Barsboldia
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''Barsboldia'' (meaning "of Barsbold", a well-known
Mongolian Mongolian may refer to: * Something of, from, or related to Mongolia, a country in Asia * Mongolian people, or Mongols * Bogd Khanate of Mongolia, the government of Mongolia, 1911–1919 and 1921–1924 * Mongolian language * Mongolian alphabet * ...
paleontologist Paleontology, also spelled as palaeontology or palæontology, is the scientific study of the life of the past, mainly but not exclusively through the study of fossils. Paleontologists use fossils as a means to classify organisms, measure geolo ...
) is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of large
hadrosaurid Hadrosaurids (), also hadrosaurs or duck-billed dinosaurs, are members of the ornithischian family Hadrosauridae. This group is known as the duck-billed dinosaurs for the flat duck-bill appearance of the bones in their snouts. The ornithopod fami ...
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutio ...
from the early Maastrichtian
Nemegt Formation The Nemegt Formation (also known as Nemegtskaya Svita) is a geological formation in the Gobi Desert of Mongolia, dating to the Late Cretaceous. The formation consists of river channel sediments and contains fossils of fish, turtles, crocodilians ...
of Ömnogöv',
Mongolia Mongolia is a landlocked country in East Asia, bordered by Russia to the north and China to the south and southeast. It covers an area of , with a population of 3.5 million, making it the world's List of countries and dependencies by po ...
. It is known from a partial
vertebra Each vertebra (: vertebrae) is an irregular bone with a complex structure composed of bone and some hyaline cartilage, that make up the vertebral column or spine, of vertebrates. The proportions of the vertebrae differ according to their spina ...
l column, partial
pelvis The pelvis (: pelves or pelvises) is the lower part of an Anatomy, anatomical Trunk (anatomy), trunk, between the human abdomen, abdomen and the thighs (sometimes also called pelvic region), together with its embedded skeleton (sometimes also c ...
, and some
ribs The rib cage or thoracic cage is an endoskeletal enclosure in the thorax of most vertebrates that comprises the ribs, vertebral column and sternum, which protect the vital organs of the thoracic cavity, such as the heart, lungs and great vessels ...
.


Discovery

In 1970, a Polish-Mongolian expedition near the Nemegt found the skeleton of an ornithopod and first assigned it to ''
Saurolophus angustirostris ''Saurolophus'' (; meaning "lizard crest") is a genus of large hadrosaurid dinosaur from the Late Cretaceous period of Asia and North America, that lived in what is now the Horseshoe Canyon and Nemegt formations about 70 million to 66 million ...
''. However, two Polish paleontologists,
Teresa Maryańska Teresa Maryańska (1937 – 3 October 2019) was a Polish paleontologist who specialized in Mongolian dinosaurs, particularly pachycephalosaurians and ankylosaurians. She is considered not only as one of Poland's but also one of the world's leadin ...
and
Halszka Osmólska Halszka Osmólska (September 15, 1930 – March 31, 2008) was a Polish paleontologist who had specialized in Mongolian dinosaurs. Early life, family and education Osmólska was born in 1930 in Poznań. In 1949, she began to study biology at Fa ...
, came to the conclusion that it was a
lambeosaurine Lambeosauridae /ˌlæmbiəˈsɔːraɪniː/ (meaning 'lambe's lizards') is an extinct group of crested hadrosauroid dinosaurs. Description Size Uncertainty surrounds the size of lambeosaurs from the European continent. Hadrosaurs found there, a ...
that had to represent a separate species. They named and described ''Barsboldia sicinskii'' based on the holotype specimen ZPAL MgD-1/110. The genus name honors the Mongolian paleontologist Rinchen Barsbold, while the species name honors Wojciech Siciński, the technician at the Warsaw Paleobiological Institute who prepared the skeleton. The holotype specimen, found in a layer of the
Nemegt Formation The Nemegt Formation (also known as Nemegtskaya Svita) is a geological formation in the Gobi Desert of Mongolia, dating to the Late Cretaceous. The formation consists of river channel sediments and contains fossils of fish, turtles, crocodilians ...
dating to the early Maastrichtian, consisted of a partial skeleton consisting of nine back
vertebra Each vertebra (: vertebrae) is an irregular bone with a complex structure composed of bone and some hyaline cartilage, that make up the vertebral column or spine, of vertebrates. The proportions of the vertebrae differ according to their spina ...
e, nine hip vertebrae, fifteen tail vertebrae, a left ilium, parts of the left and right pubis, several ribs, and a few fragments of the hind limbs, with the backbone largely articulated. The anterior and posteriormost portions of the skeleton were lost due to erosion.


Description

''Barsboldia'' was a large hadrosaur, previously estimated at in length and in body mass. In 2011, the tibial length was measured at , rivaling that of ''
Shantungosaurus ''Shantungosaurus'' (meaning "''Shandong Lizard''") is a genus of very large saurolophine hadrosaurid dinosaur found in the Late Cretaceous Wangshi Group of the Shandong Peninsula in China, containing a single species, ''Shantungosaurus gigante ...
'' at and that of ''
Magnapaulia ''Magnapaulia'' is a genus of herbivorous lambeosaurine hadrosaurid dinosaurs known from the Late Cretaceous Baja California, of northwestern Mexico. It contains a single species, ''Magnapaulia laticaudus''. ''Magnapaulia'' was first described in ...
'' at ; this indicates that ''Barsboldia'' could have possibly reached within the range of in total body length. Similar to ''
Ouranosaurus ''Ouranosaurus'' is a genus of herbivorous basal hadrosauriform dinosaur that lived during the Aptian stage of the Early Cretaceous of modern-day Niger and Cameroon. ''Ouranosaurus'' measured about long and weighed . Two rather complete fossils ...
'', the most distinctive features of ''Barsboldia'' are found in the
neural spines Each vertebra (: vertebrae) is an irregular bone with a complex structure composed of bone and some hyaline cartilage, that make up the vertebral column or spine, of vertebrates. The proportions of the vertebrae differ according to their spinal ...
. These are very tall, particularly over the hips, and were described as second only to those of ''
Hypacrosaurus altispinus ''Hypacrosaurus'' (meaning "near the highest lizard" reek υπο-, ''hypo-'' = less + ακρος, ''akros'', high because it was almost but not quite as large as ''Tyrannosaurus'') is an extinct genus of duckbill dinosaur similar in appearance ...
'' and the tips of those found in the first few vertebrae of the tail are club-shaped, possibly a sign of old age.


Phylogeny

Maryańska and Osmólska described their new genus as a
lambeosaurine Lambeosauridae /ˌlæmbiəˈsɔːraɪniː/ (meaning 'lambe's lizards') is an extinct group of crested hadrosauroid dinosaurs. Description Size Uncertainty surrounds the size of lambeosaurs from the European continent. Hadrosaurs found there, a ...
(or hollow-crested duckbill), the first from the Nemegt Formation, although it lacked a skull. However, the sacrum has a keel along the bottom, a possible lambeosaurine feature, and the bones closely resemble those of ''Hypacrosaurus''. With only one partial skeleton known, and no skull, the genus has been considered dubious or a possible lambeosaurine of uncertain placement. A newer study published in 2011 suggests that ''Barsboldia'' is actually a valid
saurolophine Saurolophinae is a subfamily (biology), subfamily of hadrosaurid dinosaurs. It has since the mid-20th century generally been called the Hadrosaurinae, a group of largely non-crested hadrosaurs related to the crested sub-family Lambeosaurinae. How ...
. The following
cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not s ...
was recovered in the 2011 phylogenetic analysis of
Hadrosauroidea Hadrosauroidea is a clade or superfamily of ornithischian dinosaurs that includes the "duck-billed" dinosaurs, or Hadrosauridae, and all dinosaurs more closely related to them than to ''Iguanodon''. Their remains have been recovered in Asia, Euro ...
by Prieto-Márquez (the relationships within Lambeosaurinae and between basal hadrosauroids aren't shown).


Paleobiology

As a hadrosaurid, ''Barsboldia'' would have been a large
bipedal Bipedalism is a form of terrestrial locomotion where an animal moves by means of its two rear (or lower) limbs or legs. An animal or machine that usually moves in a bipedal manner is known as a biped , meaning 'two feet' (from Latin ''bis'' ...
/
quadruped Quadrupedalism is a form of locomotion in which animals have four legs that are used to bear weight and move around. An animal or machine that usually maintains a four-legged posture and moves using all four legs is said to be a quadruped (fr ...
al
herbivore A herbivore is an animal anatomically and physiologically evolved to feed on plants, especially upon vascular tissues such as foliage, fruits or seeds, as the main component of its diet. These more broadly also encompass animals that eat ...
, eating
plant Plants are the eukaryotes that form the Kingdom (biology), kingdom Plantae; they are predominantly Photosynthesis, photosynthetic. This means that they obtain their energy from sunlight, using chloroplasts derived from endosymbiosis with c ...
s with a sophisticated skull that permitted a grinding motion analogous to
chewing Chewing or mastication is the process by which food is crushed and ground by the teeth. It is the first step in the process of digestion, allowing a greater surface area for digestive enzymes to break down the foods. During the mastication proc ...
, and was furnished with hundreds of continually-replaced
teeth A tooth (: teeth) is a hard, calcified structure found in the jaws (or mouths) of many vertebrates and used to break down food. Some animals, particularly carnivores and omnivores, also use teeth to help with capturing or wounding prey, tear ...
. If it was a lambeosaurine, it would have had a hollow crest formed out of expanded skull bones containing the nasal passages, with a function relating to identification by sight and sound.


See also

*
Timeline of hadrosaur research This timeline of hadrosaur research is a chronological listing of events in the History of paleontology, history of paleontology focused on the hadrosauroids, a group of herbivorous ornithopod dinosaurs popularly known as the duck-billed dinosa ...


References

{{Taxonbar, from1=Q18606529, from2=Q264109 Saurolophinae Dinosaur genera Maastrichtian dinosaurs Nemegt Formation Taxa named by Teresa Maryańska Taxa named by Halszka Osmólska Fossil taxa described in 1981 Dinosaurs of Mongolia