
A biennial plant is a
flowering plant
Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Ancient Greek, Greek words (; 'container, vessel') and (; 'seed'), meaning that the seeds are enclosed with ...
that, generally in a
temperate climate
In geography, the temperate climates of Earth occur in the middle latitudes (approximately 23.5° to 66.5° N/S of the Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ra ...
, takes two years to complete its
biological life cycle
In biology, a biological life cycle (or just life cycle when the biological context is clear) is a series of stages of the life of an organism, that begins as a zygote, often in an egg, and concludes as an adult that reproduces, producing an offsp ...
.
Background
In its first year, the biennial plant undergoes primary growth, during which its vegetative structures (leaves, stems, and roots) develop. Usually, the stem of the plant remains short and the leaves are low to the ground, forming a
rosette. After one year's growing season, the plant enters a period of
dormancy
Dormancy is a period in an organism's Biological life cycle, life cycle when growth, development, and (in animals) physical activity are temporarily stopped. This minimizes metabolism, metabolic activity and therefore helps an organism to conserv ...
for the colder months. Many biennials require a cold treatment, or
vernalization before they will flower. During the next
spring or
summer
Summer or summertime is the hottest and brightest of the four temperate seasons, occurring after spring and before autumn. At or centred on the summer solstice, daylight hours are the longest and darkness hours are the shortest, with day ...
, the stem of the biennial plant elongates greatly, or "bolts".
The plant then flowers, producing
fruit
In botany, a fruit is the seed-bearing structure in flowering plants (angiosperms) that is formed from the ovary after flowering.
Fruits are the means by which angiosperms disseminate their seeds. Edible fruits in particular have long propaga ...
s and
seed
In botany, a seed is a plant structure containing an embryo and stored nutrients in a protective coat called a ''testa''. More generally, the term "seed" means anything that can be Sowing, sown, which may include seed and husk or tuber. Seeds ...
s before it finally dies. There are far fewer biennials than either
perennial plant
In horticulture, the term perennial (''wikt:per-#Prefix, per-'' + ''wikt:-ennial#Suffix, -ennial'', "through the year") is used to differentiate a plant from shorter-lived annual plant, annuals and biennial plant, biennials. It has thus been d ...
s or
annual plant
An annual plant is a plant that completes its life cycle, from germination to the production of seeds, within one growing season, and then dies. Globally, 6% of all plant species and 15% of herbaceous plants (excluding trees and shrubs) are ...
s.
Biennials do not always follow a strict two-year life cycle: most plants in the wild can take three or more years to mature. Rosette leaf size has been found to predict when a plant may enter its second stage of flowering and seed production. Alternatively, under extreme climatic conditions, a biennial plant may complete its life cycle rapidly (e.g., in three months instead of two years).
This is quite common in vegetable or flower seedlings that were
vernalized before they were planted in the ground. This behavior leads to many normally biennial plants being treated as annuals in some areas. Conversely, an annual grown under extremely favorable conditions may have highly successful seed propagation, giving it the appearance of being biennial or perennial. Some short-lived perennials may appear to be biennial rather than perennial. True biennials flower only once, while many perennials will flower every year once mature.
Biennials grown for flowers, fruits, or seeds are grown for two years, whereas those grown for edible leaves or roots are harvested after one year—and are not kept a second year to run to seed.
Examples of biennial plants are members of the
onion
An onion (''Allium cepa'' , from Latin ), also known as the bulb onion or common onion, is a vegetable that is the most widely cultivated species of the genus '' Allium''. The shallot is a botanical variety of the onion which was classifie ...
family including
leek
A leek is a vegetable, a cultivar of ''Allium ampeloprasum'', the broadleaf wild leek (synonym (taxonomy), syn. ''Allium porrum''). The edible part of the plant is a bundle of Leaf sheath, leaf sheaths that is sometimes erroneously called a "s ...
,
[ some members of the ]cabbage
Cabbage, comprising several cultivars of '' Brassica oleracea'', is a leafy green, red (purple), or white (pale green) biennial plant grown as an annual vegetable crop for its dense-leaved heads. It is descended from the wild cabbage ( ''B.& ...
family,[ common mullein, ]parsley
Parsley, or garden parsley (''Petroselinum crispum''), is a species of flowering plant in the family Apiaceae that is native to Greece, Morocco and the former Yugoslavia. It has been introduced and naturalisation (biology), naturalized in Eur ...
, fennel,[ '' Lunaria'', silverbeet, black-eyed Susan, ]sweet William
Sweet William may refer to:
* '' Dianthus barbatus'', a species of flowering plant
* '' Mustelus antarcticus,'' a species of shark
* ''Sweet William'' (novel), a 1975 novel by Beryl Bainbridge
* ''Sweet William'' (film), a 1980 British drama fil ...
, colic weed, carrot
The carrot ('' Daucus carota'' subsp. ''sativus'') is a root vegetable, typically orange in colour, though heirloom variants including purple, black, red, white, and yellow cultivars exist, all of which are domesticated forms of the wild ...
,[ and some ]hollyhock
''Alcea'' is a genus of over 80 species of flowering plants in the mallow family Malvaceae, commonly known as the hollyhocks. They are native to Asia and Europe. The single species of hollyhock from the Americas, the Iliamna rivularis, streamban ...
s. Plant breeders have produced annual cultivars of several biennials that will flower the first year from seed, for example, foxglove
''Digitalis'' ( or ) is a genus of about 20 species of herbaceous perennial plants, shrubs, and biennials, commonly called foxgloves.
''Digitalis'' is native to Europe, Western Asia, and northwestern Africa. The flowers are tubular in sha ...
and stock
Stocks (also capital stock, or sometimes interchangeably, shares) consist of all the Share (finance), shares by which ownership of a corporation or company is divided. A single share of the stock means fractional ownership of the corporatio ...
.
See also
*
*
*
References
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