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The capybara or greater capybara (''Hydrochoerus hydrochaeris'') is the largest living rodent, native to
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a considerably smaller portion in the Northern Hemisphere. It can also be described as the southern Subregion#Americas, subregion o ...
. It is a member of the genus '' Hydrochoerus''. The only other
extant Extant or Least-concern species, least concern is the opposite of the word extinct. It may refer to: * Extant hereditary titles * Extant literature, surviving literature, such as ''Beowulf'', the oldest extant manuscript written in English * Exta ...
member is the lesser capybara (''Hydrochoerus isthmius''). Its close relatives include
guinea pig The guinea pig or domestic guinea pig (''Cavia porcellus''), also known as the cavy or domestic cavy ( ), is a species of rodent belonging to the genus ''Cavia'', family Caviidae. Animal fancy, Breeders tend to use the name "cavy" for the ani ...
s and rock cavies, and it is more distantly related to the agouti, the chinchilla, and the
nutria The nutria () or coypu () (''Myocastor coypus'') is a herbivore, herbivorous, semiaquatic rodent from South America. Classified for a long time as the only member of the family Myocastoridae, ''Myocastor'' has since been included within Echimy ...
. The capybara inhabits savannas and dense forests, and lives near bodies of water. It is a highly social species and can be found in groups as large as one hundred individuals, but usually live in groups of 10–20 individuals. The capybara is hunted for its meat and hide and also for grease from its thick fatty skin.


Etymology

Its common name is derived from Tupi , a complex agglutination of (leaf) + (slender) + (eat) + (a suffix for agent nouns), meaning "one who eats slender leaves", or "grass-eater". The genus name, ''hydrochoerus'', comes from Greek (' "water") and (' "pig, hog") and the species name, ''hydrochaeris'', comes from Greek (' "water") and (' "feel happy, enjoy"). * In page 57, Darwin says "The largest gnawing animal in the world, the ''Hydrochærus Capybara'' (the water-hog), is here also common." * See it also i
The Complete Work of Charles Darwin Online
/ref>


Classification and phylogeny

The capybara and the lesser capybara both belong to the subfamily Hydrochoerinae along with the rock cavies. The living capybaras and their extinct relatives were previously classified in their own family Hydrochoeridae. Since 2002, molecular phylogenetic studies have recognized a close relationship between ''Hydrochoerus'' and '' Kerodon'', the rock cavies, supporting placement of both genera in a subfamily of Caviidae. Paleontological classifications previously used Hydrochoeridae for all capybaras, while using Hydrochoerinae for the living genus and its closest fossil relatives, such as '' Neochoerus'', but more recently have adopted the classification of Hydrochoerinae within Caviidae. The taxonomy of fossil hydrochoerines is also in a state of flux. In recent years, the diversity of fossil hydrochoerines has been substantially reduced. This is largely due to the recognition that capybara molar teeth show strong variation in shape over the life of an individual. In one instance, material once referred to four genera and seven species on the basis of differences in molar shape is now thought to represent differently aged individuals of a single species, ''Cardiatherium paranense''. Among fossil species, the name "capybara" can refer to the many species of Hydrochoerinae that are more closely related to the modern ''Hydrochoerus'' than to the "cardiomyine" rodents like ''Cardiomys''. The fossil genera ''Cardiatherium'', ''Phugatherium'', ''Hydrochoeropsis'', and ''Neochoerus'' are all capybaras under that concept.


Description

The capybara has a heavy, barrel-shaped body and short head, with reddish-brown fur on the upper part of its body that turns yellowish-brown underneath. Its sweat glands can be found in the surface of the hairy portions of its skin, an unusual trait among rodents. The animal lacks
down hair A fur is a soft, thick growth of hair that covers the skin of almost all mammals. It consists of a combination of oily guard hair on top and thick underfur beneath. The guard hair keeps moisture from reaching the skin; the underfur acts as an ...
, and its guard hair differs little from over hair. Adult capybaras grow to in length, stand tall at the
withers Withers are the ridge between the shoulder blades of an animal, typically a quadruped. In many species, this ridge is the tallest point of the body. In horses and dogs, it is the standard place to measure the animal's height. In contrast, catt ...
, and typically weigh , with an average in the Venezuelan llanos of .Capybara
, Arkive
Females are slightly heavier than males. The top recorded weights are for a wild female from Brazil and for a wild male from Uruguay. Also, an 81 kg individual was reported in São Paulo in 2001 or 2002. The dental formula is . Capybaras have slightly webbed feet and vestigial tails. Their hind legs are slightly longer than their forelegs; they have three toes on their rear feet and four toes on their front feet. Their muzzles are blunt, with nostrils, and the eyes and ears are near the top of their heads. Its
karyotype A karyotype is the general appearance of the complete set of chromosomes in the cells of a species or in an individual organism, mainly including their sizes, numbers, and shapes. Karyotyping is the process by which a karyotype is discerned by de ...
has 2n = 66 and FN = 102, meaning it has 66 chromosomes with a total of 102 arms.


Ecology

Capybaras are semiaquatic mammals found throughout all countries of South America except
Chile Chile, officially the Republic of Chile, is a country in western South America. It is the southernmost country in the world and the closest to Antarctica, stretching along a narrow strip of land between the Andes, Andes Mountains and the Paci ...
. They live in densely forested areas near bodies of water, such as lakes, rivers, swamps, ponds, and marshes, as well as flooded savannah and along rivers in the tropical rainforest. They are superb swimmers and can hold their breath underwater for up to five minutes at a time. Capybara have flourished in cattle ranches. They roam in home ranges averaging in high-density populations. Many escapees from captivity can also be found in similar watery habitats around the world. Sightings are fairly common in
Florida Florida ( ; ) is a U.S. state, state in the Southeastern United States, Southeastern region of the United States. It borders the Gulf of Mexico to the west, Alabama to the northwest, Georgia (U.S. state), Georgia to the north, the Atlantic ...
, although a breeding population has not yet been confirmed. In 2011, one specimen was spotted on the Central Coast of California. These escaped populations occur in areas where prehistoric capybaras inhabited;
late Pleistocene The Late Pleistocene is an unofficial Age (geology), age in the international geologic timescale in chronostratigraphy, also known as the Upper Pleistocene from a Stratigraphy, stratigraphic perspective. It is intended to be the fourth division ...
capybaras inhabited Florida and '' Hydrochoerus hesperotiganites'' in
California California () is a U.S. state, state in the Western United States that lies on the West Coast of the United States, Pacific Coast. It borders Oregon to the north, Nevada and Arizona to the east, and shares Mexico–United States border, an ...
and '' Hydrochoerus gaylordi'' in
Grenada Grenada is an island country of the West Indies in the eastern Caribbean Sea. The southernmost of the Windward Islands, Grenada is directly south of Saint Vincent and the Grenadines and about north of Trinidad and Tobago, Trinidad and the So ...
, and feral capybaras in North America may actually fill the ecological niche of the Pleistocene species.


Diet and predation

Capybaras are
herbivore A herbivore is an animal anatomically and physiologically evolved to feed on plants, especially upon vascular tissues such as foliage, fruits or seeds, as the main component of its diet. These more broadly also encompass animals that eat ...
s, grazing mainly on grasses and aquatic plants, as well as fruit and tree bark. They are very selective feeders and feed on the leaves of one species and disregard other species surrounding it. They eat a greater variety of plants during the dry season, as fewer plants are available. While they eat grass during the wet season, they have to switch to more abundant reeds during the dry season. Plants that capybaras eat during the summer lose their nutritional value in the winter, so they are not consumed at that time. The capybara's jaw hinge is not perpendicular, so they chew food by grinding back-and-forth rather than side-to-side. Capybaras are autocoprophagous, meaning they eat their own feces as a source of bacterial gut flora, to help digest the
cellulose Cellulose is an organic compound with the chemical formula, formula , a polysaccharide consisting of a linear chain of several hundred to many thousands of glycosidic bond, β(1→4) linked glucose, D-glucose units. Cellulose is an important s ...
in the grass that forms their normal diet, and to extract the maximum protein and vitamins from their food. They also regurgitate food to masticate again, similar to cud-chewing by cattle. Like other rodents, a capybara's front teeth grow continually to compensate for the constant wear from eating grasses; their cheek teeth also grow continuously. Like its relative the guinea pig, the capybara does not have the capacity to synthesize
vitamin C Vitamin C (also known as ascorbic acid and ascorbate) is a water-soluble vitamin found in citrus and other fruits, berries and vegetables. It is also a generic prescription medication and in some countries is sold as a non-prescription di ...
, and capybaras not supplemented with vitamin C in captivity have been reported to develop gum disease as a sign of
scurvy Scurvy is a deficiency disease (state of malnutrition) resulting from a lack of vitamin C (ascorbic acid). Early symptoms of deficiency include weakness, fatigue, and sore arms and legs. Without treatment, anemia, decreased red blood cells, gum d ...
. The maximum lifespan of the capybara is 8 to 10 years, but in the wild capybaras usually do not live longer than four years because of predation from South American big cats such as jaguars and
cougar The cougar (''Puma concolor'') (, ''Help:Pronunciation respelling key, KOO-gər''), also called puma, mountain lion, catamount and panther is a large small cat native to the Americas. It inhabits North America, North, Central America, Cent ...
s and from non-mammalian predators such as
harpy eagle The harpy eagle (''Harpia harpyja'') is a large Neotropical realm, neotropical species of eagle. It is also called the American harpy eagle to distinguish it from the Papuan eagle, which is sometimes known as the New Guinea Harpy Eagle, New Guin ...
s, caimans, green anacondas and piranhas.


Social organization

Capybaras are known to be gregarious. While they sometimes live solitarily, they are more commonly found in groups of around 10–20 individuals, with two to four adult males, four to seven adult females, and the remainder juveniles. Capybara groups can consist of as many as 50 or 100 individuals during the dry season when the animals gather around available water sources. Males establish social bonds, dominance, or general group consensus. They can make dog-like barks when threatened or when females are herding young. Capybaras have two types of scent glands: a morrillo, located on the snout, and anal glands. Both sexes have these glands, but males have much larger morrillos and use their anal glands more frequently. The anal glands of males are also lined with detachable hairs. A crystalline form of scent secretion is coated on these hairs and is released when in contact with objects such as plants. These hairs have a longer-lasting scent mark and are tasted by other capybaras. Capybaras scent-mark by rubbing their morrillos on objects, or by walking over scrub and marking it with their anal glands. Capybaras can spread their scent farther by urinating; however, females usually mark without urinating and scent-mark less frequently than males overall. Females mark more often during the wet season when they are in estrus. In addition to objects, males also scent-mark females.


Reproduction

When in estrus, the female's scent changes subtly and nearby males begin pursuit. In addition, a female alerts males she is in estrus by whistling through her nose. During mating, the female has the advantage and mating choice. Capybaras mate only in water, and if a female does not want to mate with a certain male, she either submerges or leaves the water. Dominant males are highly protective of the females, but they usually cannot prevent some of the subordinates from copulating. The larger the group, the harder it is for the male to watch all the females. Dominant males secure significantly more matings than each subordinate, but subordinate males, as a class, are responsible for more matings than each dominant male. The lifespan of the capybara's sperm is longer than that of other rodents. Capybara
gestation Gestation is the period of development during the carrying of an embryo, and later fetus, inside viviparous animals (the embryo develops within the parent). It is typical for mammals, but also occurs for some non-mammals. Mammals during pregn ...
is 130–150 days, and produces a litter of four young on average, but may produce between one and eight in a single litter. Birth is on land and the female rejoins the group within a few hours of delivering the newborn capybaras, which join the group as soon as they are mobile. Within a week, the young can eat grass, but continue to suckle—from any female in the group—until weaned around 16 weeks. The young form a group within the main group. Alloparenting has been observed in this species. Breeding peaks between April and May in Venezuela and between October and November in
Mato Grosso Mato Grosso ( – ) is one of the states of Brazil, the List of Brazilian states by area, third largest by area, located in the Central-West Region, Brazil, Central-West region. The state has 1.66% of the Brazilian population and is responsible ...
, Brazil.


Activities

Though quite agile on land, capybaras are equally at home in the water. They are excellent swimmers, and can remain completely submerged for up to five minutes, an ability they use to evade predators. Capybaras can sleep in water, keeping only their noses out. As temperatures increase during the day, they wallow in water and then graze during the late afternoon and early evening. They also spend time wallowing in mud. They rest around midnight and then continue to graze before dawn.


Communication

Capybaras communicate using barks, chirps, whistles, huffs, and purrs.


Conservation and human interaction

Capybaras are not considered a threatened species; their population is stable throughout most of their South American range, though in some areas hunting has reduced their numbers. Capybaras are hunted for their meat and pelts in some areas, and otherwise killed by humans who see their grazing as competition for livestock. In some areas, they are farmed, which has the effect of ensuring the wetland habitats are protected. Their survival is aided by their ability to breed rapidly. Capybaras have adapted well to urbanization in South America. They can be found in many areas in zoos and parks, and may live for 12 years in captivity, more than double their wild lifespan. Capybaras are docile and usually allow humans to pet and hand-feed them, but physical contact is normally discouraged, as their ticks can be vectors to Rocky Mountain spotted fever. The
European Association of Zoos and Aquaria The European Association of Zoos and Aquaria (EAZA) is an organisation for the European zoo and aquarium community that links over 340 member organisations in 41 countries. EAZA membership is open to all zoos and aquaria across Europe that compl ...
asked Drusillas Park in Alfriston,
Sussex Sussex (Help:IPA/English, /ˈsʌsɪks/; from the Old English ''Sūþseaxe''; lit. 'South Saxons'; 'Sussex') is an area within South East England that was historically a kingdom of Sussex, kingdom and, later, a Historic counties of England, ...
, England, to keep the studbook for capybaras, to monitor captive populations in Europe. The studbook includes information about all births, deaths and movements of capybaras, as well as how they are related. Capybaras are farmed for meat and skins in South America. The meat is considered unsuitable to eat in some areas, while in other areas it is considered an important source of protein. In parts of South America, especially in Venezuela, capybara meat is popular during
Lent Lent (, 'Fortieth') is the solemn Christianity, Christian religious moveable feast#Lent, observance in the liturgical year in preparation for Easter. It echoes the 40 days Jesus spent fasting in the desert and enduring Temptation of Christ, t ...
and
Holy Week Holy Week () commemorates the seven days leading up to Easter. It begins with the commemoration of Triumphal entry into Jerusalem, Christ's triumphal entry into Jerusalem on Palm Sunday, marks the betrayal of Jesus on Spy Wednesday (Holy Wednes ...
as the Catholic Church (according to a legend) previously issued special dispensation to allow it to be eaten while other meats are generally forbidden. There is widespread perception in Venezuela that consumption of capybaras is exclusive to rural people. In August 2021, Argentine and international media reported that capybaras had been disturbing residents of Nordelta, an affluent gated community north of
Buenos Aires Buenos Aires, controlled by the government of the Autonomous City of Buenos Aires, is the Capital city, capital and largest city of Argentina. It is located on the southwest of the Río de la Plata. Buenos Aires is classified as an Alpha− glob ...
built atop the local capybara's preexisting wetland habitat. This inspired social media users to jokingly adopt the capybara as a symbol of class struggle and
communism Communism () is a political sociology, sociopolitical, political philosophy, philosophical, and economic ideology, economic ideology within the history of socialism, socialist movement, whose goal is the creation of a communist society, a ...
. Brazilian Lyme-like borreliosis likely involves capybaras as reservoirs and '' Amblyomma'' and '' Rhipicephalus'' ticks as vectors. A Capybara café in St. Augustine,
Florida Florida ( ; ) is a U.S. state, state in the Southeastern United States, Southeastern region of the United States. It borders the Gulf of Mexico to the west, Alabama to the northwest, Georgia (U.S. state), Georgia to the north, the Atlantic ...
allows visitors to interact with and give head scratches to the rodents.


In popular culture

Izu Shaboten Zoo and other zoos in Japan have prepared hot spring baths for capybaras. Video clips of the bathing capybaras have gained millions of views. The capybaras have influenced an anime character named ''Kapibara-san'', and a series of merchandise such as plush toys. Capybaras have long been a figure in meme culture, particularly in the 2020s. In 2022, Peronists in Argentina presented them as figures of class struggle after the disturbances in Nordelta. Common meme formats pair capybaras with the song " After Party" by Don Toliver.


See also

* '' Josephoartigasia monesi'', an extinct species identified as the largest known rodent ever * Kurloff cell, a type of cell found in capybaras and guinea pigs * '' Capybara Walking'', a historical animal locomotion film by
Eadweard Muybridge Eadweard Muybridge ( ; 9 April 1830 – 8 May 1904, born Edward James Muggeridge) was an English photographer known for his pioneering work in photographic studies of motion, and early work in motion-picture Movie projector, projection. He ...


Notes


References


External links

*
Animal Diversity Web ''Hydrochoerus hydrochaeris''

Capybara information
{{Taxonbar, from=Q131538 Mammals described in 1766 Cavies Herbivorous mammals Mammals of Argentina Mammals of Bolivia Rodents of Brazil Mammals of Colombia Mammals of Ecuador Mammals of French Guiana Mammals of Guyana Mammals of Paraguay Mammals of Peru Mammals of Uruguay Mammals of Venezuela Articles containing video clips Semiaquatic mammals Taxa named by Carl Linnaeus 2020s in Internet culture Internet memes introduced in the 2020s