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''Bittacus'' is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of hangingflies in the
order Order, ORDER or Orders may refer to: * A socio-political or established or existing order, e.g. World order, Ancien Regime, Pax Britannica * Categorization, the process in which ideas and objects are recognized, differentiated, and understood ...
Mecoptera Mecoptera (from the Greek language, Greek: ''mecos'' = "long", ''ptera'' = "wings") is an Order (biology), order of insects in the superorder Holometabola with about six hundred species in nine Family (biology), families worldwide. Mecopterans a ...
. Members of the genus have a
cosmopolitan distribution In biogeography, a cosmopolitan distribution is the range of a taxon that extends across most or all of the surface of the Earth, in appropriate habitats; most cosmopolitan species are known to be highly adaptable to a range of climatic and en ...
. The genus has existed since at least the earliest Late Cretaceous. ''Bittacus'' is considered "grossly
paraphyletic Paraphyly is a taxonomic term describing a grouping that consists of the grouping's last common ancestor and some but not all of its descendant lineages. The grouping is said to be paraphyletic ''with respect to'' the excluded subgroups. In co ...
" and serves as a catch-all for many distantly related species of hangingflies.


Characteristics

Members of this genus have long legs, the front pair of which are modified for grasping and are used to hang from vegetation. The other two pairs have tarsal claws and are modified for catching
prey Predation is a biological interaction in which one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not ki ...
. There are two pairs of equal sized, membranous
wings A wing is a type of fin that produces both lift and drag while moving through air. Wings are defined by two shape characteristics, an airfoil section and a planform. Wing efficiency is expressed as lift-to-drag ratio, which compares the bene ...
with dark mottling. The mouthparts are modified for chewing. The insects superficially resemble
crane flies A crane fly is any member of the dipteran superfamily Tipuloidea, which contains the living families Cylindrotomidae, Limoniidae, Pediciidae and Tipulidae, as well as several extinct families. "Winter crane flies", members of the family T ...
.


Species

The following species are listed in the ''World Checklist of Extant Mecoptera Species'': *'' Bittacus aequalis'' Navás, 1914 – Kenya *'' Bittacus africanus'' Esben-Petersen, 1915 – Zaire *'' Bittacus alluaudi'' Navás, 1914 – Kenya, Tanzania, Ethiopia *'' Bittacus andinus'' Londt and Byers, 1974 – Bolivia, Peru *'' Bittacus angrensis'' Souza Lopes and Mangabeira, 1942 – Brazil *''
Bittacus annae ''Bittacus'' is a genus of hangingflies in the order Mecoptera. Members of the genus have a cosmopolitan distribution. The genus has existed since at least the earliest Late Cretaceous. ''Bittacus'' is considered "grossly paraphyletic" and serves ...
'' Londt, 1972 – South Africa *'' Bittacus appendiculatus'' Esben-Petersen, 1927 – China (Yunnan) *'' Bittacus armatus'' Tjeder, 1956 – Zimbabwe, South Africa *'' Bittacus banksi'' Esben-Petersen, 1915 – Costa Rica, El Salvador, Guatemala, Honduras, Mexico, Panama *''
Bittacus berlandi ''Bittacus'' is a genus of hangingflies in the order Mecoptera. Members of the genus have a cosmopolitan distribution. The genus has existed since at least the earliest Late Cretaceous. ''Bittacus'' is considered "grossly paraphyletic" and serves ...
'' Capra, 1939 – Tanzania, Kenya *''
Bittacus bicornis ''Bittacus'' is a genus of hangingflies in the Order (biology), order Mecoptera. Members of the genus have a cosmopolitan distribution. The genus has existed since at least the earliest Late Cretaceous. ''Bittacus'' is considered "grossly Paraphy ...
'' Londt, 1993 – South Africa *''
Bittacus boraceiensis ''Bittacus'' is a genus of hangingflies in the order Mecoptera. Members of the genus have a cosmopolitan distribution. The genus has existed since at least the earliest Late Cretaceous. ''Bittacus'' is considered "grossly paraphyletic" and serves ...
'' -Morgante, 1967 – Brazil (São Paulo) *''
Bittacus boranicus ''Bittacus'' is a genus of hangingflies in the order Mecoptera. Members of the genus have a cosmopolitan distribution. The genus has existed since at least the earliest Late Cretaceous. ''Bittacus'' is considered "grossly paraphyletic" and serves ...
'' Capra, 1939 – Ethiopia *''
Bittacus brasiliensis ''Bittacus'' is a genus of hangingflies in the order Mecoptera. Members of the genus have a cosmopolitan distribution. The genus has existed since at least the earliest Late Cretaceous. ''Bittacus'' is considered "grossly paraphyletic" and serves ...
'' Klug, 1838 – Argentina, Brazil *''
Bittacus brunneus ''Bittacus'' is a genus of hangingflies in the order Mecoptera. Members of the genus have a cosmopolitan distribution. The genus has existed since at least the earliest Late Cretaceous. ''Bittacus'' is considered "grossly paraphyletic" and serves ...
'' Esben-Petersen, 1927 – Paraguay *''
Bittacus bullatus ''Bittacus'' is a genus of hangingflies in the order Mecoptera. Members of the genus have a cosmopolitan distribution. The genus has existed since at least the earliest Late Cretaceous. ''Bittacus'' is considered "grossly paraphyletic" and serve ...
'' Londt, 1972 – South Africa *''
Bittacus burgeoni ''Bittacus'' is a genus of hangingflies in the order Mecoptera. Members of the genus have a cosmopolitan distribution. The genus has existed since at least the earliest Late Cretaceous. ''Bittacus'' is considered "grossly paraphyletic" and serves ...
'' Navás, 1930 – Zaire *''
Bittacus burmanus ''Bittacus'' is a genus of hangingflies in the order Mecoptera. Members of the genus have a cosmopolitan distribution. The genus has existed since at least the earliest Late Cretaceous. ''Bittacus'' is considered "grossly paraphyletic" and serves ...
'' Tjeder, 1974 – Burma *''
Bittacus byersi ''Bittacus'' is a genus of hangingflies in the order Mecoptera. Members of the genus have a cosmopolitan distribution. The genus has existed since at least the earliest Late Cretaceous. ''Bittacus'' is considered "grossly paraphyletic" and serves ...
'' Londt, 1972 – South Africa *'' Bittacus capensis'' (Thunberg), 1784 – South Africa *'' Bittacus caprai'' Londt, 1972 – Zambia, Zimbabwe *'' Bittacus carpenteri'' Cheng, 1957 – China (Sichuan) *'' Bittacus chevalieri'' (Navás), 1908 – Chad, Mali, Senegal, French Somaliland *'' Bittacus chilensis'' Klug, 1838 – Chile *'' Bittacus chlorostigma'' MacLachlan, 1881 – United States (California, Oregon) *'' Bittacus chujoi'' Issiki and Cheng, 1947 – Taiwan *'' Bittacus cirratus'' Tjeder, 1956 – China (Manchuria,
Jiangxi ; Gan: ) , translit_lang1_type2 = , translit_lang1_info2 = , translit_lang1_type3 = , translit_lang1_info3 = , image_map = Jiangxi in China (+all claims hatched).svg , mapsize = 275px , map_caption = Location ...
) *'' Bittacus contumax'' Tjeder, 1956 – China *'' Bittacus coreanus'' Issiki, 1929 – China (
Jiangsu Jiangsu is a coastal Provinces of the People's Republic of China, province in East China. It is one of the leading provinces in finance, education, technology, and tourism, with its capital in Nanjing. Jiangsu is the List of Chinese administra ...
), Korea *'' Bittacus cottrelli'' Londt, 1972 – South Africa *'' Bittacus discors'' Navás, 1914 – Kenya, Somalia *'' Bittacus disternum'' Byers, 1996 – Costa Rica *'' Bittacus diversinervis'' Souza Lopes and Mangabeira, 1942 – Brazil *'' Bittacus elisabethae'' Navás, 1930 – Zaire *'' Bittacus eremus'' Lambkin, 1988 – Australia *'' Bittacus erythrostigma'' Byers, 1975 – Uganda, Zaire *'' Bittacus femoralis'' Klug, 1838 – Brazil *'' Bittacus flavescens'' Klug, 1838 – Brazil, Venezuela *'' Bittacus formosanus'' Issiki, 1927 – Taiwan *'' Bittacus fritzi'' Williner, 1990 – Argentina *'' Bittacus fumosus'' Esben-Petersen, 1913 – Kenya, Zimbabwe, Tanzania, Zambia *'' Bittacus geniculatus'' Erichson, 1848 – Brazil, Guyana *'' Bittacus golbachi'' Williner, 1990 – Argentina *'' Bittacus gressitti'' Cheng, 1957 – China (
Guangdong ) means "wide" or "vast", and has been associated with the region since the creation of Guang Prefecture in AD 226. The name "''Guang''" ultimately came from Guangxin ( zh, labels=no, first=t, t= , s=广信), an outpost established in Han dynasty ...
) *'' Bittacus hageni'' Brauer, 1860 – Austria, Belgium, France *'' Bittacus henryi'' Kimmins, 1928 – Sri Lanka *'' Bittacus homburgerae'' Navás, 1933 – French Guinea *'' Bittacus indicus'' Walker 1853 – India *'' Bittacus insularis'' Esben-Petersen 1915 – Sri Lanka *'' Bittacus issikii'' Miyamoto 1979 – Japan *'' Bittacus italicus'' (Müller) 1766 -Belgium, Bosnia, Germany, Poland, Romania, Spain, Switzerland, Ukraine *'' Bittacus kagoshimaensis'' Issiki 1929 -Japan *'' Bittacus kimminsi'' Tjeder 1956 – South Africa *'' Bittacus kunenensis'' Wood 1933 – Namibia *'' Bittacus laevipes'' Navás 1909 – Japan *'' Bittacus latipennis'' Gerstaecker 1885 – India *'' Bittacus leptocaudus'' Byers 1965 – Thailand *'' Bittacus leptocercus'' Navás 1934 – Tanzania *'' Bittacus lineatus'' Navás 1914 – Kenya *'' Bittacus livingstonei'' Londt 1981 – Malawi *'' Bittacus maculatus'' Issiki 1927 – Taiwan *'' Bittacus maculosus'' Byers 1965 – Brazil, Trinidad *'' Bittacus malaisei'' Tjeder 1974 – Burma *'' Bittacus marginatus'' Miyake 1913 – Japan *'' Bittacus mastrillii'' Navás 1913 – Japan *'' Bittacus mexicanus'' Klug 1838 – Mexico (Oaxaca, San Luis Potosí) *'' Bittacus milleri'' Londt 1978 – South Africa *'' Bittacus montanus'' Weele 1910 – Angola, Cameroon, Kenya, Malawi, Zimbabwe, Rwanda, Tanzania, Uganda, Zaire, Zambia *''
Bittacus moschinus ''Bittacus'' is a genus of hangingflies in the order Mecoptera. Members of the genus have a cosmopolitan distribution. The genus has existed since at least the earliest Late Cretaceous. ''Bittacus'' is considered "grossly paraphyletic" and serves ...
'' Navás 1914 – Malawi, Tanzania, Zambia *'' Bittacus natalensis'' Wood 1933 – Mozambique, Malawi, Zimbabwe, South Africa *'' Bittacus nebulosus'' Klug 1838 – Mozambique, South Africa *'' Bittacus nipponicus'' Navás 1909 – Japan *'' Bittacus nodosus'' Rust and Byers 1976 – India, Pakistan *'' Bittacus occidentis'' Walker 1853 – United States *'' Bittacus omega'' Morgante 1967 – Brazil *'' Bittacus oreinus'' Navás 1914 – Ethiopia *'' Bittacus panamensis'' Byers 1958 – Costa Rica, Panama, Venezuela *'' Bittacus peninsularis'' Byers 1996 – Mexico (Baja California Sur) *'' Bittacus peringueyi'' Esben-Petersen 1913 – South Africa *'' Bittacus peterseni'' Kimmins 1938 – South Africa *'' Bittacus pieli'' Navás 1935 – China (Jiangxi) *'' Bittacus pignatellii'' Navás 1932 – Panama *'' Bittacus pilicornis'' Westwood 1846 – Canada, United States *'' Bittacus pinguipalpi'' Wood 1933 – Namibia *'' Bittacus pintoi'' Souza Lopes and Mangabeira 1942 – Brazil *'' Bittacus planus'' Cheng 1949 – China (
Shaanxi Shaanxi is a Provinces of China, province in north Northwestern China. It borders the province-level divisions of Inner Mongolia to the north; Shanxi and Henan to the east; Hubei, Chongqing, and Sichuan to the south; and Gansu and Ningxia to t ...
) *'' Bittacus pobeguini'' (Navás) 1908 – Zaire, Ghana, Ivory Coast, Nigeria, Uganda *'' Bittacus pollex'' Byers and Roggero 1992 – Panama *'' Bittacus punctiger'' Westwood 1846 – United States *'' Bittacus rossi'' Londt 1977 – Zaire *'' Bittacus saigusai'' Miyamoto 1984 – Japan *'' Bittacus schoutedeni'' Esben-Petersen 1913 – Zaire *'' Bittacus selysi'' Esben-Petersen 1917 – South Africa *'' Bittacus sinensis'' Walker 1853 – China (
Zhejiang ) , translit_lang1_type2 = , translit_lang1_info2 = ( Hangzhounese) ( Ningbonese) (Wenzhounese) , image_skyline = 玉甑峰全貌 - panoramio.jpg , image_caption = View of the Yandang Mountains , image_map = Zhejiang i ...
, Jiangsu), Korea, Japan *'' Bittacus sinicus'' Issiki 1931 – China (Sichuan) *'' Bittacus sjostedti'' Weele 1910 – Kenya, Tanzania *'' Bittacus smithersi'' Londt 1972 – South Africa *'' Bittacus sobrinus'' Tjeder 1956 – South Africa *'' Bittacus sonani'' Issiki 1929 – Taiwan *'' Bittacus spatulatus'' Byers 1996 – Costa Rica, Nicaragua *'' Bittacus stanleyi'' Byers 1968 – Zaire, Malawi *'' Bittacus stigmaterus'' Say 1823 – United States *'' Bittacus striatus'' Issiki 1927 – Taiwan *''
Bittacus strigosus ''Bittacus strigosus'', the striped scorpionfly, is a species of hangingfly in the family Bittacidae, living in North America. References External links

* Bittacus Articles created by Qbugbot Insects described in 1861 Insects of Nor ...
'' Hagen 1861 – United States *'' Bittacus sylvaticus'' Byers 1996 – Mexico *'' Bittacus takaoensis'' Miyake 1913 – Japan *'' Bittacus taraiensis'' Penny 1969 – India *'' Bittacus tarsalis'' Miyamoto 1984 – Japan *'' Bittacus testaceus'' Klug 1838 – Zimbabwe, South Africa *'' Bittacus texanus'' Banks 1908 – United States (Kansas, New Mexico, Texas) *'' Bittacus tienmushana'' Cheng 1957 – China (Zhejiang) *'' Bittacus tjederi'' Londt 1970 – South Africa *'' Bittacus triangularis'' Issiki 1929 – Korea, Manchuria *'' Bittacus tuxeni'' Byers 1975 – Guinea, Malawi *'' Bittacus ussuriensis'' Plutenko 1985 – Far Eastern Russia *'' Bittacus vexilliferus'' Byers 1970 – China (Sichuan) *'' Bittacus vumbanus'' Smithers 1960 – Zimbabwe *'' Bittacus wahlbergi'' Londt 1972 – South Africa *'' Bittacus walkeri'' Esben-Petersen 1915 – South Africa *'' Bittacus weelei'' Esben-Petersen 1913 – Angola, Ethiopia, Ghana, Malawi, Nigeria, Tanzania, Uganda, Zaire, Zambia *'' Bittacus zambezinus'' Navás 1931 – Mozambique, Malawi, Zimbabwe, South Africa *'' Bittacus zelichi'' Williner 1990 – Argentina *'' Bittacus zulu'' Londt 1972 – South Africa


Extinct species

* †'' Bittacus lepiduscretaceus'' Li et al. 2018
Burmese amber Burmese amber, also known as Burmite or Kachin amber, is amber from the Hukawng Valley in northern Myanmar. The amber is dated to around 100 million years ago, during the latest Albian to earliest Cenomanian ages of the mid-Cretaceous period. Th ...
, Myanmar,
Cenomanian The Cenomanian is, in the International Commission on Stratigraphy's (ICS) geological timescale, the oldest or earliest age (geology), age of the Late Cretaceous epoch (geology), Epoch or the lowest stage (stratigraphy), stage of the Upper Cretace ...
* †'' Bittacus succinus'' Carpenter 1954
Baltic amber Baltic amber or succinite is amber from the Baltic region, home of its largest known deposits. It was produced sometime during the Eocene epoch, but exactly when is controversial. It has been estimated that this forested region provided the re ...
,
Eocene The Eocene ( ) is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (Ma). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene'' comes ...


References

{{Authority control Hangingflies Insects described in 1805