Blackberry (other)
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The blackberry is an edible fruit produced by many
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
in the genus ''
Rubus ''Rubus'' is a large and diverse genus of flowering plants in the rose family, Rosaceae, subfamily Rosoideae, most commonly known as brambles. Fruits of various species are known as raspberries, blackberries, dewberries, and bristleberries. ...
'' in the
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Rosaceae Rosaceae (), the rose family, is a family of flowering plants that includes 4,828 known species in 91 genera. The name is derived from the type genus '' Rosa''. The family includes herbs, shrubs, and trees. Most species are deciduous, but som ...
, hybrids among these species within the subgenus ''Rubus'', and hybrids between the subgenera ''Rubus'' and ''Idaeobatus''. The taxonomy of blackberries has historically been confused because of hybridization and
apomixis In botany, apomixis is asexual development of seed or embryo without fertilization. However, other definitions include replacement of the seed by a plantlet or replacement of the flower by bulbils. Apomictically produced offspring are geneti ...
so that species have often been grouped together and called
species aggregate In biology, a species complex is a group of closely related organisms that are so similar in appearance and other features that the boundaries between them are often unclear. The taxa in the complex may be able to hybridize readily with each oth ...
s. Blackberry fruit production is abundant with annual volumes of per possible, making this plant commercially attractive. ''
Rubus armeniacus ''Rubus armeniacus'', the Himalayan blackberry or Armenian blackberry, is a species of ''Rubus'' in the blackberry group ''Rubus'' subgenus ''Rubus'' series ''Discolores'' (P.J. Müll.) Focke. It is native to Armenia and northern Iran, and widel ...
'' ("Himalayan" blackberry) is considered a
noxious weed A noxious weed, harmful weed or injurious weed is a weed that has been designated by an agricultural or other governing authority as a plant that is harmful to agricultural or horticultural crops, natural habitats or ecosystems, or humans or lives ...
and
invasive species An invasive species is an introduced species that harms its new environment. Invasive species adversely affect habitats and bioregions, causing ecological, environmental, and/or economic damage. The term can also be used for native spec ...
in many regions of the
Pacific Northwest The Pacific Northwest (PNW; ) is a geographic region in Western North America bounded by its coastal waters of the Pacific Ocean to the west and, loosely, by the Rocky Mountains to the east. Though no official boundary exists, the most common ...
of Canada and the United States, where it grows out of control in urban and suburban parks and woodlands.


Description

The blackberry looks similar to its
raspberry The raspberry is the edible fruit of several plant species in the genus ''Rubus'' of the Rosaceae, rose family, most of which are in the subgenus ''Rubus#Modern classification, Idaeobatus''. The name also applies to these plants themselves. Ras ...
relatives. What distinguishes the blackberry is the connection between the berry and the torus ( receptacle or stem). When a raspberry is picked, the torus remains on the vine. This is why there is a hollow core in the raspberry fruit. When a blackberry is picked, the torus stays with the fruit. The term ''
bramble ''Rubus'' is a large and diverse genus of flowering plants in the rose family, Rosaceae, subfamily Rosoideae, most commonly known as brambles. Fruits of various species are known as raspberries, blackberries, dewberries, and bristleberries. I ...
'', a word referring to any impenetrable
thicket A thicket is a very dense stand of trees or tall shrubs, often dominated by only one or a few species, to the exclusion of all others. They may be formed by species that shed large numbers of highly viable seeds that are able to germinate in th ...
, has in some circles traditionally been applied specifically to the blackberry or its products, though in the United States it applies to all members of the genus ''Rubus''. In the western US, the term ''caneberry'' is used to refer to blackberries and raspberries as a group rather than the term bramble. '' Briar'' or ''brier'' may be used to refer to the dense vines of the plant, though this name is used for other thorny thickets (such as ''
Smilax ''Smilax'' is a genus of about 300–350 species, found in the tropics and subtropics worldwide. They are climbing flowering plants, many of which are woody and/or thorny, in the monocotyledon family (biology), family Smilacaceae, native through ...
'') as well. The usually black fruit is not a
berry A berry is a small, pulpy, and often edible fruit. Typically, berries are juicy, rounded, brightly colored, sweet, sour or tart, and do not have a stone or pit although many pips or seeds may be present. Common examples of berries in the cul ...
in the
botanical Botany, also called plant science, is the branch of natural science and biology studying plants, especially Plant anatomy, their anatomy, Plant taxonomy, taxonomy, and Plant ecology, ecology. A botanist or plant scientist is a scientist who s ...
sense, as it is termed botanically as an
aggregate fruit A raspberry_beetle.html" ;"title="raspberry fruit (shown with a raspberry beetle">raspberry fruit (shown with a raspberry beetle larva) is an aggregate fruit, an aggregate of drupelets. image:Aquilegia vulgaris 004.JPG, The fruit of an ''Aquilegi ...
, composed of small
drupe In botany, a drupe (or stone fruit) is a type of fruit in which an outer fleshy part (exocarp, or skin, and mesocarp, or flesh) surrounds a single shell (the ''pip'' (UK), ''pit'' (US), ''stone'', or ''pyrena'') of hardened endocarp with a seed ...
lets. It is a widespread and well-known group of over 375 species, many of which are closely related apomictic microspecies
native Native may refer to: People * '' Jus sanguinis'', nationality by blood * '' Jus soli'', nationality by location of birth * Indigenous peoples, peoples with a set of specific rights based on their historical ties to a particular territory ** Nat ...
throughout Europe, northwestern Africa,
temperate In geography, the temperate climates of Earth occur in the middle latitudes (approximately 23.5° to 66.5° N/S of the Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ran ...
western and central Asia and North and South America.


Plants

Blackberries are
perennial In horticulture, the term perennial ('' per-'' + '' -ennial'', "through the year") is used to differentiate a plant from shorter-lived annuals and biennials. It has thus been defined as a plant that lives more than 2 years. The term is also ...
plants bearing
biennial Biennial means (an event) lasting for two years or occurring every two years. The related term biennium is used in reference to a period of two years. In particular, it can refer to: * Biennial plant, a plant which blooms in its second year and t ...
stems (called ''canes'') from their roots. In its first year, a new stem, the ''primocane'', reaches a full length of about trailing on the ground and bearing large palmate compound
leaves A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, ...
with 5–7 new leaves; it does not produce any flowers. In its second year, the cane is a ''floricane'' with a non-growing stem. The lateral buds open to produce flowering laterals. First- and second-year shoots produce short-curved, sharp thorns. Thornless cultivars have been developed during the early 21st century. Unmanaged plants tend to aggregate in a dense tangle of stems and branches, which can be controlled in gardens or farms using trellises. Blackberry shrubs can tolerate poor
soil Soil, also commonly referred to as earth, is a mixture of organic matter, minerals, gases, water, and organisms that together support the life of plants and soil organisms. Some scientific definitions distinguish dirt from ''soil'' by re ...
s, spreading readily in wasteland, ditches, and roadsides. The
flower Flowers, also known as blooms and blossoms, are the reproductive structures of flowering plants ( angiosperms). Typically, they are structured in four circular levels, called whorls, around the end of a stalk. These whorls include: calyx, m ...
s bloom in late spring and early summer on the tips of branches. Each flower is about in diameter, with five white-pink
petal Petals are modified leaves that form an inner whorl surrounding the reproductive parts of flowers. They are often brightly coloured or unusually shaped to attract pollinators. All of the petals of a flower are collectively known as the ''corol ...
s. The fruit only develops around ovules fertilized by the male
gamete A gamete ( ) is a Ploidy#Haploid and monoploid, haploid cell that fuses with another haploid cell during fertilization in organisms that Sexual reproduction, reproduce sexually. Gametes are an organism's reproductive cells, also referred to as s ...
from a pollen grain. The most likely cause of undeveloped ovules is inadequate
pollinator A pollinator is an animal that moves pollen from the male anther of a flower to the female carpel, stigma of a flower. This helps to bring about fertilization of the ovules in the flower by the male gametes from the pollen grains. Insects are ...
visits. Infection with raspberry bushy dwarf virus sometimes causes no visible symptoms and sometimes causes incomplete drupelet development and crumbly fruit.


Genetics

The loci controlling the primocane fruiting was mapped in the F Locus, on LG7, whereas thorns/thornlessness was mapped on LG4. Better understanding of the genetics is useful for genetic screening of cross-breds, and for genetic engineering purposes.


Ecology

Blackberry leaves are food for certain
caterpillar Caterpillars ( ) are the larval stage of members of the order Lepidoptera (the insect order comprising butterflies and moths). As with most common names, the application of the word is arbitrary, since the larvae of sawflies (suborder ...
s; some grazing mammals, especially deer, are also very fond of the leaves. Caterpillars of the
concealer moth Oecophoridae (concealer moths) is a family of small moths in the superfamily Gelechioidea. The phylogeny and systematics of gelechoid moths are still not fully resolved, and the circumscription of the Oecophoridae is strongly affected by this. ...
'' Alabonia geoffrella'' have been found feeding inside dead blackberry shoots. When mature, the berries are eaten and their seeds dispersed by mammals, such as the red fox, American black bear, and the Eurasian badger, as well as by small birds. Blackberries grow wild throughout most of Europe. They are an important element in the ecology of many countries, and harvesting the berries is a common pastime. However, their vigorous growth and tendency to grow unchecked if not managed correctly mean that the plants are also considered a weed, sending down roots from branches that touch the ground, and sending up suckers from the roots. In some parts of the world, such as in Australia, Chile, New Zealand, and the
Pacific Northwest The Pacific Northwest (PNW; ) is a geographic region in Western North America bounded by its coastal waters of the Pacific Ocean to the west and, loosely, by the Rocky Mountains to the east. Though no official boundary exists, the most common ...
of North America, some blackberry species, particularly ''
Rubus armeniacus ''Rubus armeniacus'', the Himalayan blackberry or Armenian blackberry, is a species of ''Rubus'' in the blackberry group ''Rubus'' subgenus ''Rubus'' series ''Discolores'' (P.J. Müll.) Focke. It is native to Armenia and northern Iran, and widel ...
'' (Himalayan blackberry) and '' Rubus laciniatus'' (evergreen blackberry), are
naturalized Naturalization (or naturalisation) is the legal act or process by which a non-national of a country acquires the nationality of that country after birth. The definition of naturalization by the International Organization for Migration of the ...
and considered an
invasive species An invasive species is an introduced species that harms its new environment. Invasive species adversely affect habitats and bioregions, causing ecological, environmental, and/or economic damage. The term can also be used for native spec ...
and a
noxious weed A noxious weed, harmful weed or injurious weed is a weed that has been designated by an agricultural or other governing authority as a plant that is harmful to agricultural or horticultural crops, natural habitats or ecosystems, or humans or lives ...
. Blackberry fruits are red when unripe, leading to an old expression that "blackberries are red when they're green".


Cultivation

Worldwide, Mexico is the leading producer of blackberries, with nearly the entire crop being produced for export into the off-season fresh markets in North America and Europe. Until 2018, the Mexican market was almost entirely based on the cultivar 'Tupy' (often spelled 'Tupi', but the EMBRAPA program in Brazil from which it was released prefers the 'Tupy' spelling), but Tupy fell out of favor in some Mexican growing regions. In the US,
Oregon Oregon ( , ) is a U.S. state, state in the Pacific Northwest region of the United States. It is a part of the Western U.S., with the Columbia River delineating much of Oregon's northern boundary with Washington (state), Washington, while t ...
is the leading commercial blackberry producer, producing on in 2017. Numerous
cultivar A cultivar is a kind of Horticulture, cultivated plant that people have selected for desired phenotypic trait, traits and which retains those traits when Plant propagation, propagated. Methods used to propagate cultivars include division, root a ...
s have been selected for commercial and amateur cultivation in Europe and the United States. Since the many species form
hybrid Hybrid may refer to: Science * Hybrid (biology), an offspring resulting from cross-breeding ** Hybrid grape, grape varieties produced by cross-breeding two ''Vitis'' species ** Hybridity, the property of a hybrid plant which is a union of two diff ...
s easily, there are numerous cultivars with more than one species in their ancestry.


History

Modern hybridization and
cultivar A cultivar is a kind of Horticulture, cultivated plant that people have selected for desired phenotypic trait, traits and which retains those traits when Plant propagation, propagated. Methods used to propagate cultivars include division, root a ...
development took place mostly in the US. In 1880, a hybrid blackberry-raspberry named the
loganberry The loganberry (''Rubus'' × ''loganobaccus'') is a hybrid of the North American blackberry ('' Rubus ursinus'') and the European raspberry (''Rubus idaeus''), accidentally bred in 1881 by James Harvey Logan, for whom they are named. They are ...
was developed in
Santa Cruz, California Santa Cruz (Spanish language, Spanish for "Holy Cross") is the largest city and the county seat of Santa Cruz County, California, Santa Cruz County, in Northern California. As of the 2020 United States census, 2020 census, the city population ...
, by an American judge and
horticulturalist Horticulture (from ) is the art and science of growing fruits, vegetables, flowers, trees, shrubs and ornamental plants. Horticulture is commonly associated with the more professional and technical aspects of plant cultivation on a smaller and mo ...
,
James Harvey Logan James Harvey Logan (December 8, 1841 – July 16, 1928) was a judge in Santa Cruz, California, and an amateur botanist credited with the 1881 creation of the loganberry, a cross between the raspberry and the blackberry. He was District Atto ...
. One of the first thornless varieties was developed in 1921, but the berries lost much of their flavor. Common thornless cultivars developed from the 1990s to the early 21st century by the
US Department of Agriculture The United States Department of Agriculture (USDA) is an executive department of the United States federal government that aims to meet the needs of commercial farming and livestock food production, promotes agricultural trade and producti ...
enabled efficient machine-harvesting, higher yields, larger and firmer fruit, and improved flavor, including the ''Triple Crown'', ''Black Diamond'', ''Black Pearl'', and ''Nightfall'', a
marionberry The marionberry (''Rubus L.'' subgenus ''Rubus'') is a cultivar of blackberry released in 1956 by the USDA Agricultural Research Service breeding program in cooperation with Oregon State University. It is named after Marion County, Oregon, whe ...
.


Hybrids

'Marion' (marketed as "
marionberry The marionberry (''Rubus L.'' subgenus ''Rubus'') is a cultivar of blackberry released in 1956 by the USDA Agricultural Research Service breeding program in cooperation with Oregon State University. It is named after Marion County, Oregon, whe ...
") is an important cultivar that was selected from seedlings from a cross between 'Chehalem' and 'Olallie' (commonly called "
Olallieberry The olallieberry ( ), sometimes spelled ollalieberry, olallaberry, olalliberry, ollalaberry or ollaliberry,< is the marketing name for the 'Olallie'
") berries. 'Olallie' in turn is a cross between
loganberry The loganberry (''Rubus'' × ''loganobaccus'') is a hybrid of the North American blackberry ('' Rubus ursinus'') and the European raspberry (''Rubus idaeus''), accidentally bred in 1881 by James Harvey Logan, for whom they are named. They are ...
and
youngberry The youngberry is a complex hybrid between three different berry species from the genus ''Rubus'' of the rose family: raspberry, blackberry, and dewberry. The berries of the plant are eaten fresh or used to make juice, jam, and in recipes. The ...
. 'Marion', 'Chehalem' and 'Olallie' are just three of many trailing blackberry cultivars developed by the
United States Department of Agriculture The United States Department of Agriculture (USDA) is an executive department of the United States federal government that aims to meet the needs of commercial farming and livestock food production, promotes agricultural trade and producti ...
Agricultural Research Service The Agricultural Research Service (ARS) is the principal in-house research agency of the United States Department of Agriculture (USDA). ARS is one of four agencies in USDA's Research, Education and Economics mission area. ARS is charged with ext ...
(USDA-ARS) blackberry breeding program at
Oregon State University Oregon State University (OSU) is a Public university, public Land-grant university, land-grant research university in Corvallis, Oregon, United States. OSU offers more than 200 undergraduate degree programs and a variety of graduate and doctor ...
in
Corvallis, Oregon Corvallis ( ) is a city in and the county seat of Benton County, Oregon, Benton County in central western Oregon, United States. It is the principal city of the Corvallis, Oregon Metropolitan Statistical Area, which encompasses all of Benton Co ...
. The most recent cultivars released from this program are the thornless cultivars 'Black Diamond', 'Black Pearl', and 'Nightfall' as well as the early-ripening 'Obsidian' and 'Metolius'. 'Black Diamond' is now the leading cultivar being planted in the Pacific Northwest. Some of the other cultivars from this program are 'Newberry', 'Waldo', 'Siskiyou', 'Black Butte', ' Kotata', 'Pacific', and 'Cascade'. Varieties with good commercial characteristics developed in
Arkansas Arkansas ( ) is a landlocked state in the West South Central region of the Southern United States. It borders Missouri to the north, Tennessee and Mississippi to the east, Louisiana to the south, Texas to the southwest, and Oklahoma ...
are grown in nurseries in
Oklahoma Oklahoma ( ; Choctaw language, Choctaw: , ) is a landlocked U.S. state, state in the South Central United States, South Central region of the United States. It borders Texas to the south and west, Kansas to the north, Missouri to the northea ...
. Such blackberries are easy to grow, and may produce fruit for a decade or more. These varieties have diverse flavors varying from sweet to tart.


Trailing

Trailing blackberries are vigorous and crown-forming, require a trellis for support, and are less cold-hardy than erect or semi-erect blackberries. In addition to the
Pacific Northwest The Pacific Northwest (PNW; ) is a geographic region in Western North America bounded by its coastal waters of the Pacific Ocean to the west and, loosely, by the Rocky Mountains to the east. Though no official boundary exists, the most common ...
, these types do well in similar climates, such as the United Kingdom, New Zealand, Chile, and the Mediterranean countries.


Thornless

Semi-erect, prickle-free blackberries were first developed at the
John Innes Centre The John Innes Centre (JIC), located in Norwich, Norfolk, England, is an independent centre for research and training in plant and microbial science founded in 1910. It is a registered charity (No 223852) grant-aided by the Biotechnology and B ...
in
Norwich Norwich () is a cathedral city and district of the county of Norfolk, England, of which it is the county town. It lies by the River Wensum, about north-east of London, north of Ipswich and east of Peterborough. The population of the Norwich ...
, UK, and subsequently by the USDA-ARS in
Beltsville, Maryland Beltsville is a census-designated place (CDP) in northern Prince George's County, Maryland, United States. The community was named for Truman Belt, a local landowner. The 2020 census counted 20,133 residents. Beltsville includes the unincorporate ...
. These are crown-forming and very vigorous and need a trellis for support. Cultivars include 'Black Satin', 'Chester Thornless', 'Dirksen Thornless', 'Hull Thornless', 'Loch Maree', 'Loch Ness', 'Loch Tay', 'Merton Thornless', 'Smoothstem', and 'Triple Crown'. 'Loch Ness' and 'Loch Tay' have gained the RHS's
Award of Garden Merit The Award of Garden Merit (AGM) is a long-established award for plants by the British Royal Horticultural Society (RHS). It is based on assessment of the plants' performance under UK growing conditions. It includes the full range of cultivated p ...
. The cultivar 'Cacanska Bestrna' (also called 'Cacak Thornless') has been developed in Serbia and has been planted on many thousands of hectares there.


Erect

The
University of Arkansas The University of Arkansas (U of A, UArk, or UA) is a Public university, public Land-grant university, land-grant research university in Fayetteville, Arkansas, United States. It is the Flagship campus, flagship campus of the University of Arkan ...
has developed cultivars of erect blackberries. These types are less vigorous than the semi-erect types and produce new canes from root initials (therefore they spread underground like raspberries). There are prickly and prickle-free cultivars from this program, including 'Navaho', 'Ouachita', 'Cherokee', 'Apache', 'Arapaho', and 'Kiowa'. They are also responsible for developing the primocane fruiting blackberries such as 'Prime-Jan' and 'Prime-Jim'.


Primocane

In raspberries, these types are called primocane fruiting, fall fruiting, or everbearing. 'Prime-Jim' and 'Prime-Jan' were released in 2004 by the
University of Arkansas The University of Arkansas (U of A, UArk, or UA) is a Public university, public Land-grant university, land-grant research university in Fayetteville, Arkansas, United States. It is the Flagship campus, flagship campus of the University of Arkan ...
and are the first cultivars of primocane fruiting blackberry. They grow much like the other erect cultivars described above; however, the canes that emerge in the spring will flower in midsummer and fruit in late summer or fall. The fall crop has its highest quality when it ripens in cool mild climates such as in California or the Pacific Northwest. 'Illini Hardy', a semi-erect prickly cultivar introduced by the
University of Illinois The University of Illinois Urbana-Champaign (UIUC, U of I, Illinois, or University of Illinois) is a public university, public land-grant university, land-grant research university in the Champaign–Urbana metropolitan area, Illinois, United ...
, is cane hardy in zone 5, where blackberry production has traditionally been problematic since canes often failed to survive the winter.


Mexico and Chile

Blackberry production in Mexico expanded considerably in the early 21st century. In 2017, Mexico had 97% of the
market share Market share is the percentage of the total revenue or sales in a Market (economics), market that a company's business makes up. For example, if there are 50,000 units sold per year in a given industry, a company whose sales were 5,000 of those ...
for fresh blackberries imported into the United States, while Chile had 61% of the market share for American imports of frozen blackberries. While once based on the cultivar 'Brazos', an old erect blackberry cultivar developed in Texas in 1959, the Mexican industry is now dominated by the Brazilian 'Tupy' released in the 1990s. The 'Tupy' has the erect blackberry 'Comanche', and a "wild Uruguayan blackberry" as parents. Since there are no native blackberries in Uruguay, the suspicion is that the widely grown '
Boysenberry The boysenberry is a cross between the European raspberry (''Rubus idaeus''), European blackberry ('' Rubus fruticosus''), American dewberry ('' Rubus aboriginum''), and loganberry (''Rubus'' × ''loganobaccus''). Description Boysenberrie ...
' is the male parent. To produce these blackberries in regions of Mexico where there is no winter chilling to stimulate flower bud development, chemical defoliation and application of growth regulators are used to bring the plants into bloom.


Diseases and pests

Because blackberries belong to the same genus as raspberries, they share the same diseases, including anthracnose (a type of
canker A plant canker is a small area of dead tissue, which grows slowly, often over years. Some cankers are of only minor consequence, but others are ultimately lethal and therefore can have major economic implications for agriculture and horticultur ...
), which can cause the berry to have uneven ripening. Sap flow may also be slowed. They also share the same remedies, including the Bordeaux mixture, a combination of lime, water and
copper(II) sulfate Copper(II) sulfate is an inorganic compound with the chemical formula . It forms hydrates , where ''n'' can range from 1 to 7. The pentahydrate (''n'' = 5), a bright blue crystal, is the most commonly encountered hydrate of copper(II) sulfate, whi ...
. The rows between blackberry plants must be free of weeds, blackberry suckers and grasses, which may lead to pests or diseases. Fruit growers are selective when planting blackberry bushes because wild blackberries may be infected, and gardeners are recommended to purchase only certified disease-free plants. The spotted-wing drosophila, '' Drosophila suzukii'', is a serious pest of blackberries. Unlike its vinegar fly relatives, which are primarily attracted to rotting or fermented fruit, ''D. suzukii'' attacks fresh, ripe fruit by laying eggs under the soft skin. The larvae hatch and grow in the fruit, destroying the fruit's commercial value. Another pest is ''Amphorophora rubi'', known as the blackberry aphid, which eats not just blackberries but raspberries as well. ''Byturus tomentosus'' (
raspberry beetle The raspberry beetle (''Byturus tomentosus'') is a species of beetle in the fruitworm family Byturidae. It is a major pest that is widespread in north-central Europe, affecting raspberry, blackberry and loganberry plants. This species is rela ...
), ''
Lampronia corticella ''Lampronia corticella'', the raspberry moth, is a moth of the family Prodoxidae. The species was first described by Carl Linnaeus in his 1758 10th edition of ''Systema Naturae''. It is found in most of Europe, except Iceland, the Iberian Peni ...
'' (raspberry moth) and ''
Anthonomus rubi ''Anthonomus rubi'', the strawberry-blossom weevil or strawberry blossom weevil, is a species of weevil found in Europe and western Asia. It feeds on plants of the family Rosaceae and is an important pest of strawberry (''Fragaria'' × ''ananassa ...
'' (strawberry blossom weevil) are also known to infest blackberries.


Uses


Nutrients

Raw blackberries are 88% water, 10%
carbohydrate A carbohydrate () is a biomolecule composed of carbon (C), hydrogen (H), and oxygen (O) atoms. The typical hydrogen-to-oxygen atomic ratio is 2:1, analogous to that of water, and is represented by the empirical formula (where ''m'' and ''n'' ...
s, 1%
protein Proteins are large biomolecules and macromolecules that comprise one or more long chains of amino acid residue (biochemistry), residues. Proteins perform a vast array of functions within organisms, including Enzyme catalysis, catalysing metab ...
, and 0.5% fat (table). In a reference amount, raw cultivated blackberries supply 43
calorie The calorie is a unit of energy that originated from the caloric theory of heat. The large calorie, food calorie, dietary calorie, kilocalorie, or kilogram calorie is defined as the amount of heat needed to raise the temperature of one liter o ...
s and rich contents (20% or more of the
Daily Value In the U.S. and Canada, the Reference Daily Intake (RDI) is used in nutrition labeling on food and dietary supplement products to indicate the daily intake level of a nutrient that is considered to be sufficient to meet the requirements of 97â ...
(DV) of
dietary fiber Dietary fiber (fibre in English in the Commonwealth of Nations, Commonwealth English) or roughage is the portion of plant-derived food that cannot be completely broken down by human digestive enzymes. Dietary fibers are diverse in chemical co ...
,
manganese Manganese is a chemical element; it has Symbol (chemistry), symbol Mn and atomic number 25. It is a hard, brittle, silvery metal, often found in minerals in combination with iron. Manganese was first isolated in the 1770s. It is a transition m ...
(31% DV),
vitamin C Vitamin C (also known as ascorbic acid and ascorbate) is a water-soluble vitamin found in citrus and other fruits, berries and vegetables. It is also a generic prescription medication and in some countries is sold as a non-prescription di ...
(25% DV), and
vitamin K Vitamin K is a family of structurally similar, fat-soluble vitamers found in foods and marketed as dietary supplements. The human body requires vitamin K for post-translational modification, post-synthesis modification of certain proteins ...
(19% DV) (table).


Seed composition

Blackberries contain numerous large seeds that consumers do not always prefer. The seeds contain oil rich in
omega-3 Omega−3 fatty acids, also called omega−3 oils, ω−3 fatty acids or ''n''−3 fatty acids, are polyunsaturated fatty acids (PUFAs) characterized by the presence of a double bond three atoms away from the terminal methyl group in their ...
( alpha-linolenic acid) and omega-6 (
linoleic acid Linoleic acid (LA) is an organic compound with the formula . Both alkene groups () are ''cis''. It is a fatty acid sometimes denoted 18:2 (n−6) or 18:2 ''cis''-9,12. A linoleate is a salt or ester of this acid. Linoleic acid is a polyunsat ...
) fats as well as
protein Proteins are large biomolecules and macromolecules that comprise one or more long chains of amino acid residue (biochemistry), residues. Proteins perform a vast array of functions within organisms, including Enzyme catalysis, catalysing metab ...
, dietary fiber,
carotenoid Carotenoids () are yellow, orange, and red organic pigments that are produced by plants and algae, as well as several bacteria, archaea, and fungi. Carotenoids give the characteristic color to pumpkins, carrots, parsnips, corn, tomatoes, cana ...
s,
ellagitannins image:Castalagin.svg, 130px, Castalagin is a representative ellagitannin, characterized by coupled gallic acid substituents The ellagitannins are a diverse class of hydrolyzable tannins, a type of polyphenol formed primarily from the oxidative link ...
, and
ellagic acid Ellagic acid is a polyphenol found in numerous fruits and vegetables. It is the dilactone of hexahydroxydiphenic acid. Name The name comes from the French term ''acide ellagique'', from the word ''galle'' spelled backward because it can be o ...
.


Culinary use

The ripe fruit is commonly used in desserts, jams, jelly, wine, and
liqueur A liqueur ( , ; ) is an alcoholic drink composed of Liquor, spirits (often rectified spirit) and additional flavorings such as sugar, fruits, herbs, and spices. Often served with or after dessert, they are typically heavily sweetened and un-age ...
s. It may be mixed with other berries and fruits for pies and
crumble A crumble (British English) or crisp (American English) is a dessert with a crumbly cake-like topping, sometimes with oats, baked over a fruit filling. Apple and rhubarb are two popular varieties. Savoury fillings such as meat, cheese or vege ...
s.


Phytochemical research

Blackberries contain numerous
phytochemicals Phytochemicals are naturally-occurring chemicals present in or extract, extracted from plants. Some phytochemicals are nutrients for the plant, while others are metabolites produced to enhance plant survivability and reproduction. The field ...
including
polyphenols Polyphenols () are a large family of naturally occurring phenols. They are abundant in plants and structurally diverse. Polyphenols include phenolic acids, flavonoids, tannic acid, and ellagitannin, some of which have been used historically as ...
,
flavonoids Flavonoids (or bioflavonoids; from the Latin word ''flavus'', meaning yellow, their color in nature) are a class of polyphenolic secondary metabolites found in plants, and thus commonly consumed in the diets of humans. Chemically, flavonoids ...
,
anthocyanin Anthocyanins (), also called anthocyans, are solubility, water-soluble vacuole, vacuolar pigments that, depending on their pH, may appear red, purple, blue, or black. In 1835, the German pharmacist Ludwig Clamor Marquart named a chemical compou ...
s,
salicylic acid Salicylic acid is an organic compound with the formula HOC6H4COOH. A colorless (or white), bitter-tasting solid, it is a precursor to and a active metabolite, metabolite of acetylsalicylic acid (aspirin). It is a plant hormone, and has been lis ...
,
ellagic acid Ellagic acid is a polyphenol found in numerous fruits and vegetables. It is the dilactone of hexahydroxydiphenic acid. Name The name comes from the French term ''acide ellagique'', from the word ''galle'' spelled backward because it can be o ...
, and
fiber Fiber (spelled fibre in British English; from ) is a natural or artificial substance that is significantly longer than it is wide. Fibers are often used in the manufacture of other materials. The strongest engineering materials often inco ...
. Anthocyanins in blackberries are responsible for their rich dark color. One report placed blackberries at the top of more than 1,000 polyphenol-rich foods consumed in the United States, but this concept of a health benefit from consuming dark-colored foods like blackberries remains scientifically unverified and not accepted for
health claim A health claim on a food label and in food marketing is a claim by a manufacturer of food products that their food will reduce the risk of developing a disease or condition. For example, it is claimed by the manufacturers of oat cereals that oa ...
s on food labels.


Historical uses

One of the earliest known instances of blackberry consumption comes from the remains of the
Haraldskær Woman The Haraldskær Woman (or Haraldskjaer Woman) is the name given to a bog body of a woman preserved in a bog in Jutland, Denmark, and dating from about 490 BC (pre-Roman Iron Age). Workers found the body in 1835 while excavating peat on the Harald ...
, the naturally preserved
bog body A bog body is a human cadaver that has been naturally mummified in a peat bog. Such bodies, sometimes known as bog people, are both geographically and chronologically widespread, having been dated to between 8000 BC and the Second World War. Fi ...
of a Danish woman dating from approximately 2,500 years ago. Forensic evidence found blackberries in her stomach contents, among other foods. The use of blackberries to make wines and cordials was documented in the London Pharmacopoeia in 1696. In the culinary world, blackberries have long been used alongside other fruits to make pies, jellies, and jams. Blackberry plants were used for
traditional medicine Traditional medicine (also known as indigenous medicine or folk medicine) refers to the knowledge, skills, and practices rooted in the cultural beliefs of various societies, especially Indigenous groups, used for maintaining health and treatin ...
by Greeks, other European peoples, and aboriginal Americans. A 1771 document described brewing blackberry leaves, stem, and bark for
stomach ulcers Peptic ulcer disease is when the inner part of the stomach's gastric mucosa (lining of the stomach), the first part of the small intestine, or sometimes the lower esophagus, gets damaged. An ulcer in the stomach is called a gastric ulcer, while ...
. Blackberry fruit, leaves, and stems have been used to dye fabrics and hair. Native Americans have even been known to use the stems to make rope. The shrubs have also been used for barriers around buildings, crops, and livestock. The wild plants have sharp, thick prickles, which offer some protection against enemies and large animals.


In culture

Folklore in the United Kingdom and Ireland tells us that blackberries should not be picked after Old Michaelmas Day (11 October) as the
devil A devil is the mythical personification of evil as it is conceived in various cultures and religious traditions. It is seen as the objectification of a hostile and destructive force. Jeffrey Burton Russell states that the different conce ...
(or a
Púca The púca ( Irish for ''spirit/ghost''; plural púcaí), puca (Old English for ''goblin''), also pwca, pookah, phouka, and puck, is a creature of Celtic, English, and Channel Islands folklore. Considered to be bringers both of good and bad f ...
) has made them unfit to eat by stepping, spitting or fouling on them. There is some value in this legend as autumn's wetter and cooler weather often allows the fruit to become infected by various
mold A mold () or mould () is one of the structures that certain fungus, fungi can form. The dust-like, colored appearance of molds is due to the formation of Spore#Fungi, spores containing Secondary metabolite#Fungal secondary metabolites, fungal ...
s such as ''
Botryotinia ''Botryotinia'' is a genus of ascomycete fungi causing several plant diseases. The anamorphs of ''Botryotinia'' are mostly included in the " imperfect fungi" genus ''Botrytis''. The genus contains 22 species and one hybrid. Plant diseases caus ...
'' which give the fruit an unpleasant look and may be toxic. According to some traditions, a blackberry's deep purple color represents Jesus's blood and the
crown of thorns According to the New Testament, a woven crown of thorns ( or ) was placed on the head of Jesus during the Passion of Jesus, events leading up to his crucifixion of Jesus, crucifixion. It was one of the Arma Christi, instruments of the Passion, e ...
was made of brambles, although other thorny plants, such as ''
Crataegus ''Crataegus'' (), commonly called hawthorn, quickthorn, thornapple, Voss, E. G. 1985. ''Michigan Flora: A guide to the identification and occurrence of the native and naturalized seed-plants of the state. Part II: Dicots (Saururaceae–Cornacea ...
'' (hawthorn) and ''
Euphorbia milii ''Euphorbia milii'', the crown-of-thorns, Christ plant or Christ's thorn, is a species of flowering plant in the spurge family, Euphorbiaceae, native to Madagascar. The specifies commemorates Baron Milius, once Governor of Réunion, who introduce ...
'' (crown of thorns plant), have been proposed as the material for the crown.


See also

*
Rubus plicatus ''Rubus plicatus'' is a species of blackberry native to Europe. Taxonomy The type specimen of the genus ''Rubus'' was selected from among Linnaeus's specimens by Nathaniel Lord Britton and Addison Brown in 1913 as a herbarium specimen labelled ' ...
, a common European species of blackberry * Elmleaf blackberry, another common European species of blackberry * Pacific blackberry, a North American species of blackberry *
Rubus fruticosus ''Rubus fruticosus'' L. is the ambiguous name of a European blackberry species in the genus ''Rubus'' (part of the rose family). The name has been interpreted in several ways: *The species represented by the type specimen of ''Rubus fruticosu ...
, an ambiguous name used by
Carl Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...
that applied to multiple species *
Mulberry ''Morus'', a genus of flowering plants in the family Moraceae, consists of 19 species of deciduous trees commonly known as mulberries, growing wild and under cultivation in many temperate world regions. Generally, the genus has 64 subordinat ...
, similar fruit appearance, but a tree rather than a
bramble ''Rubus'' is a large and diverse genus of flowering plants in the rose family, Rosaceae, subfamily Rosoideae, most commonly known as brambles. Fruits of various species are known as raspberries, blackberries, dewberries, and bristleberries. I ...
* Redberry mite, a common pest of North American blackberry crops


References


Further reading

*


External links

* {{Commons-inline Berries Medicinal plants Plant common names Rubus Crown of thorns Plants used in Native American cuisine