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Capsalidae Yamaguti, S. (1963) Systema Helminthum IV. Monogenea and Aspidocotylea. London-New York, Interscience Publishers. 699 pp. is a family of
monopisthocotylea The Monopisthocotylea are an inferior class of parasitic flatworms in the class Monogenea.WoRMS (2019). Monopisthocotylea. Accessed at: http://www.marinespecies.org/aphia.php?p=taxdetails&id=119219 on 2019-02-08 Yamaguti, S. (1963). Systema Helmi ...
n
monogenea Monogeneans, members of the class Monogenea, are a group of ectoparasitic flatworms commonly found on the skin, gills, or fins of fish. They have a direct lifecycle and do not require an intermediate host. Adults are hermaphrodites, meaning they ...
ns, which includes about 200
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
. The
monophyly In biological cladistics for the classification of organisms, monophyly is the condition of a taxonomic grouping being a clade – that is, a grouping of organisms which meets these criteria: # the grouping contains its own most recent comm ...
of the Capsalidae is supported by possession of accessory
sclerites A sclerite (Greek , ', meaning " hard") is a hardened body part. In various branches of biology the term is applied to various structures, but not as a rule to vertebrate anatomical features such as bones and teeth. Instead it refers most commonly ...
in the
haptor The haptor is the attachment organ of the monogeneans, a group of parasitic Platyhelminthes. The haptor is sometimes called opisthaptor (from ''opistho-'': behind) to emphasize that it is located in the posterior part of the body, and to diffe ...
(the posterior attachment organ), and was confirmed by
molecular phylogeny Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to ...
. Their
oncomiracidium An oncomiracidium is the ciliated and free-living larva of a monogenean, a type of parasitic flatworm commonly found on fish. It is similar to the miracidium of Trematoda Trematoda is a class of flatworms known as trematodes, and commonly as ...
(the free larva) is distinct from that of other families. Capsalids are
parasite Parasitism is a Symbiosis, close relationship between species, where one organism, the parasite, lives (at least some of the time) on or inside another organism, the Host (biology), host, causing it some harm, and is Adaptation, adapted str ...
on various organs of marine
fish A fish (: fish or fishes) is an aquatic animal, aquatic, Anamniotes, anamniotic, gill-bearing vertebrate animal with swimming fish fin, fins and craniate, a hard skull, but lacking limb (anatomy), limbs with digit (anatomy), digits. Fish can ...
(
teleosts Teleostei (; Greek ''teleios'' "complete" + ''osteon'' "bone"), members of which are known as teleosts (), is, by far, the largest group of ray-finned fishes (class Actinopterygii), with 96% of all extant species of fish. The Teleostei, which i ...
and
elasmobranchs Elasmobranchii () is a subclass of Chondrichthyes or cartilaginous fish, including modern sharks ( division Selachii), and batomorphs (division Batomorphi, including rays, skates, and sawfish). Members of this subclass are characterised by ha ...
), including skin,
fins A fin is a thin component or appendage attached to a larger body or structure. Fins typically function as foil (fluid mechanics), foils that produce lift (force), lift or thrust, or provide the ability to steer or stabilize motion while travelin ...
and
gills A gill () is a respiratory organ that many aquatic organisms use to extract dissolved oxygen from water and to excrete carbon dioxide. The gills of some species, such as hermit crabs, have adapted to allow respiration on land provided they are ...
. Several capsalid species, such a ''
Neobenedenia ''Neobenedenia'' is a genius of monopisthocotylean monogenean flatworm parasites. The most common species, ''Neobenedenia melleni'', causes problematic disease in public aquariums. It was named after ichthyologist Ida May Mellen (1877–1970) ...
'' spp. are pathogenic, especially on
mariculture Mariculture, sometimes called marine farming or marine aquaculture, is a branch of aquaculture involving the cultivation of marine organisms for food and other animal products, in seawater. Subsets of it include ( offshore mariculture), fish fa ...
d fish.


Included genera

Genera as recognized i
WorMs
are listed below. Recent molecular analyses have shown that several genera, which were defined on morphological characters, are not monophyletic. ''
Menziesia ''Rhododendron'' sect. ''Sciadorhodion'' is a subsection of Rhododendron subg. Azaleastrum, subgenus ''Azaleastrum'' in the genus ''Rhododendron''. It comprises 16 species of deciduous shrubss native to East Asia and North America. Taxonomy Plan ...
'' and ''
Nitzschia ''Nitzschia'' is a common pennate marine and freshwater diatom. In the scientific literature, this genus, named after Christian Ludwig Nitzsch, is sometimes referred to incorrectly as ''Nitzchia'', and it has many species described, which all hav ...
'' have their equivalent in the
botanical nomenclature Botanical nomenclature is the formal, scientific naming of plants. It is related to, but distinct from taxonomy (biology), taxonomy. Plant taxonomy is concerned with grouping and classifying plants; Botany, botanical nomenclature then provides na ...
: ''
Menziesia ''Rhododendron'' sect. ''Sciadorhodion'' is a subsection of Rhododendron subg. Azaleastrum, subgenus ''Azaleastrum'' in the genus ''Rhododendron''. It comprises 16 species of deciduous shrubss native to East Asia and North America. Taxonomy Plan ...
'' (a
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Ancient Greek, Greek words (; 'container, vessel') and (; 'seed'), meaning that the seeds are enclosed with ...
) and ''
Nitzschia ''Nitzschia'' is a common pennate marine and freshwater diatom. In the scientific literature, this genus, named after Christian Ludwig Nitzsch, is sometimes referred to incorrectly as ''Nitzchia'', and it has many species described, which all hav ...
'' (a
diatom A diatom (Neo-Latin ''diatoma'') is any member of a large group comprising several Genus, genera of algae, specifically microalgae, found in the oceans, waterways and soils of the world. Living diatoms make up a significant portion of Earth's B ...
). * ''
Allobenedenia ''Allobenedenia'' is a genus of monopisthocotylean monogeneans, included in the family Capsalidae.Yamaguti, S. (1963). Systema Helminthum Volume IV Monogenea and Aspidocotylea: John Wiley & Sons.WoRMS (2018). Allobenedenia Yamaguti, 1963. Accesse ...
'' Yamaguti, 1963 * ''
Alloencotyllabe ''Alloencotyllabe'' is a genus of parasitic monogenean in the family Capsalidae. It is monotypic, containing the sole species ''Alloencotyllabe caranxi''.WoRMS (2023)''Alloencotyllabe'' Khalil & Abdul-Salam, 1988 Accessed 2023-06-16. The specific ...
'' Khalil & Abdul-Salam, 1988 * ''
Allomegalocotyla ''Allomegalocotyla'' is a genus of monopisthocotylean monogeneans, included in the family Capsalidae.Yamaguti, S. (1963). Systema Helminthum Volume IV Monogenea and Aspidocotylea: John Wiley & Sons.WoRMS (2018). Allobenedenia Yamaguti, 1963. Acce ...
'' Yamaguti, 1963 * ''
Allometabenedeniella Capsalidae Yamaguti, S. (1963) Systema Helminthum IV. Monogenea and Aspidocotylea. London-New York, Interscience Publishers. 699 pp. is a family of monopisthocotylean monogeneans, which includes about 200 species. The monophyly of the Capsalida ...
'' Velasquez, 1982 * ''
Ancyrocotyle CapsalidaeSatyu Yamaguti, Yamaguti, S. (1963) Systema Helminthum IV. Monogenea and Aspidocotylea. London-New York, Interscience Publishers. 699 pp. is a family of monopisthocotylean monogeneans, which includes about 200 species. The monophyly o ...
'' Parona & Monticelli, 1903 * ''
Benedenia CapsalidaeSatyu Yamaguti, Yamaguti, S. (1963) Systema Helminthum IV. Monogenea and Aspidocotylea. London-New York, Interscience Publishers. 699 pp. is a family of monopisthocotylean monogeneans, which includes about 200 species. The monophyly o ...
'' Diesing, 1858 * ''
Benedeniella Capsalidae Yamaguti, S. (1963) Systema Helminthum IV. Monogenea and Aspidocotylea. London-New York, Interscience Publishers. 699 pp. is a family of monopisthocotylean monogeneans, which includes about 200 species. The monophyly of the Capsalida ...
'' Johnston, 1929 * ''
Branchobdella Capsalidae Yamaguti, S. (1963) Systema Helminthum IV. Monogenea and Aspidocotylea. London-New York, Interscience Publishers. 699 pp. is a family of monopisthocotylean monogeneans, which includes about 200 species. The monophyly of the Capsalida ...
'' Kearn, Whittington & Evans-Gowing, 2007 * '' Calicobenedenia''
Kritsky ''Kritsky'' is a genus of digeneans in the family Aporocotylidae or blood flukes.WoRMS (2018). Kritsky Orelis-Ribeiro & Bullard, 2016. Accessed at: http://www.marinespecies.org/aphia.php?p=taxdetails&id=871002 on 2018-09-23 The name of the genus ...
& Fennessy, 1999
* ''
Capsala Capsalidae Yamaguti, S. (1963) Systema Helminthum IV. Monogenea and Aspidocotylea. London-New York, Interscience Publishers. 699 pp. is a family of monopisthocotylean monogeneans, which includes about 200 species. The monophyly of the Capsalida ...
'' Bosc, 1811 * '' Capsaloides'' Price, 1936 * '' Dioncopseudobenedenia'' Yamaguti, 1965 * '' Encotyllabe'' Diesing, 1850 * ''
Entobdella CapsalidaeSatyu Yamaguti, Yamaguti, S. (1963) Systema Helminthum IV. Monogenea and Aspidocotylea. London-New York, Interscience Publishers. 699 pp. is a family of monopisthocotylean monogeneans, which includes about 200 species. The monophyly o ...
'' Blainville in
Lamarck Jean-Baptiste Pierre Antoine de Monet, chevalier de Lamarck (1 August 1744 – 18 December 1829), often known simply as Lamarck (; ), was a French naturalist, biologist, academic, and soldier. He was an early proponent of the idea that biolo ...
, 1818
* '' Interniloculus'' Suriano & Beverley-Burton, 1979 * ''
Lagenivaginopseudobenedenia ''Lagenivaginopseudobenedenia'' is a genus of monopisthocotylean monogenean (a group of ectoparasitic flatworms), included in the family Capsalidae.Whittington, I. D. 2004: The Capsalidae (Monogenea: Monopisthocotylea): a review of diversity, cla ...
'' Yamaguti, 1966 * ''
Listrocephalos CapsalidaeSatyu Yamaguti, Yamaguti, S. (1963) Systema Helminthum IV. Monogenea and Aspidocotylea. London-New York, Interscience Publishers. 699 pp. is a family of monopisthocotylean monogeneans, which includes about 200 species. The monophyly o ...
'' Bullard, Payne & Braswell, 2004 * ''
Macrophyllida Capsalidae Yamaguti, S. (1963) Systema Helminthum IV. Monogenea and Aspidocotylea. London-New York, Interscience Publishers. 699 pp. is a family of monopisthocotylean monogeneans, which includes about 200 species. The monophyly of the Capsalida ...
'' Johnston, 1929 * ''
Mediavagina Capsalidae Yamaguti, S. (1963) Systema Helminthum IV. Monogenea and Aspidocotylea. London-New York, Interscience Publishers. 699 pp. is a family of monopisthocotylean monogeneans, which includes about 200 species. The monophyly of the Capsalida ...
'' Lawler & Hargis, 1968 * ''
Megalobenedenia Capsalidae Yamaguti, S. (1963) Systema Helminthum IV. Monogenea and Aspidocotylea. London-New York, Interscience Publishers. 699 pp. is a family of monopisthocotylean monogeneans, which includes about 200 species. The monophyly of the Capsalida ...
'' Egorova, 1994 * ''
Megalocotyle Capsalidae Yamaguti, S. (1963) Systema Helminthum IV. Monogenea and Aspidocotylea. London-New York, Interscience Publishers. 699 pp. is a family of monopisthocotylean monogeneans, which includes about 200 species. The monophyly of the Capsalida ...
'' Folda, 1928 * ''
Menziesia ''Rhododendron'' sect. ''Sciadorhodion'' is a subsection of Rhododendron subg. Azaleastrum, subgenus ''Azaleastrum'' in the genus ''Rhododendron''. It comprises 16 species of deciduous shrubss native to East Asia and North America. Taxonomy Plan ...
'' Gibson, 1976 * '' Metabenedeniella'' Yamaguti, 1958 * ''
Nasicola Capsalidae Yamaguti, S. (1963) Systema Helminthum IV. Monogenea and Aspidocotylea. London-New York, Interscience Publishers. 699 pp. is a family of monopisthocotylean monogeneans, which includes about 200 species. The monophyly of the Capsalida ...
'' Yamaguti, 1968 * ''
Neobenedenia ''Neobenedenia'' is a genius of monopisthocotylean monogenean flatworm parasites. The most common species, ''Neobenedenia melleni'', causes problematic disease in public aquariums. It was named after ichthyologist Ida May Mellen (1877–1970) ...
'' Yamaguti, 1963 * '' Neobenedeniella'' Yamaguti, 1963 * ''
Neoentobdella Capsalidae Yamaguti, S. (1963) Systema Helminthum IV. Monogenea and Aspidocotylea. London-New York, Interscience Publishers. 699 pp. is a family of monopisthocotylean monogeneans, which includes about 200 species. The monophyly of the Capsalida ...
'' Kearn & Whittington, 2005 * ''
Nitzschia ''Nitzschia'' is a common pennate marine and freshwater diatom. In the scientific literature, this genus, named after Christian Ludwig Nitzsch, is sometimes referred to incorrectly as ''Nitzchia'', and it has many species described, which all hav ...
''
Baer Baer (or Bär, from ) or Van Baer is a surname. Notable people with the surname include: Baer * Alan Baer, American tuba player * Arthur "Bugs" Baer (1886–1969), American journalist and humorist * Buddy Baer (1915–1986), American boxer * Byro ...
, 1826
* ''
Oligoncobenedenia Capsalidae Yamaguti, S. (1963) Systema Helminthum IV. Monogenea and Aspidocotylea. London-New York, Interscience Publishers. 699 pp. is a family of monopisthocotylean monogeneans, which includes about 200 species. The monophyly of the Capsalida ...
'' Yamaguti, 1965 * '' Pseudoallobenedenia'' Yamaguti, 1966 * '' Pseudobenedenia'' Johnston, 1931 * '' Pseudobenedeniella'' Timofeeva, Gaevskaja & Kovaliova, 1987 * '' Pseudobenedenoides'' Szidat, 1969 * ''
Pseudoentobdella Capsalidae Yamaguti, S. (1963) Systema Helminthum IV. Monogenea and Aspidocotylea. London-New York, Interscience Publishers. 699 pp. is a family of monopisthocotylean monogeneans, which includes about 200 species. The monophyly of the Capsalida ...
'' Yamaguti, 1963 * ''
Pseudomegalocotyla Capsalidae Yamaguti, S. (1963) Systema Helminthum IV. Monogenea and Aspidocotylea. London-New York, Interscience Publishers. 699 pp. is a family of monopisthocotylean monogeneans, which includes about 200 species. The monophyly of the Capsalida ...
'' Yamaguti, 1963 * ''
Pseudonitzschia Capsalidae Yamaguti, S. (1963) Systema Helminthum IV. Monogenea and Aspidocotylea. London-New York, Interscience Publishers. 699 pp. is a family of monopisthocotylean monogeneans, which includes about 200 species. The monophyly of the Capsalida ...
'' Yamaguti, 1968 * '' Sessilorbis'' Mamaev, 1970 * ''
Sprostonia Capsalidae Yamaguti, S. (1963) Systema Helminthum IV. Monogenea and Aspidocotylea. London-New York, Interscience Publishers. 699 pp. is a family of monopisthocotylean monogeneans, which includes about 200 species. The monophyly of the Capsalida ...
'' Bychowsky, 1957 * ''
Sprostoniella Capsalidae Yamaguti, S. (1963) Systema Helminthum IV. Monogenea and Aspidocotylea. London-New York, Interscience Publishers. 699 pp. is a family of monopisthocotylean monogeneans, which includes about 200 species. The monophyly of the Capsalida ...
'' Bychowsky & Nagibina, 1967 * ''
Tetrasepta Capsalidae Yamaguti, S. (1963) Systema Helminthum IV. Monogenea and Aspidocotylea. London-New York, Interscience Publishers. 699 pp. is a family of monopisthocotylean monogeneans, which includes about 200 species. The monophyly of the Capsalida ...
'' Suriano, 1975 * '' Trilobiodiscus'' Bychowsky & Nagibina, 1967 * ''
Trimusculotrema Capsalidae Yamaguti, S. (1963) Systema Helminthum IV. Monogenea and Aspidocotylea. London-New York, Interscience Publishers. 699 pp. is a family of monopisthocotylean monogeneans, which includes about 200 species. The monophyly of the Capsalida ...
'' Whittington & Barton, 1990 * '' Tristoma''
Cuvier Jean Léopold Nicolas Frédéric, baron Cuvier (23 August 1769 – 13 May 1832), known as Georges Cuvier (; ), was a French natural history, naturalist and zoology, zoologist, sometimes referred to as the "founding father of paleontology". Cuv ...
, 1817
* ''
Tristomella Capsalidae Yamaguti, S. (1963) Systema Helminthum IV. Monogenea and Aspidocotylea. London-New York, Interscience Publishers. 699 pp. is a family of monopisthocotylean monogeneans, which includes about 200 species. The monophyly of the Capsalida ...
'' Guiart, 1938 * '' Trochopella''
Euzet Euzet (; ) is a commune in the Gard department in southern France. Population See also *Communes of the Gard department *Occitania (administrative region) Occitania ( ; ; ) is the southernmost Regions of France, administrative region ...
& Trilles, 1962
* ''
Trochopus Capsalidae Yamaguti, S. (1963) Systema Helminthum IV. Monogenea and Aspidocotylea. London-New York, Interscience Publishers. 699 pp. is a family of monopisthocotylean monogeneans, which includes about 200 species. The monophyly of the Capsalida ...
'' Diesing, 1850


References

{{Taxonbar, from=Q5019492 Monopisthocotylea Platyhelminthes families