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''Blechnum'', known as hard fern, is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
of
fern A fern (Polypodiopsida or Polypodiophyta ) is a member of a group of vascular plants (plants with xylem and phloem) that reproduce via spores and have neither seeds nor flowers. The polypodiophytes include all living pteridophytes exce ...
s in the family
Blechnaceae Blechnaceae is a family of ferns in the order Polypodiales, with a cosmopolitan distribution. Its status as a family and the number of genera included have both varied considerably. In the Pteridophyte Phylogeny Group classification of 201 ...
, subfamily Blechnoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). Two very different circumscriptions of the genus are used by different authors. In the PPG I system, based on Gasper et al. (2016), ''Blechnum'' is one of 18 genera in the subfamily Blechnoideae, and has about 30 species. Other sources use a very broadly defined ''Blechnum'' s.l., including accepting only two other genera in the subfamily. The genus then has about 250 species. In the PPG I circumscription, the genus is mostly
neotropical The Neotropical realm is one of the eight biogeographic realms constituting Earth's land surface. Physically, it includes the tropical terrestrial ecoregions of the Americas and the entire South American temperate zone. Definition In bioge ...
, with a few southern African species.


Description

Plants in the genus ''Blechnum'' (as circumscribed in the PPG I classification) are mainly terrestrial or grow on rocks; few are
epiphyte An epiphyte is an organism that grows on the surface of a plant and derives its moisture and nutrients from the air, rain, water (in marine environments) or from debris accumulating around it. The plants on which epiphytes grow are called phoroph ...
s. Their
rhizome In botany and dendrology, a rhizome (; , ) is a modified subterranean plant stem that sends out roots and shoots from its nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from axillary buds and grow ...
s may be upright or creeping and have scales with entire margins or at most a few very small teeth. They generally form
stolon In biology, stolons (from Latin '' stolō'', genitive ''stolōnis'' – "branch"), also known as runners, are horizontal connections between organisms. They may be part of the organism, or of its skeleton; typically, animal stolons are external s ...
s, which is a characteristic of the genus. The sterile and fertile fronds are usually of the same form or at most slightly different. The blades of the fronds are of one colour and are usually pinnatisect or unipinnate, rarely entire. The leaf veins are usually free, dividing one to three times, each ending near the frond margin in an enlarged tip. The sori have indusia (scale-like coverings).


Species

Using the PPG I circumscription, , the ''Checklist of Ferns and Lycophytes of the World'' accepted the following species and hybrids. *'' Blechnum anthracinum'' R.C.Moran *'' Blechnum appendiculatum'' Willd. *''
Blechnum arcuatum ''Blechnum arcuatum'', known locally as ''quil-quil'', is a fern native with a natural range in Chile ranging from Ñuble River (~36° S) in the north to Aysén Region (~47° S) including adjacent areas of Argentina. It is found from sea level up ...
'' J.Rémy ex Fée *''
Blechnum areolatum ''Blechnum'', known as hard fern, is a genus of ferns in the family Blechnaceae, subfamily Blechnoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). Two very different circumscriptions of the genus are used by d ...
'' V.A.O.Dittrich & Salino *''
Blechnum asplenioides ''Blechnum'', known as hard fern, is a genus of ferns in the family Blechnaceae, subfamily Blechnoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). Two very different circumscriptions of the genus are used by ...
'' Sw. *'' Blechnum auriculatum'' Cav. *''
Blechnum australe ''Blechnum'', known as hard fern, is a genus of ferns in the family Blechnaceae, subfamily Blechnoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). Two very different circumscriptions of the genus are used by ...
'' L. *''
Blechnum austrobrasilianum ''Blechnum'', known as hard fern, is a genus of ferns in the family Blechnaceae, subfamily Blechnoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). Two very different circumscriptions of the genus are used by d ...
'' de la Sota * ''Blechnum'' × ''caudatum'' Cav. * ''Blechnum'' × ''confluens'' Schltdl. & Cham. * ''Blechnum'' × ''falciculatum'' C.Presl *''
Blechnum gracile ''Blechnum'', known as hard fern, is a genus of ferns in the family Blechnaceae, subfamily Blechnoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). Two very different circumscriptions of the genus are used by d ...
'' Kaulf. *''
Blechnum gracilipes ''Blechnum'', known as hard fern, is a genus of ferns in the family Blechnaceae, subfamily Blechnoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). Two very different circumscriptions of the genus are used by d ...
'' (Rosenst.) M.Kessler & A.R.Sm. *''
Blechnum guayanense ''Blechnum'', known as hard fern, is a genus of ferns in the family Blechnaceae, subfamily Blechnoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). Two very different circumscriptions of the genus are used by d ...
'' A.Rojas *''
Blechnum hastatum ''Blechnum'', known as hard fern, is a genus of ferns in the family Blechnaceae, subfamily Blechnoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). Two very different circumscriptions of the genus are used by d ...
'' Kaulf. *''
Blechnum heringeri ''Blechnum'', known as hard fern, is a genus of ferns in the family Blechnaceae, subfamily Blechnoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). Two very different circumscriptions of the genus are used by d ...
'' Brade *''
Blechnum laevigatum ''Blechnum'', known as hard fern, is a genus of ferns in the family Blechnaceae, subfamily Blechnoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). Two very different circumscriptions of the genus are used by d ...
'' Cav. *''
Blechnum lanceola ''Blechnum'', known as hard fern, is a genus of ferns in the family Blechnaceae, subfamily Blechnoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). Two very different circumscriptions of the genus are used by d ...
'' Sw. * ''Blechnum'' × ''leopoldense'' (Dutra) V.A.O.Dittrich & Salino *'' Blechnum longipilosum'' V.A.O.Dittrich & Salino *''
Blechnum ludificans ''Blechnum'', known as hard fern, is a genus of ferns in the family Blechnaceae, subfamily Blechnoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). Two very different circumscriptions of the genus are used by ...
'' Herter *'' Blechnum malacothrix'' Maxon & Morton *'' Blechnum meridense'' Klotzsch *''
Blechnum occidentale ''Blechnum occidentale'' is a fern in the family Blechnaceae. Its native range is from Mexico through Central America and the Caribbean to tropical South America. ''Blechnum scaberulum'' and ''Blechnum sodiroi'' have been regarded as separate ...
'' L. * ''Blechnum'' × ''pampasicum'' de la Sota & M.L.Durán *'' Blechnum polypodioides'' Raddi *'' Blechnum punctulatum'' Sw. *'' Blechnum tomentosum'' M.Kessler & A.R.Sm.


References


Further reading

* Huxley, A., ed. (1992). ''New RHS Dictionary of Gardening''. Macmillan * Moore, D. M. (1983). ''Flora of Tierra del Fuego''


External links


Trees and shrubs in the Andes of Ecuador: ''Blechnum''
{{Taxonbar, from=Q2049575 Blechnaceae Fern genera Taxa named by Carl Linnaeus