''Canaanimico'' is an extinct

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

of medium-sized

New World monkey New World monkeys are the five families of primates that are found in the tropical regions of Mexico, Central and South America: Callitrichidae, Cebidae, Aotidae, Pitheciidae, and Atelidae. The five families are ranked together as the Ceboi ...

s from the

Late Oligocene The Chattian is, in the geologic timescale The geologic time scale or geological time scale (GTS) is a representation of time based on the rock record of Earth. It is a system of chronological dating that uses chronostratigraphy (the pro ...

(approximately 26.5 Ma,

Deseadan The Deseadan () age is a period of geologic time (29.0–21.0 Ma) within the Oligocene epoch of the Paleogene to the Early Miocene epoch of the Neogene, used more specifically within the SALMA classification of South America. It follows the Tingu ...

in the SALMA classification) fossiliferous fluvio-

lacustrine A lake is often a naturally occurring, relatively large and fixed body of water on or near the Earth's surface. It is localized in a basin or interconnected basins surrounded by dry land. Lakes lie completely on land and are separate from t ...

Chambira Formation The Chambira River is a major tributary of the Marañón River, and has been the traditional territory of the Urarina peoples for at least the past 350 years. Located in the Amazon jungle of Peru, otherwise known as the Selva, the Chambira is a tr ...

of the

Ucayali Basin The Ucayali River (, ) is the main headstream of the Amazon River. It rises about north of Lake Titicaca, in the Arequipa region of Peru and becomes the Amazon at the confluence of the Marañón close to Nauta city. The city of Pucallpa is lo ...

in Amazonian

Peru Peru, officially the Republic of Peru, is a country in western South America. It is bordered in the north by Ecuador and Colombia, in the east by Brazil, in the southeast by Bolivia, in the south by Chile, and in the south and west by the Pac ...

. The genus was described by Marivaux et al. in 2016 and the

type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...

is ''C. amazonensis''.Marivaux et al., 2016''Canaanimico''
at

Fossilworks Fossilworks was a portal which provides query, download, and analysis tools to facilitate access to the Paleobiology Database, a large relational database assembled by hundreds of paleontologists from around the world. History Fossilworks was cr ...

.org

Classification

Marivaux et al. placed the genus in the

subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end botanical subfamily names with "-oideae", and zo ...

Soriacebinae, of the family Homunculidae,Homunculidae
at

.org while Silvestro et al. (2017) attribute an ''

incertae sedis or is a term used for a taxonomy (biology), taxonomic group where its broader relationships are unknown or undefined. Alternatively, such groups are frequently referred to as "enigmatic taxa". In the system of open nomenclature, uncertainty ...

'' status for the family and subfamily to the genus.Silvestro et al., 2017, p.13 Marivaux et al. proposed a close relation with the

Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...

South American genera '' Soriacebus'' (

Santacrucian The Santacrucian age is a period of geologic time (17.5 – 16.3 Mya (unit), Ma) within the Early Miocene epoch of the Neogene, used more specifically with South American land mammal age, SALMA classification in South America. It follows the Colhue ...

) and '' Mazzonicebus'' (

Colhuehuapian The Colhuehuapian age is a period of geologic time (21.0–17.5 Ma) within the Early Miocene epoch of the Neogene, used more specifically within the SALMA classification in South America. It follows the Deseadan and precedes the Santacrucian ag ...

).Urbani et al., 2018, p.33

Description

''Canaanimico'' was described by Marivaux et al. on the basis of two isolated upper

molars The molars or molar teeth are large, flat tooth, teeth at the back of the mouth. They are more developed in mammal, mammals. They are used primarily to comminution, grind food during mastication, chewing. The name ''molar'' derives from Latin, '' ...

collected at the Contamana fossil locality. The authors inferred that based on

dental microwear Dental microwear analysis is a method to infer diet and behavior in extinct animals, especially in fossil specimens. It has been used on a variety of taxa, including hominids, victoriapithecids, amphicyonids, canids, ursids, hyaenids, hyaenodont ...

patterns recorded on one upper molar ''Canaanimico'' possibly was a fruit and hard-object eater. Silvestro et al. estimated a body mass of for ''Canaanimico'', making it a medium-sized New World monkey.Silvestro et al., 2017, p.14

Paleoecology

Fossils of ''Canaanimico'' were found in the

(

)

of the

in Amazonian Peru. The formation, a sequence of red

claystone Mudrocks are a class of fine-grained siliciclastic sedimentary rocks. The varying types of mudrocks include siltstone, claystone, mudstone and shale. Most of the particles of which the stone is composed are less than and are too small to ...

s and

paleosol In Earth science, geoscience, paleosol (''palaeosol'' in Great Britain and Australia) is an ancient soil that formed in the past. The definition of the term in geology and paleontology is slightly different from its use in soil science. In geo ...

s with intercalated conglomerates and

sandstone Sandstone is a Clastic rock#Sedimentary clastic rocks, clastic sedimentary rock composed mainly of grain size, sand-sized (0.0625 to 2 mm) silicate mineral, silicate grains, Cementation (geology), cemented together by another mineral. Sand ...

s,Antoine et al., 2016, p.40 and

gypsum Gypsum is a soft sulfate mineral composed of calcium sulfate Hydrate, dihydrate, with the chemical formula . It is widely mined and is used as a fertilizer and as the main constituent in many forms of plaster, drywall and blackboard or sidewalk ...

layers,Castro Medina, 2010, p.55 has been dated on the basis of

zircon Zircon () is a mineral belonging to the group of nesosilicates and is a source of the metal zirconium. Its chemical name is zirconium(IV) silicate, and its corresponding chemical formula is Zr SiO4. An empirical formula showing some of th ...

s in a

tuff Tuff is a type of rock made of volcanic ash ejected from a vent during a volcanic eruption. Following ejection and deposition, the ash is lithified into a solid rock. Rock that contains greater than 75% ash is considered tuff, while rock co ...

bed in the formation, providing an age of 26.56 ± 0.07 Ma.Antoine et al., 2016, p.35 The formation was deposited in a

tectonically Tectonics ( via Latin ) are the processes that result in the structure and properties of the Earth's crust and its evolution through time. The field of ''planetary tectonics'' extends the concept to other planets and moons. These processes ...

relatively calmCastro Medina, 2010, p.71 tropical fluvio-

environment with

oxbow lake An oxbow lake is a U-shaped lake or stream pool, pool that forms when a wide meander of a river is meander cutoff, cut off, creating a free-standing body of water. The word "oxbow" can also refer to a U-shaped bend in a river or stream, whether ...

s.Antoine et al., 2016, p.41Boivin et al., 2016, p.103 The Chambira Formation has provided a rich mammal assemblage of '' Chambiramys shipiborum'', '' C. sylvaticus'', '' Deseadomys cf. arambourgi'', '' Loretomys minutus'', '' Palaeosteiromys amazonensis'', '' Plesiosteiromys newelli'', '' Scleromys praecursor'', '' Ucayalimys crassidens'', '' Abderites sp.'', '' aff. Eosallamys sp.'', '' cf. Neoglyptatelus sp.'', and indeterminate Adelphomyinae,

Anthropoidea The simians, anthropoids, or higher primates are an infraorder (Simiiformes ) of primates containing all animals traditionally called monkeys and apes. More precisely, they consist of the parvorders Platyrrhini (New World monkeys) and Catar ...

, Astrapotheriidae,

Caenolestidae The family Caenolestidae contains the seven surviving species of shrew opossum: small, shrew-like marsupials that are confined to the Andes mountains of South America. The order is thought to have diverged from the ancestral marsupial line very e ...

Caviomorpha Caviomorpha is the rodent parvorder that unites all New World hystricognaths. It is supported by both fossil and molecular evidence. The Caviomorpha was for a time considered to be a separate order outside the Rodentia, but is now accepted as ...

Chinchilloidea Caviomorpha is the rodent parvorder that unites all New World hystricognaths. It is supported by both fossil and molecular evidence. The Caviomorpha was for a time considered to be a separate order outside the Rodentia, but is now accepted as ...

Emballonuridae Emballonuridae is a family of microbats, many of which are referred to as sac-winged or sheath-tailed bats. They are widely distributed in tropical and subtropical regions around the world. The earliest fossil records are from the Eocene. Desc ...

, Erethizontoidea,

Herpetotheriidae Herpetotheriidae is an extinct family of metatherians, closely related to marsupials. Species of this family are generally reconstructed as terrestrial, and are considered morphologically similar to modern opossums. They are suggested to have b ...

, Interatheriinae,

Litopterna Litopterna (from "smooth heel") is an extinction, extinct order of South American native ungulates that lived from the Paleocene to the Pleistocene-Holocene around 62.5 million to 12,000 years ago (or possibly as late as 3,500 years ago), and we ...

Marsupialia Marsupials are a diverse group of mammals belonging to the infraclass Marsupialia. They are natively found in Australasia, Wallacea, and the Americas. One of marsupials' unique features is their reproductive strategy: the young are born in a ...

Microbiotheria Microbiotheria is an australidelphian marsupial order that encompasses two families, Microbiotheriidae and Woodburnodontidae, and is represented by only one extant species, the monito del monte, and a number of extinct species known from foss ...

Mylodontidae Mylodontidae is a family of extinct South American and North American ground sloths within the suborder Folivora of order Pilosa, living from around 23 million years ago (Mya) to 11,000 years ago. This family is most closely related to another fa ...

, Mylodontoidea,

Notoungulata Notoungulata is an extinct order of ungulates that inhabited South America from the early Paleocene to the end of the Pleistocene, living from approximately 61 million to 11,000 years ago. Notoungulates were morphologically diverse, with forms re ...

Octodontoidea Caviomorpha is the rodent parvorder that unites all New World hystricognaths. It is supported by both fossil and molecular phylogeny, molecular evidence. The Caviomorpha was for a time considered to be a separate order outside the Rodentia, but i ...

, Palaeothentidae,

Pampatheriidae Pampatheriidae ("Pampas beasts") is an extinct family of large cingulates related to armadillos. They first appeared in South America during the mid-Miocene, and '' Holmesina'' and '' Pampatherium'' spread to North America during the Pleistocen ...

Rhinolophoidea Rhinolophoidea is a superfamily (taxonomy), superfamily of bats. It contains the following families: Craseonycteridae, Hipposideridae, Megadermatidae, Rhinolophidae, Rhinonycteridae, and Rhinopomatidae. It is one of two superfamilies that compris ...

Tolypeutinae Tolypeutinae is a subfamily of armadillos in the family Chlamyphoridae, consisting of the giant, three-banded and naked-tailed armadillos. __TOC__ Taxonomy It contains the following genera: *'' Cabassous'' *'' Kuntinaru'' *'' Priodontes'' *' ...

Toxodontidae Toxodontidae is an extinct family of notoungulate mammals, known from the Oligocene to the Holocene (11,000 BP) of South America, with one genus, '' Mixotoxodon'', also known from the Pleistocene of Central America and southern North America (a ...

Typotheria Notoungulata is an extinct order of ungulates that inhabited South America from the early Paleocene to the end of the Pleistocene, living from approximately 61 million to 11,000 years ago. Notoungulates were morphologically diverse, with forms re ...

and

Vespertilionoidea Vespertilionoidea is a superfamily of bats containing five families: Cistugidae, Miniopteridae, Molossidae, Natalidae, and Vespertilionidae. It is one of three superfamilies in the suborder Yangochiroptera, the others being Noctilionoidea ...

.Boivin et al., 2016CTA-61
at

.orgCTA-32
at

.org Additionally,

crocodylia Crocodilia () is an Order (biology), order of semiaquatic, predatory reptiles that are known as crocodilians. They first appeared during the Late Cretaceous and are the closest living relatives of birds. Crocodilians are a type of crocodylomorp ...

ns and

turtle Turtles are reptiles of the order (biology), order Testudines, characterized by a special turtle shell, shell developed mainly from their ribs. Modern turtles are divided into two major groups, the Pleurodira (side necked turtles) and Crypt ...

s of '' ?Balanerodus sp.'', '' Podocnemis sp.'', '' cf. Purussaurus sp.'', '' cf. Sebecus sp.'', and indeterminate

frogs A frog is any member of a diverse and largely semiaquatic group of short-bodied, tailless amphibian vertebrates composing the order Anura (coming from the Ancient Greek , literally 'without tail'). Frog species with rough skin texture due to ...

Booidea The Booidea, also known as booid snakes, are a superfamily of snakes that contains boas (family Boidae) and other closely related boa-like snakes (but not pythons, which are in a separate superfamily called Pythonoidea). As of 2017, Booidea cont ...

Caimaninae A caiman ( (also spelled cayman) from Taíno ''kaiman'') is an alligatorid belonging to the subfamily Caimaninae, one of two primary lineages within the Alligatoridae family, the other being alligators. Caimans are native to Central and ...

Colubroidea Colubroidea is a superfamily of snakes in the clade Colubroides that includes Colubridae, with some studies splitting Colubridae into multiple families that make up Colubroidea. Historically, Colubroidea also included other caenophidian snakes su ...

Gavialoidea Gavialoidea is one of three superfamily (zoology), superfamilies of crocodylians, the other two being Alligatoroidea and Crocodyloidea. Although many extinct species are known, only the gharial ''Gavialis gangeticus'' and the false gharial ''Tomi ...

and

snakes Snakes are elongated Limbless vertebrate, limbless reptiles of the suborder Serpentes (). Cladistically Squamata, squamates, snakes are ectothermic, amniote vertebrates covered in overlapping Scale (zoology), scales much like other members of ...

were found in the formation, as well as fossil fishes of '' cf. Hydrolycus sp.'', '' Leporinus sp.'', '' cf. Phractocephalus sp.'', '' Potamotrygon sp.'', and indeterminate

Actinopterygii Actinopterygii (; ), members of which are known as ray-finned fish or actinopterygians, is a class (biology), class of Osteichthyes, bony fish that comprise over 50% of living vertebrate species. They are so called because of their lightly built ...

Cichlidae Cichlids () are a large, diverse, and widespread family of percomorph fish in the family Cichlidae, order Cichliformes. At least 1,760 species have been scientifically described, making it one of the largest vertebrate families, with on ...

Erythrinidae The Erythrinidae are a family of fishes found in rivers and other freshwater habitats from Costa Rica south as far as Argentina. They are common and are caught with hooks by fishermen, partially because of their voracious behaviour. They are som ...

Loricariidae Loricariidae is the largest family (biology), family of catfish (order Siluriformes), with over 90 genus, genera and just over 680 species. Loricariids originate from freshwater habitats of Costa Rica, Panama, and tropical and subtropical South A ...

and

Pimelodidae The Pimelodidae, commonly known as the long-whiskered catfishes, are a family of catfishes (order Siluriformes). Taxonomy The family Pimelodidae has undergone much revision. Currently, it contains about 30 genera and about 90 recognized and know ...

References

Bibliography

* * * * * *

Classification

Description

Paleoecology

See also

References

Bibliography

Further reading