''Bisticeratops'' (meaning "

Bisti/De-Na-Zin Wilderness The Bisti/De-Na-Zin Wilderness is a wilderness area located in San Juan County in the U.S. state of New Mexico. Established in 1984, the Wilderness is a desolate area of steeply eroded badlands managed by the Bureau of Land Management, ex ...

horned face") is a

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

chasmosaurine Chasmosaurinae is a subfamily of ceratopsid dinosaurs. They were one of the most successful groups of herbivores of their time. Chasmosaurines appeared in the early Campanian, and became extinct, along with all other non- avian dinosaurs, during ...

ceratopsia Ceratopsia or Ceratopia ( or ; Ancient Greek, Greek: "horned faces") is a group of herbivore, herbivorous, beaked dinosaurs that thrived in what are now North America, Asia and Europe, during the Cretaceous Period (geology), Period, although ance ...

n from outcrops of the

Campanian The Campanian is the fifth of six ages of the Late Cretaceous epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campa ...

age

Kirtland Formation The Kirtland Formation (originally the Kirtland Shale) is a Sedimentary rock, sedimentary geological formation. Description The Kirtland Formation is the product of alluvial muds and overbank sand deposits from the many channels draining the ...

found in the Bisti/De-Na-Zin Wilderness in northwestern

New Mexico New Mexico is a state in the Southwestern United States, Southwestern region of the United States. It is one of the Mountain States of the southern Rocky Mountains, sharing the Four Corners region with Utah, Colorado, and Arizona. It also ...

, United States. The type and only

species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...

is ''B. froeseorum'', known from a nearly complete skull.

Discovery and naming

Geological map of the southeast San Juan Basin

The ''Bisticeratops''

holotype A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...

specimen, NMMNH P-50000, was discovered in 1975 in layers of the

(Farmington Member) in the

area of

San Juan Basin The San Juan Basin is a geologic structural basin located near the Four Corners region of the Southwestern United States. The basin covers 7,500 square miles and resides in northwestern New Mexico, southwestern Colorado, and parts of Utah a ...

, northwestern

, United States, which dates to the late

age of the late Cretaceous period. The specimen consists of a mostly complete

skull The skull, or cranium, is typically a bony enclosure around the brain of a vertebrate. In some fish, and amphibians, the skull is of cartilage. The skull is at the head end of the vertebrate. In the human, the skull comprises two prominent ...

, missing the parietal, left

squamosal The squamosal is a skull bone found in most reptiles, amphibians, and birds. In fishes, it is also called the pterotic bone. In most tetrapods, the squamosal and quadratojugal bones form the cheek series of the skull. The bone forms an ancestra ...

, postorbital horncores (only the cast of the right postorbital horncore is preserved), left

jugal The jugal is a skull bone found in most reptiles, amphibians and birds. In mammals, the jugal is often called the malar or zygomatic. It is connected to the quadratojugal and maxilla, as well as other bones, which may vary by species. Anatomy ...

, left quadrate,

predentary Ornithischia () is an extinct clade of mainly herbivorous dinosaurs characterized by a pelvic structure superficially similar to that of birds. The name ''Ornithischia'', or "bird-hipped", reflects this similarity and is derived from the Ancient ...

, and both

dentaries In jawed vertebrates, the mandible (from the Latin ''mandibula'', 'for chewing'), lower jaw, or jawbone is a bone that makes up the lowerand typically more mobilecomponent of the mouth (the upper jaw being known as the maxilla). The jawbone i ...

. The holotype specimen was originally thought to be a specimen of ''

Pentaceratops ''Pentaceratops'' ("five-horned face") is a genus of herbivorous ceratopsid dinosaur from the late Cretaceous Period of what is now North America. Fossils of this animal were first discovered in 1921, but the genus was named in 1923 when its ty ...

'', despite being two million years younger than other specimens of that genus. When the description of ''

Sierraceratops ''Sierraceratops'' (meaning " Sierra horned face") is a genus of chasmosaurine ceratopsian from the Late Cretaceous Hall Lake Formation of New Mexico, United States. The genus contains a single species, ''Sierraceratops turneri'', known from a p ...

'' was initially published online in 2021, the name "Bisticeratops froeseorum" was leaked in a cladogram, but edited out when physically published in early 2022. ''Bisticeratops froeseorum'' was formally described by Dalman ''et al''. in a separate paper published the same year. The generic name, "''Bisticeratops''", combines "Bisti", a reference to the location where the holotype was discovered, with the

Greek Greek may refer to: Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group *Greek language, a branch of the Indo-European language family **Proto-Greek language, the assumed last common ancestor of all kno ...

''keras'' (meaning "horn"), and ''ops'' (meaning "face"). The

specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...

honors

Edgar Froese Edgar Willmar Froese (; 6 June 1944 – 20 January 2015) was a German musical artist and electronic music pioneer, best known for founding the electronic music group Tangerine Dream in 1967. Froese was the only continuous member of the gro ...

, the founder of the band

Tangerine Dream Tangerine Dream is a German electronic music band founded in 1967 by Edgar Froese. The group has seen many personnel changes over the years, with Froese the only constant member until his death in January 2015. The best-known lineup of the grou ...

, and his son

Jerome Jerome (; ; ; – 30 September 420), also known as Jerome of Stridon, was an early Christian presbyter, priest, Confessor of the Faith, confessor, theologian, translator, and historian; he is commonly known as Saint Jerome. He is best known ...

, former member of Tangerine Dream and founder of the band Loom. Their music is said to have "provided through the years" and inspired the description.

Classification

Using the same

phylogenetic analyses In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as Computational phylogenetics, phylogenetic inference. It infers the relationship among organ ...

used to find the position of ''

'', Dalman ''et al''. (2022) recovered ''Bisticeratops'' as a sister taxon to an unnamed ceratopsian from the Almond Formation. The

cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not s ...

below displays the results of the

phylogenetic In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical dat ...

analyses by Dalman, Jasinski, & Lucas.

Paleobiology

The holotype preserves bite marks from tyrannosaurids, some of which show signs of healing.

Paleoenvironment

The fossil remains of ''Bisticeratops'' were recovered from the Farmington Member of the

. This area represents the product of alluvial muds and overbank sand deposits from the many channels draining the coastal plain that existed on the inland seashore of North America during the late Cretaceous period. No other dinosaurs have been described from the Farmington Member, but taxa from other sub-units include

kritosaurin Saurolophinae is a subfamily of hadrosaurid dinosaurs. It has since the mid-20th century generally been called the Hadrosaurinae, a group of largely non-crested hadrosaurs related to the crested sub-family Lambeosaurinae. However, the name Hadros ...

and

lambeosaurine Lambeosauridae /ˌlæmbiəˈsɔːraɪniː/ (meaning 'lambe's lizards') is an extinct group of crested hadrosauroid dinosaurs. Description Size Uncertainty surrounds the size of lambeosaurs from the European continent. Hadrosaurs found there, a ...

hadrosaurs,

ankylosaurids Ankylosauridae () is a family of armored dinosaurs within Ankylosauria, and is the sister group to Nodosauridae. The oldest known ankylosaurids date to around 122 million years ago and went extinct 66 million years ago during the Cretaceous–Pal ...

, chasmosaurine ceratopsids,

pachycephalosaurids Pachycephalosauria (; from Greek παχυκεφαλόσαυρος for 'thick headed lizards') is a clade of ornithischian dinosaurs. Along with Ceratopsia, it makes up the clade Marginocephalia. With the exception of two species, most pachycepha ...

dromaeosaurids Dromaeosauridae () is a family of feathered coelurosaurian theropod dinosaurs. They were generally small to medium-sized feathered carnivores that flourished in the Cretaceous Period. The name Dromaeosauridae means 'running lizards', from Gree ...

, an indeterminate

ornithomimid Ornithomimidae (meaning "bird-mimics") is an extinct family of theropod dinosaurs which bore a superficial resemblance to modern ostriches. Ornithomimids were fast, omnivorous or herbivorous dinosaurs known mainly from the Late Cretaceous Period ...

tyrannosauroids Tyrannosauroidea (meaning 'tyrant lizard forms') is a superfamily (or clade) of coelurosaurian theropod dinosaurs that includes the family Tyrannosauridae as well as more basal relatives. Tyrannosauroids lived on the Laurasian supercontinent b ...

azhdarchids Azhdarchidae (from the Persian language, Persian word , , a dragon-like creature in Persian mythology) is a family (biology), family of pterosaurs known primarily from the Late Cretaceous Period, though an isolated vertebra apparently from an azh ...

crurotarsans Crurotarsi is a clade of archosauriform reptiles that includes crocodilians and stem-crocodilians and possibly bird-line archosaurs too if the extinct, crocodile-like phytosaurs are more distantly related to crocodiles than traditionally thought ...

turtles Turtles are reptiles of the order Testudines, characterized by a special shell developed mainly from their ribs. Modern turtles are divided into two major groups, the Pleurodira (side necked turtles) and Cryptodira (hidden necked turtle ...

, and

cartilaginous Cartilage is a resilient and smooth type of connective tissue. Semi-transparent and non-porous, it is usually covered by a tough and fibrous membrane called perichondrium. In tetrapods, it covers and protects the ends of long bones at the joints ...

and

bony fish Osteichthyes ( ; ), also known as osteichthyans or commonly referred to as the bony fish, is a Biodiversity, diverse clade of vertebrate animals that have endoskeletons primarily composed of bone tissue. They can be contrasted with the Chondricht ...

from the De-Na-Zin and Hunter Wash members.

References

{{Taxonbar, from=Q113534609 Chasmosaurinae Dinosaur genera Campanian dinosaurs Kirtland Formation Fossil taxa described in 2022 Dinosaurs of the United States