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The blackpoll warbler (''Setophaga striata'') is a
New World warbler The New World warblers or wood-warblers are a group of small, often colorful, passerine birds that make up the family Parulidae and are restricted to the New World. They are not closely related to Old World warblers or Australian warblers. Mos ...
. Breeding males are mostly black and white. They have a prominent black cap, white cheeks and white wing bars. The blackpoll breeds in forests of northern North America, from
Alaska Alaska ( ; russian: Аляска, Alyaska; ale, Alax̂sxax̂; ; ems, Alas'kaaq; Yup'ik: ''Alaskaq''; tli, Anáaski) is a state located in the Western United States on the northwest extremity of North America. A semi-exclave of the U.S. ...
throughout most of
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...
, to the mountains of New York and
New England New England is a region comprising six states in the Northeastern United States: Connecticut, Maine, Massachusetts, New Hampshire, Rhode Island, and Vermont. It is bordered by the state of New York (state), New York to the west and by the Can ...
. They are a common migrant through much of North America. In fall, they fly south to the
Greater Antilles The Greater Antilles ( es, Grandes Antillas or Antillas Mayores; french: Grandes Antilles; ht, Gwo Zantiy; jam, Grieta hAntiliiz) is a grouping of the larger islands in the Caribbean Sea, including Cuba, Hispaniola, Puerto Rico, Jamaica, a ...
and the northeastern coasts of
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the sout ...
in a non-stop long-distance migration over open water, averaging 2500 km, one of the longest distance non-stop overwater flights ever recorded for a migratory songbird. Rare vagrants to western
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a subcontinent of Eurasia and it is located entirel ...
, they are one of the more frequent transatlantic
passerine A passerine () is any bird of the order Passeriformes (; from Latin 'sparrow' and '-shaped'), which includes more than half of all bird species. Sometimes known as perching birds, passerines are distinguished from other orders of birds by th ...
wanderers.


Etymology

The genus name ''Setophaga'' is from
Ancient Greek Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic p ...
''ses'', "moth", and ', "eating", and the specific ''striata'' means "striped".


Description

The blackpoll warbler is a fairly small bird which attains the weight of a ball point pen. However, it is one of the larger of the diverse genus ''
Setophaga ''Setophaga'' is a genus of birds of the New World warbler family Parulidae. It contains at least 33 species. The males in breeding plumage are often highly colorful. The ''Setophaga'' warblers are an example of adaptive radiation with the vario ...
'' (formerly ''Dendroica''). In the species, body length can vary from and wingspan can range from . Body mass can vary from , with an average bird anywhere between . Among standard measurements, the wing chord is , the
tail The tail is the section at the rear end of certain kinds of animals’ bodies; in general, the term refers to a distinct, flexible appendage to the torso. It is the part of the body that corresponds roughly to the sacrum and coccyx in mammal ...
is , the
bill Bill(s) may refer to: Common meanings * Banknote, paper cash (especially in the United States) * Bill (law), a proposed law put before a legislature * Invoice, commercial document issued by a seller to a buyer * Bill, a bird or animal's beak Plac ...
is and the tarsus is . The summer male blackpoll warblers have dark-streaked brown backs, white faces and black crowns. Their underparts are white with black streaks, and they display two white wing bars. The adult females essentially resemble washed-out versions of the summer males, and in particular, the females lack the strong head patterns, and their crowns and faces are shades of gray. Another outstanding physical characteristic of the species are the bright orange, pink legs. Non-breeding birds of this species have greenish heads, dark-streaked greenish upperparts and yellowish breasts, with the yellow extending to the belly in young birds. Their wing bars are always present.


Habitat

In the southern portion of their breeding range, blackpoll warblers can be found on the higher elevations of mountains in woodland or brushy areas. They also spend their summers on the wooded coastal islands of Maine and the Maritime Provinces. Farther north they have been reported throughout the boreal coniferous forest. Blackpolls breed nearer to the tundra than any other warbler.


Behavior

Although fairly large for a warbler, blackpoll warblers are fairly easy to miss because of their relatively inactive foraging style and tendency to perch in dense foliage near the canopy of the trees. They are more often heard than seen, though their song is one of the highest pitched known. Their songs are simple repetitions of high ''tsi'' notes. Their calls are thin ''sit''s.


Foraging and diet

The blackpoll has a deliberate feeding style with occasional flitting, hovering and hawking around branches. They are primarily insectivorous. The species appears to be quite a generalist, preying on a great diversity of adult and larval insects and spiders. Documented insect prey for the species includes lice,
locust Locusts (derived from the Vulgar Latin ''locusta'', meaning grasshopper) are various species of short-horned grasshoppers in the family Acrididae that have a swarming phase. These insects are usually solitary, but under certain circumstanc ...
s, cankerworms,
mosquito Mosquitoes (or mosquitos) are members of a group of almost 3,600 species of small flies within the family Culicidae (from the Latin ''culex'' meaning " gnat"). The word "mosquito" (formed by ''mosca'' and diminutive ''-ito'') is Spanish for "li ...
es,
webworm The fall webworm (''Hyphantria cunea'') is a moth in the family Erebidae known principally for its larval stage, which creates the characteristic webbed nests on the tree limbs of a wide variety of hardwoods in the late summer and fall. It is con ...
s,
ant Ants are eusocial insects of the family Formicidae and, along with the related wasps and bees, belong to the order Hymenoptera. Ants evolved from vespoid wasp ancestors in the Cretaceous period. More than 13,800 of an estimated total of ...
s,
termite Termites are small insects that live in colonies and have distinct castes (eusocial) and feed on wood or other dead plant matter. Termites comprise the infraorder Isoptera, or alternatively the epifamily Termitoidae, within the order Blatto ...
s,
gnat A gnat () is any of many species of tiny flying insects in the dipterid suborder Nematocera, especially those in the families Mycetophilidae, Anisopodidae and Sciaridae. They can be both biting and non-biting. Most often they fly in large ...
s,
aphid Aphids are small sap-sucking insects and members of the superfamily Aphidoidea. Common names include greenfly and blackfly, although individuals within a species can vary widely in color. The group includes the fluffy white woolly aphids. A t ...
s and
sawflies Sawflies are the insects of the suborder Symphyta within the order Hymenoptera, alongside ants, bees, and wasps. The common name comes from the saw-like appearance of the ovipositor, which the females use to cut into the plants where they lay ...
. It has been suggested that this species may be a
spruce budworm ''Choristoneura'' is a genus of moths in the family Tortricidae. Several species are serious pests of conifers, such as spruce and are known as spruce budworms. Species *'' Choristoneura adumbratanus'' (Walsingham, 1900) *'' Choristoneura afri ...
specialist, but there is no obvious connection between population trends of the two species. The blackpoll will opt for
berries A berry is a small, pulpy, and often edible fruit. Typically, berries are juicy, rounded, brightly colored, sweet, sour or tart, and do not have a stone or pit, although many pips or seeds may be present. Common examples are strawberries, rasp ...
in migration and during winter. They often forage high in trees, and sometimes catch insects while in flight.


Reproduction

Their breeding habitats are coniferous woodlands, especially those in which spruce trees grow. The bird's breeding ranges extend to the
taiga Taiga (; rus, тайга́, p=tɐjˈɡa; relates to Mongolic and Turkic languages), generally referred to in North America as a boreal forest or snow forest, is a biome characterized by coniferous forests consisting mostly of pines, spruc ...
. Blackpoll warblers commonly nest in a relatively low site of a conifer. They lay 3–5 eggs in a cup-shaped nest, rarely up to 9. The eggs are incubated for around 12 days and the young leave the nest when they are only 10 days old, before they can fly well. Their parents feed them for a total of around two weeks. Mated females usually begin second nests right away and leave post-fledging parental duties to their mates. The high incidence of double brooding, coupled with and partly a function of low nest predation and parasitism rates, results in high annual productivity for this species.


Migration

The blackpoll warbler's transoceanic flight has been the subject of over twenty-five scientific studies. Sources of data include radar observations, bird banding and weights taken, dead birds recovered from field sites and fatal obstacles. It is unknown if they feed on insects while in flight. Blackpoll warblers have the longest migration of any species of
New World warbler The New World warblers or wood-warblers are a group of small, often colorful, passerine birds that make up the family Parulidae and are restricted to the New World. They are not closely related to Old World warblers or Australian warblers. Mos ...
. This is likely the reason that they are one of the later warblers to appear in spring migration, after one or more short overwater flights and a relatively prolonged movement overland after through North America anytime from early May to mid-June. The peak of their migration is in late May, when most warblers are on their breeding grounds. In the fall the birds migrate from their breeding grounds across the northern latitudes. They converge on the Northeastern United States south to Virginia starting in mid-August.Baird, James (1999)
''Returning to the Tropics: The Epic Autumn Flight of the Blackpoll Warbler''
in Kenneth P. Able's Gathering of Angels. Ithaca, New York: Cornell University Press, pp. 63–78.
Most blackpolls fly directly from northeastern North America over the Atlantic Ocean to their winter range. Data from nocturnal accidents, banding stations and sightings have shown that blackpolls are rare autumn migrants south of
Cape Hatteras Cape Hatteras is a cape located at a pronounced bend in Hatteras Island, one of the barrier islands of North Carolina. Long stretches of beach, sand dunes, marshes, and maritime forests create a unique environment where wind and waves shap ...
, North Carolina, whereas north of Cape Hatteras they are common. Part of the fall migratory route of the blackpoll warbler is over the
Atlantic Ocean The Atlantic Ocean is the second-largest of the world's five oceans, with an area of about . It covers approximately 20% of Earth's surface and about 29% of its water surface area. It is known to separate the " Old World" of Africa, Europe ...
from the northeastern
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territori ...
to the Greater and
Lesser Antilles The Lesser Antilles ( es, link=no, Antillas Menores; french: link=no, Petites Antilles; pap, Antias Menor; nl, Kleine Antillen) are a group of islands in the Caribbean Sea. Most of them are part of a long, partially volcanic island arc bet ...
, or northern
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the sout ...
. Island stopovers at Bermuda and other places are evidence of migratory pathways. To accomplish this flight, the blackpoll warbler nearly doubles its body mass in staging areas and takes advantage of a shift in prevailing wind direction to direct it to its destination. When they fly southward over the Atlantic they burn, 0.08 g of fat every hour. This route averages over water, requiring a potentially nonstop flight of around 72 to 88 hours. They travel at a speed of about . Blackpolls can weigh more than when they leave the United States and lose 4 or more grams by the time they reach South America. Some of the blackpolls land in Bermuda before going on. Some birds, often with lower body weights, do not make it. Using a tiny
light level geolocator A light level geolocator, light-level logger or GLS (global location sensor) is essentially a lightweight, electronic archival tracking device, usually used in bird migration research to map migration routes, identify important staging areas, and ...
biologists have proven that the black poll flies an average of 2540 km (2270 to 2770 km) non-stop over an average of 62 h, up to 3 days, corresponding to about 41 km/h. In 2013, 37 blackpolls from Vermont and Nova Scotia carried a miniaturized geolocator weighing 0.5 g with harness on their back. The device recorded light-levels, from which longitudes and latitudes could be estimated, and in 2014 the scientists recovered five of the original 37. Four of the five birds departed from western Nova Scotia between September 25 and October 21, and traveled at speeds between 10.7 and 13.4 meters per second. The study revealed that the spring migration overland and the autumn routes overwater were "dramatically different". When the flight distance per body mass was compared to other birds, only the
ruby-throated hummingbird The ruby-throated hummingbird (''Archilochus colubris'') is a species of hummingbird that generally spends the winter in Central America, Mexico, and Florida, and migrates to Canada and other parts of Eastern North America for the summer to bree ...
might travel more kilometres per gram (estimated around 210–280 km/g vs. 233 km/g for blackpolls).


References


External links



– Cornell Lab of Ornithology

– USGS Patuxent Bird Identification InfoCenter

for: Grenada, Guyana at bird-stamps.org * * {{Taxonbar, from=Q27075924
blackpoll warbler The blackpoll warbler (''Setophaga striata'') is a New World warbler. Breeding males are mostly black and white. They have a prominent black cap, white cheeks and white wing bars. The blackpoll breeds in forests of northern North America, from Ala ...
Native birds of Alaska Birds of Canada Birds of South America Birds of the Dominican Republic
blackpoll warbler The blackpoll warbler (''Setophaga striata'') is a New World warbler. Breeding males are mostly black and white. They have a prominent black cap, white cheeks and white wing bars. The blackpoll breeds in forests of northern North America, from Ala ...