Buteogallus Daggetti
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''Buteogallus daggetti'', occasionally called "Daggett's eagle" or the "walking eagle", is an extinct species of long-legged
hawk Hawks are birds of prey of the family Accipitridae. They are very widely distributed and are found on all continents, except Antarctica. The subfamily Accipitrinae includes goshawks, sparrowhawks, sharp-shinned hawks, and others. This ...
which lived in southwest North America during the
Pleistocene The Pleistocene ( ; referred to colloquially as the ''ice age, Ice Age'') is the geological epoch (geology), epoch that lasted from to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fin ...
. Initially believed to be some sort of carrion-eating
eagle Eagle is the common name for the golden eagle, bald eagle, and other birds of prey in the family of the Accipitridae. Eagles belong to several groups of Genus, genera, some of which are closely related. True eagles comprise the genus ''Aquila ( ...
, it was for some time placed in the distinct genus ''Wetmoregyps'', named for
Alexander Wetmore Frank Alexander Wetmore (June 18, 1886 – December 7, 1978) was an American ornithologist and avian paleontologist. He was the sixth Secretary of the Smithsonian Institution. He was also an elected member of both the American Philosophical Soc ...
. It probably resembled a larger version of the modern-day savanna hawk, with its long legs possibly used like the
secretarybird The secretarybird or secretary bird (''Sagittarius serpentarius'') is a large bird of prey that is Endemism, endemic to Africa. It is mostly terrestrial, spending most of its time on the ground, and is usually found in the open grasslands and s ...
of Africa to hunt for small reptiles from a safe distance. It died out about 13,000 years ago. Fossils of ''B. daggetti'' were discovered in the La Brea and
Carpinteria Carpinteria (; , meaning "Carpentry") is a small seaside city in southeastern Santa Barbara County, California. Located on the Central Coast of California, it had a population of 13,264 at the 2020 census. Carpinteria is a popular surf destin ...
lagerstätte A Fossil-Lagerstätte (, from ''Lager'' 'storage, lair' '' Stätte'' 'place'; plural ''Lagerstätten'') is a sedimentary deposit that preserves an exceptionally high amount of palaeontological information. ''Konzentrat-Lagerstätten'' preserv ...
in southern California, and in
Nuevo León Nuevo León, officially the Free and Sovereign State of Nuevo León, is a Administrative divisions of Mexico, state in northeastern Mexico. The state borders the Mexican states of Tamaulipas, Coahuila, Zacatecas, and San Luis Potosí, San Luis ...
in Mexico. Its habitat included grasslands, marshlands, brushy savannas and ponds. It probably ate mostly small reptiles such as snakes. As is often the case with birds of prey, the female seems to have been larger than the male.


Description

In life, ''B. daggetti'' is thought to have resembled the related savanna hawk, also of the genus ''Buteogallus'': ''B. daggetti'' had long legs which earned it the nickname "walking eagle". Miller (1915) concluded that these legs had little lifting power and weakly grasping claws compared to eagles that hunt on the wing. Conversely, the points of strong muscle attachment on the shaft of the tarsus suggest walking proficiency. He likened this "degeneracy" to the relationship of the
roadrunner The roadrunners (genus ''Geococcyx''), also known as chaparral birds or chaparral cocks, are two species of fast-running ground cuckoos with long tails and crests. They are found in the southwestern and south-central United States, Mexico and C ...
to tree-dwelling
cuckoo Cuckoos are birds in the Cuculidae ( ) family, the sole taxon in the order Cuculiformes ( ). The cuckoo family includes the common or European cuckoo, roadrunners, koels, malkohas, couas, coucals, and anis. The coucals and anis are somet ...
s. As with many birds, its females were larger than the males, an example of
sexual dimorphism Sexual dimorphism is the condition where sexes of the same species exhibit different Morphology (biology), morphological characteristics, including characteristics not directly involved in reproduction. The condition occurs in most dioecy, di ...
. However, it is some 40 percent larger than the savanna hawk. This gives it a total length of , including of tail; its
wingspan The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the opposite wingtip. For example, the Boeing 777–200 has a wingspan of , and a wandering albatross (''Diomedea exulans'') caught in 1965 had a wingsp ...
would have been . Body weight is estimated at . This makes it quite a heavy raptor, slightly greater than the present-day
secretarybird The secretarybird or secretary bird (''Sagittarius serpentarius'') is a large bird of prey that is Endemism, endemic to Africa. It is mostly terrestrial, spending most of its time on the ground, and is usually found in the open grasslands and s ...
of Africa. This African raptor shares the long legs of ''B. daggetti'' and their lifestyles may have been similar (see "
Ecology Ecology () is the natural science of the relationships among living organisms and their Natural environment, environment. Ecology considers organisms at the individual, population, community (ecology), community, ecosystem, and biosphere lev ...
" below).


History

''Buteogallus daggetti'' was first described by Loye Holmes Miller in 1915 as ''Morphnus daggetti'', from a
tarsometatarsus The tarsometatarsus is a bone that is only found in the lower leg of birds and some non-avian dinosaurs. It is formed from the fusion of several bird bones found in other types of animals, and homologous to the mammalian tarsus (ankle bones) a ...
found in the
La Brea Tar Pits La Brea Tar Pits comprise an active Paleontological site, paleontological research site in urban Los Angeles. Hancock Park was formed around a group of tar pits where natural Bitumen, asphalt (also called asphaltum, bitumen, or pitch; ''brea'' ...
. The bone's great length, comparable to a
great blue heron The great blue heron (''Ardea herodias'') is a large wading bird in the heron family Ardeidae, common near the shores of open water and in wetlands over most of North and Central America, as well as far northwestern South America, the Caribbea ...
's, gave him the impression of an "eagle on stilts" and led to comparisons with the
secretarybird The secretarybird or secretary bird (''Sagittarius serpentarius'') is a large bird of prey that is Endemism, endemic to Africa. It is mostly terrestrial, spending most of its time on the ground, and is usually found in the open grasslands and s ...
. The
specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
pays tribute to F.S. Daggett of the
Natural History Museum of Los Angeles County Nature is an inherent character or constitution, particularly of the ecosphere or the universe as a whole. In this general sense nature refers to the laws, elements and phenomena of the physical world, including life. Although humans are par ...
(at the time the Museum of History, Science, and Art). By 1925, Miller had assigned three more tarsometatarsi and an incomplete
tibiotarsus The tibiotarsus is the large bone between the femur and the tarsometatarsus in the leg of a bird. It is the fusion of the proximal part of the tarsus with the tibia. A similar structure also occurred in the Mesozoic Heterodontosauridae. These ...
to the genus. In 1932,
Hildegarde Howard Hildegarde Howard (April 3, 1901 – February 28, 1998) was an American pioneer in paleornithology. She was mentored by the famous ornithologist, Joseph Grinnell, at the Museum of Vertebrate Zoology (MVZ) and in avian paleontology.Joyce Harvey & ...
assigned a
coracoid A coracoid is a paired bone which is part of the shoulder assembly in all vertebrates except therian mammals (marsupials and placentals). In therian mammals (including humans), a coracoid process is present as part of the scapula, but this is n ...
and (tentatively) some
phalanges The phalanges (: phalanx ) are digit (anatomy), digital bones in the hands and foot, feet of most vertebrates. In primates, the Thumb, thumbs and Hallux, big toes have two phalanges while the other Digit (anatomy), digits have three phalanges. ...
to ''Wetmoregyps'' from specimens at the tar pits. She noted these fossils were difficult to distinguish from those of
Woodward's eagle Woodward's eagle (''Buteogallus woodwardi'') is an extinct species of black hawk that lived in North America and the Caribbean during the Late Pleistocene. Remains have been found in the La Brea Tar Pits in the United States and in Cuba. Despite ...
(''Buteogallus woodwardi''), a huge bird of prey in the same genus as the savanna hawk. In 1928 and 1931, having obtained more fragmentary tarsometarsi from the
Carpinteria Carpinteria (; , meaning "Carpentry") is a small seaside city in southeastern Santa Barbara County, California. Located on the Central Coast of California, it had a population of 13,264 at the 2020 census. Carpinteria is a popular surf destin ...
asphalt, he reexamined his conclusions and found the bird more similar to '' Caracara'' and the
great black hawk The great black hawk (''Buteogallus urubitinga'') is a bird of prey in the family Accipitridae, which also includes the eagles, hawks, and Old World vultures. Taxonomy The great black hawk was formally described in 1788 by the German naturalist ...
(''Buteogallus urubitinga''). He nevertheless assigned it to the new genus ''Wetmoregyps'', possibly because it was much bigger than those aforementioned birds. In 1943 Miller wrote on two coracoids and a tarsometatarsus from San Josecito Cavern in
Nuevo León Nuevo León, officially the Free and Sovereign State of Nuevo León, is a Administrative divisions of Mexico, state in northeastern Mexico. The state borders the Mexican states of Tamaulipas, Coahuila, Zacatecas, and San Luis Potosí, San Luis ...
, Mexico.
Storrs L. Olson Storrs Lovejoy Olson (April 3, 1944 – January 20, 2021) was an American biologist and ornithologist who spent his career at the Smithsonian Institution, retiring in 2008. One of the world's foremost avian paleontologists, he was best known ...
studied the bird in 2007 and placed it in the genus ''Buteogallus'' owing to its similarity to the savanna hawk.


Behavior and ecology

''Buteogallus daggetti'' habitats comprised open grasslands, marshlands, and savannas from sea level to . Because of its large size and long legs, ''B. daggetti'' is theorized to have lived rather like the modern-day
secretarybird The secretarybird or secretary bird (''Sagittarius serpentarius'') is a large bird of prey that is Endemism, endemic to Africa. It is mostly terrestrial, spending most of its time on the ground, and is usually found in the open grasslands and s ...
. Its diet would have been composed mostly of snakes and other small reptiles, which it would have kicked to death with its long legs. Not just good weapons, these legs would have kept the bird at a safe distance from its struggling prey. Olson's paper guessed that ''B. daggetti'' may have been attracted to grass fires as the savanna hawk is today. These fires chase out small animals, which then make easy prey for the raptors. Olson disputes claims that ''B. daggetti'' was primarily a scavenger, and that its extinction was somehow linked to the decline of the North American
Pleistocene megafauna The Late Pleistocene to the beginning of the Holocene saw the extinction of the majority of the world's megafauna, typically defined as animal species having body masses over , which resulted in a collapse in faunal density and diversity acro ...
. He likewise found Miller's vision of the bird as a forest-dweller "spurious".


References

{{Taxonbar, from=Q5208537 Buteogallus Pleistocene birds of North America Extinct birds of North America Taxa named by Loye H. Miller