental Humidity is the concentration of water vapor present in the air. Water vapor, the gaseous state of water, is generally invisible to the human eye. Humidity indicates the likelihood for precipitation, dew, or fog to be present. Humidity depe ...

. The Carolinian forest of Ontario has the warmest average annual temperatures, the longest frost-free seasons, and the mildest winters of any region in Ontario and Canada in general. This distinctive climate is largely due to the nearby

Great Lakes The Great Lakes, also called the Great Lakes of North America, are a series of large interconnected freshwater lakes spanning the Canada–United States border. The five lakes are Lake Superior, Superior, Lake Michigan, Michigan, Lake Huron, H ...

which moderate the temperature of the surrounding land.

Status

The high

fertility Fertility in colloquial terms refers the ability to have offspring. In demographic contexts, fertility refers to the actual production of offspring, rather than the physical capability to reproduce, which is termed fecundity. The fertility rate ...

of the land has seen the region become highly developed and populated, with agricultural, industrial, commercial and urban areas. Today, the Carolinian Zone contains major cities and is home to one quarter of Canada's population despite being 0.25% of the total land area. The deforestation of the region for this development has led to significant

habitat loss Habitat destruction (also termed habitat loss or habitat reduction) occurs when a natural habitat is no longer able to support its native species. The organisms once living there have either moved elsewhere, or are dead, leading to a decrease ...

and fragmentation, leaving the remaining portions of land scattered and disconnected, with some areas still threatened by human development. In total, it is estimated that forest cover has been reduced from 80% to 11.3% while wetlands reduced from 28.3% to 5.1%. In addition to habitat loss and fragmentation, the native Carolinian species are also being threatened by invasive species such as

garlic mustard ''Alliaria petiolata'', or garlic mustard, is a biennial flowering plant in the mustard family (Brassicaceae). It is native to Europe, western and central Asia, north-western Africa, Morocco, Iberia and the British Isles, north to northern S ...

and buckthorn, and

overgrazing Overgrazing occurs when plants are exposed to intensive grazing for extended periods of time, or without sufficient recovery periods. It can be caused by either livestock in poorly managed agricultural applications, game reserves, or nature ...

white-tailed deer The white-tailed deer (''Odocoileus virginianus''), also known Common name, commonly as the whitetail and the Virginia deer, is a medium-sized species of deer native to North America, North, Central America, Central and South America. It is the ...

. These factors have contributed to the Carolinian zone becoming the most threatened region in Ontario, with over 125 species of plants and animals listed as either vulnerable, threatened or endangered by the federal or provincial governments. This is over one-third of all vulnerable,

threatened A threatened species is any species (including animals, plants and fungi) which is vulnerable to extinction in the near future. Species that are threatened are sometimes characterised by the population dynamics measure of ''critical depensatio ...

endangered An endangered species is a species that is very likely to become extinct in the near future, either worldwide or in a particular political jurisdiction. Endangered species may be at risk due to factors such as habitat loss, poaching, inv ...

species in Canada.

Conservation efforts

Some parts of the remaining natural area in the Carolinian zone are protected in an effort to conserve the region and its unique, diverse biota. For example, today there are many protected areas including Point Pelee National Park, 21 provincial parks, and many conservation areas. Some of the best preserved areas of Canada's Carolinian forest are located in Windsor's Ojibway Prairie Complex and

Rondeau Provincial Park Rondeau Provincial Park is the second oldest provincial park in Ontario, Canada, having been established with an order in council on 8 September 1894. The park is located in Southwestern Ontario, on an 8 km long crescentic sand spit exten ...

near

Morpeth, Ontario Morpeth is a community in Chatham-Kent, Ontario, Canada. It is located between Blenheim and St. Thomas on Highway 3. History The community is named after Lord Morpeth, who was once a guest of Col. Thomas Talbot. Morpeth was an area of notab ...

; another is the Niagara Glen Nature Reserve near

Niagara Falls, Ontario Niagara Falls is a city in Ontario, Canada, adjacent to, and named after, Niagara Falls. As of the Canada 2021 Census, 2021 census, the city had a population of 94,415. The city is located on the Niagara Peninsula along the western bank of the ...

. There are also organizations like th
Carolinian Canada Coalition
who aim to restore the region as much as possible. Carolinian forests are complex ecosystems that cannot be replicated in gardens or other small areas. The only way to preserve this unique habitat is continue to keep larger areas of land protected from development and agriculture. Other conservation efforts have included the reduction of the white-tailed deer population from their peak density of 55 deer km⁻² to 7 deer km^−2, between the years of 1996-2009. However, studies have found that while this helps reduce further forest damage, continued decline in the forest canopy can still occur, indicating the recovery from overgrazing by herbivores on the forest canopy is a long process without immediate results. Maintaining a lower density of white-tailed deer, while increasing the seed sources of native trees and protecting tree saplings in herbivore exclosures are all suggested ways of helping the Carolinian forest recover or at least conserve what is remaining.

Examples of species

Fauna: *White-tailed deer *

American black bear The American black bear (''Ursus americanus''), or simply black bear, is a species of medium-sized bear which is Endemism, endemic to North America. It is the continent's smallest and most widely distributed bear species. It is an omnivore, with ...

*Virginia opossum * American badger *

Gray fox The gray fox (''Urocyon cinereoargenteus''), or grey fox, is an omnivorous mammal of the family Canidae, widespread throughout North America and Central America. This species and its only congener (biology), congener, the diminutive island fox ...

Bobcat The bobcat (''Lynx rufus''), also known as the wildcat, bay lynx, or red lynx, is one of the four extant species within the medium-sized wild cat genus '' Lynx''. Native to North America, it ranges from southern Canada through most of the c ...

*Southern flying squirrel *

Wild turkey The wild turkey (''Meleagris gallopavo'') is an upland game bird native to North America, one of two extant species of Turkey (bird), turkey and the heaviest member of the order Galliformes. It is the ancestor to the domestic turkey (''M. g. dom ...

Ruffed grouse The ruffed grouse (''Bonasa umbellus'') is a medium-sized grouse occurring in forests from the Appalachian Mountains across Canada to Alaska. It is the most widely distributed game bird in North America. It is non-migratory. It is the only spe ...

Hooded warbler The hooded warbler (''Setophaga citrina'') is a New World warbler. It breeds in eastern North America across the eastern United States and into southernmost Canada (Ontario). It is migratory, wintering in Central America and the West Indies. Ho ...

Prothonotary warbler The prothonotary warbler (''Protonotaria citrea'') is a small songbird of the New World warbler family. It is named for its plumage, which resembles the yellow robes once worn by papal clerks (named prothonotaries) in the Roman Catholic Church. ...

*Canada warbler *

Carolina wren The Carolina wren (''Thryothorus ludovicianus'') is a common species of wren that is a resident in the Eastern United States, the extreme south of Ontario, Canada, and the extreme northeast of Mexico. Severe winters restrict the northern limits ...

Yellow-breasted chat The yellow-breasted chat (''Icteria virens'') is a large songbird found in America, and is the only member of the family Icteriidae. It was once a member of the New World warbler family Parulidae, but in 2017, the American Ornithological Socie ...

Northern cardinal The northern cardinal (''Cardinalis cardinalis''), also commonly known as the common cardinal, red cardinal, or simply cardinal, is a bird in the genus ''Cardinalis''. It can be found in southeastern Canada, through the eastern United States fro ...

* Red bellied woodpecker *

Red-tailed hawk The red-tailed hawk (''Buteo jamaicensis'') is a bird of prey that breeds throughout most of North America, from the interior of Alaska and northern Canada to as far south as Panama and the West Indies. It is one of the most common members of ...

American barn owl The American barn owl (''Tyto furcata'') is usually considered a subspecies group and together with the western barn owl group, the eastern barn owl group, and sometimes the Andaman masked owl, make up the Tyto, barn owl, cosmopolitan in range. ...

Passenger pigeon The passenger pigeon or wild pigeon (''Ectopistes migratorius'') is an bird extinction, extinct species of Columbidae, pigeon that was endemic to North America. Its common name is derived from the French word ''passager'', meaning "passing by" ...

( formerly) *

Carolina parakeet The Carolina parakeet (''Conuropsis carolinensis''), or Carolina conure, is an extinct species of small green neotropical parrot with a bright yellow head, reddish orange face, and pale beak that was native to the Eastern, Midwest, and Plains ...

(formerly) *

Eastern box turtle The eastern box turtle (''Terrapene carolina carolina'') is a subspecies within a group of hinge-shelled turtles normally called box turtles. ''T. c. carolina'' is native to the Eastern United States. The eastern box turtle is a subspec ...

Spotted turtle The spotted turtle (''Clemmys guttata''), the only species of the genus ''Clemmys'', is a small, semi-aquatic turtle that reaches a carapace length of upon adulthood. Their broad, smooth, low dark-colored upper shell, or carapace, ranges in its ...

* Eastern hognose snake *Eastern spiny softshell turtle * Eastern fox snake * Red salamander * Karner blue butterfly Flora: *

Eastern prickly pear ''Opuntia humifusa'', commonly known as the devil's-tongue, eastern prickly pear or Indian fig, is a cactus of the genus '' Opuntia'' present in parts of the eastern United States and northeastern Mexico. Description As is the case in other ' ...

cactus *Tulip tree *''

Sassafras ''Sassafras'' is a genus of three extant and one extinct species of deciduous trees in the family Lauraceae, native to eastern North America and eastern Asia.Wolfe, Jack A. & Wehr, Wesley C. 1987. The sassafras is an ornamental tree. "Middle Eo ...

'' * Miami mist *Pawpaw *

Kentucky coffeetree The Kentucky coffeetree (''Gymnocladus dioicus''), also known as American coffee berry, Kentucky mahogany, nicker tree, and stump tree, is a tree in the subfamily Caesalpinioideae of the legume family Fabaceae, native to the Midwest, Upper South, ...

Flowering dogwood ''Cornus florida'', the flowering dogwood, is a species of flowering tree in the family Cornaceae native to eastern North America and northern Mexico. An endemic population once spanned from southernmost coastal Maine south to northern Florida ...

Black walnut ''Juglans nigra'', the eastern American black walnut, is a species of deciduous tree in the walnut family, Juglandaceae, native to central and eastern North America, growing mostly in riparian zones. Black walnut is susceptible to thousand can ...

American chestnut The American chestnut (''Castanea dentata'') is a large, fast-growing deciduous tree of the Fagaceae, beech family native to eastern North America. As is true of all species in the genus ''Chestnut, Castanea'', the American chestnut produces Bur ...

American sycamore ''Platanus occidentalis'', also known as American sycamore, American planetree, western plane, occidental plane, buttonwood, and water beech, is a species of ''Platanus'' native to the eastern and central United States, the mountains of northeast ...

Red mulberry ''Morus rubra'', commonly known as the red mulberry, is a species of mulberry native to eastern and central North America. It is found from Ontario, Minnesota, and Vermont south to southern Florida, and west as far as southeastern South Dakota, ...

Black gum Black gum may refer to several species of plants: * ''Nyssa sylvatica ''Nyssa sylvatica'', commonly known as tupelo, black tupelo, black gum or sour gum, is a medium-sized deciduous tree native to eastern North America from the coastal Northeas ...

*Red maple *

Eastern white pine ''Pinus strobus'', commonly called the eastern white pine, northern white pine, white pine, Weymouth pine (British), and soft pine is a large pine native to eastern North America. It occurs from Newfoundland, Canada, west through the Great Lake ...

Eastern hemlock ''Tsuga canadensis'', also known as eastern hemlock, eastern hemlock-spruce, or Canadian hemlock, and in the French-speaking regions of Canada as ''pruche du Canada'', is a coniferous tree native to eastern North America. It is the state tree of ...

Honey locust The honey locust (''Gleditsia triacanthos''), also known as the thorny locust or thorny honeylocust, is a deciduous tree in the family Fabaceae, native to central North America where it is mostly found in the moist soil of river valleys. Honey ...

Rivers and creeks

Ausable River Au Sable, Ausable or Aux Sable may refer to a documentary film or various places: * Au Sable (documentary), a film about the Michigan river, canoe race and people Illinois * Aux Sable Township, Grundy County, Illinois Michigan *Au Sable Town ...

* Catfish Creek *

Credit River The Credit River is a river in southern Ontario, which flows from headwaters above the Niagara Escarpment near Orangeville and Caledon East to empty into Lake Ontario at Port Credit, Mississauga. It drains an area of approximately . The total l ...

* Kettle Creek * Grand River * Rouge River *

Sydenham River The Sydenham River is a river in Chatham-Kent, Lambton County and Middlesex County in southwestern Ontario, Canada, flowing southwest from its source west of London, Ontario and emptying into Lake Saint Clair. The length of the river is and it ...

Thames River The River Thames ( ), known alternatively in parts as the River Isis, is a river that flows through southern England including London. At , it is the longest river entirely in England and the second-longest in the United Kingdom, after the ...

* Twenty Mile Creek *

Welland River The Welland River (originally called the Chippawa Creek) is a river in the Golden Horseshoe that passes through the Southern Ontario cities of Welland and Niagara Falls. It flows from its source, a Drainage Basin just south of Hamilton, On ...

* Nith River

Other

Rouge Park Rouge National Urban Park is a national urban park in Ontario, Canada. The park is centred around the Rouge River and its tributaries in the Greater Toronto Area. The southern portion of the park is situated around the mouth of the river in ...

* Barker's Bush *

Point Pelee National Park Point Pelee National Park (; ) is a National parks of Canada, national park in Essex County, Ontario, Essex County in southwestern Ontario, Canada where it extends into Lake Erie. The word is French for 'bald'. Point Pelee consists of a peninsu ...

References

{{Reflist

External links

Carolinian CanadaCarolinian invasive species
Temperate broadleaf and mixed forests in the United States Ecoregions of the United States Ecoregions of Canada Forests of Ontario Forests of the United States

Flora of the Eastern United States Flora (: floras or florae) is all the plant life present in a particular region or time, generally the naturally occurring ( indigenous) native plants. The corresponding term for animals is ''fauna'', and for fungi, it is '' funga''. Sometimes ...

Temperate broadleaf and mixed forests in Canada