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Ayres's hawk-eagle (''Hieraaetus ayresii''), also referred to as Ayres' eagle,Newman, K (1998) Newman's Birds of Southern Africa. Halfway House: Southern Book Publishers. . is a medium-sized
bird of prey Birds of prey or predatory birds, also known as raptors, are hypercarnivorous bird species that actively hunt and feed on other vertebrates (mainly mammals, reptiles and other smaller birds). In addition to speed and strength, these predators h ...
in the family
Accipitridae The Accipitridae is one of the three families within the order Accipitriformes, and is a family of small to large birds with strongly hooked bills and variable morphology based on diet. They feed on a range of prey items from insects to medium-s ...
. It is native to African woodlands. Its name honors
South Africa South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring coun ...
n
ornithologist Ornithology is a branch of zoology that concerns the "methodological study and consequent knowledge of birds with all that relates to them." Several aspects of ornithology differ from related disciplines, due partly to the high visibility and th ...
Thomas Ayres.


Description

The adult male has blackish upperparts which are mottled with white, and usually has a white forehead and
supercilium The supercilium is a plumage feature found on the heads of some bird species. It is a stripe which runs from the base of the bird's beak above its eye, finishing somewhere towards the rear of the bird's head.Dunn and Alderfer (2006), p. 10 Also k ...
. The upper-wing coverts are similar. The tail is ashy grey with a broad black tip and three to four narrower dark bars.
Primary feathers Flight feathers (''Pennae volatus'') are the long, stiff, asymmetrically shaped, but symmetrically paired pennaceous feathers on the wings or tail of a bird; those on the wings are called remiges (), singular remex (), while those on the tai ...
and
secondary feathers Flight feathers (''Pennae volatus'') are the long, stiff, asymmetrically shaped, but symmetrically paired pennaceous feathers on the wings or tail of a bird; those on the wings are called remiges (), singular remex (), while those on the tai ...
are black. The underparts are white, with heavy dark brown spots and blotches on the breast and belly, becoming sparser on thighs and vent. The legs are well-feathered and pure white. The under-wing coverts are brown marked with white, the under side of flight feathers is dark and heavily barred lacking any noticeable grey patch. The eyes are yellow to orange, the
cere The beak, bill, or rostrum is an external anatomical structure found mostly in birds, but also in turtles, non-avian dinosaurs and a few mammals. A beak is used for eating, preening, manipulating objects, killing prey, fighting, probing for food, ...
and feet yellow, the bill is bluish horn coloured becoming paler towards the base, with a black tip. This species has a small but at times pronounced crest. The males are smaller than the females which are also darker and usually more densely spotted on the underparts, and have a smaller amount of white on the forehead and supercilium. There are two phases, the normal as described above and a
melanistic The term melanism refers to black pigment and is derived from the gr, μελανός. Melanism is the increased development of the dark-colored pigment melanin in the skin or hair. Pseudomelanism, also called abundism, is another variant of pi ...
phase, which is mostly black with white markings. The juvenile Ayres's hawk-eagle is dark brown on the back and coverts, similarly dark on the crown, slightly paler necked, largely white below from the throat to the crissum and legs with sparse dark brown markings variably along the chest, belly and flanks. When maturing, the young hawk-eagles often manifest a duskier, browner colour with somewhat of a rufous cast from below before moulting in its mature, adult plumage. Mostly the species is considered confusable with the
African hawk-eagle The African hawk-eagle (''Aquila spilogaster'') is a large bird of prey. Like all eagles, it belongs to the family ''Accipitridae''. This species’ feathered legs mark it as a member of the Aquilinae subfamily.Lerner, H., Christidis, L., Gamauf, ...
which is larger and lankier with a more protruding head, a longer tail, more sparsely marked underparts, and has more heavily marked wings with a white window above. It shares white "landing lights" at the fore of the wings with the similarly sized
booted eagle The booted eagle (''Hieraaetus pennatus'', also classified as ''Aquila pennata'') is a medium-sized mostly migratory bird of prey with a wide distribution in the Palearctic and southern Asia, wintering in the tropics of Africa and Asia, with a ...
but that species is usually rather differently marked overall (either paler and browner or all dark in dark morphs) and also similarly sized black sparrowhawks are mentioned as a confusion species but sparrowhawks are rather differently proportioned with bare, dull green-yellow legs. The Ayres's hawk-eagle is a medium-sized raptor but quite a small eagle, about the weight of a
peregrine falcon The peregrine falcon (''Falco peregrinus''), also known as the peregrine, and historically as the duck hawk in North America, is a cosmopolitan bird of prey ( raptor) in the family Falconidae. A large, crow-sized falcon, it has a blue-grey ba ...
albeit with a slightly greater length and wingspan more akin to a
common buzzard The common buzzard (''Buteo buteo'') is a medium-to-large bird of prey which has a large range. A member of the genus '' Buteo'', it is a member of the family Accipitridae. The species lives in most of Europe and extends its breeding range acr ...
, with a total length of and has a wingspan of .Kemp, A. C., P. F. D. Boesman, and J. S. Marks (2020). ''Ayres's Hawk-Eagle (Hieraaetus ayresii)'', version 1.0. In Birds of the World (J. del Hoyo, A. Elliott, J. Sargatal, D. A. Christie, and E. de Juana, Editors). Cornell Lab of Ornithology, Ithaca, NY, USA. A small sample of males have been known to weigh from and females weigh from . The average weight of four adult males was while that of 10 adult females was , juvenile Ayres's being some 5% lighter in spite of slightly greater wing sizes.Hartley, R. R., & Mundy, P. J. (2003). Morphometrics and status of Ayres's Hawk-eagle in Zimbabwe. Journal of Raptor Research, 37(1), 44-54. The wing chord is and from in males and females, respectively. The tail and tarsus length is from and from in males and and from in females. The species thus manifests quite strong
sexual dimorphism Sexual dimorphism is the condition where the sexes of the same animal and/or plant species exhibit different morphological characteristics, particularly characteristics not directly involved in reproduction. The condition occurs in most an ...
in favor of the female, which does not overlap in standard measurements with the smaller male. Bill length is from , averaging in males and in females. The enlarged hallux claw on the rear toe is from , averaging in males and in females, quite large for the small size of the eagle.Parry, S. J. (2001). ''The booted eagles (Aves: Accipitridae): perspectives in evolutionary biology''. University of London, University College London (United Kingdom).


Distribution and habitat

Ayres's hawk-eagle has a patchy sub-Saharan distribution ranging from
Sierra Leone Sierra Leone,)]. officially the Republic of Sierra Leone, is a country on the southwest coast of West Africa. It is bordered by Liberia to the southeast and Guinea surrounds the northern half of the nation. Covering a total area of , Sierra ...
east to
Somalia Somalia, , Osmanya script: 𐒈𐒝𐒑𐒛𐒐𐒘𐒕𐒖; ar, الصومال, aṣ-Ṣūmāl officially the Federal Republic of SomaliaThe ''Federal Republic of Somalia'' is the country's name per Article 1 of thProvisional Constituti ...
, and south to northern
Namibia Namibia (, ), officially the Republic of Namibia, is a country in Southern Africa. Its western border is the Atlantic Ocean. It shares land borders with Zambia and Angola to the north, Botswana to the east and South Africa to the south and ...
and northeast
South Africa South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring coun ...
. Its central range is from the southern
Democratic Republic of the Congo The Democratic Republic of the Congo (french: République démocratique du Congo (RDC), colloquially "La RDC" ), informally Congo-Kinshasa, DR Congo, the DRC, the DROC, or the Congo, and formerly and also colloquially Zaire, is a country in ...
and southern
Uganda }), is a landlocked country in East Africa. The country is bordered to the east by Kenya, to the north by South Sudan, to the west by the Democratic Republic of the Congo, to the south-west by Rwanda, and to the south by Tanzania. The sou ...
down almost throughout
Kenya ) , national_anthem = " Ee Mungu Nguvu Yetu"() , image_map = , map_caption = , image_map2 = , capital = Nairobi , coordinates = , largest_city = Nairobi ...
,
Tanzania Tanzania (; ), officially the United Republic of Tanzania ( sw, Jamhuri ya Muungano wa Tanzania), is a country in East Africa within the African Great Lakes region. It borders Uganda to the north; Kenya to the northeast; Comoro Islands an ...
,
Zambia Zambia (), officially the Republic of Zambia, is a landlocked country at the crossroads of Central, Southern and East Africa, although it is typically referred to as being in Southern Africa at its most central point. Its neighbours are ...
and
Mozambique Mozambique (), officially the Republic of Mozambique ( pt, Moçambique or , ; ny, Mozambiki; sw, Msumbiji; ts, Muzambhiki), is a country located in southeastern Africa bordered by the Indian Ocean to the east, Tanzania to the north, Mal ...
as well as much of
Angola , national_anthem = "Angola Avante"() , image_map = , map_caption = , capital = Luanda , religion = , religion_year = 2020 , religion_ref = , coordina ...
to northeastern
Namibia Namibia (, ), officially the Republic of Namibia, is a country in Southern Africa. Its western border is the Atlantic Ocean. It shares land borders with Zambia and Angola to the north, Botswana to the east and South Africa to the south and ...
, northern
Botswana Botswana (, ), officially the Republic of Botswana ( tn, Lefatshe la Botswana, label= Setswana, ), is a landlocked country in Southern Africa. Botswana is topographically flat, with approximately 70 percent of its territory being the Kalah ...
and northern
Zimbabwe Zimbabwe (), officially the Republic of Zimbabwe, is a landlocked country located in Southeast Africa, between the Zambezi and Limpopo Rivers, bordered by South Africa to the south, Botswana to the south-west, Zambia to the north, and Moz ...
. In the rainy season moves out of denser and taller deciduous
woodland A woodland () is, in the broad sense, land covered with trees, or in a narrow sense, synonymous with wood (or in the U.S., the ''plurale tantum'' woods), a low-density forest forming open habitats with plenty of sunlight and limited shade (see ...
s of central Africa into more open albeit treed
savanna A savanna or savannah is a mixed woodland-grassland (i.e. grassy woodland) ecosystem characterised by the trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach the ground to ...
habitat further south, and probably into coastal East Africa; as the rains cause the leaves to emerge turning woodland into
forest A forest is an area of land dominated by trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, and ecological function. The United Nations' ...
s and tree savanna into woodland. It may then enter towns in South Africa to prey mainly on
doves Columbidae () is a bird family consisting of doves and pigeons. It is the only family in the order Columbiformes. These are stout-bodied birds with short necks and short slender bills that in some species feature fleshy ceres. They primarily ...
and feral pigeons. A similar north to south movement is expected in west Africa, where species has been recorded as a
vagrant Vagrancy is the condition of homelessness without regular employment or income. Vagrants (also known as bums, vagabonds, rogues, tramps or drifters) usually live in poverty and support themselves by begging, scavenging, petty theft, tempora ...
west to
Senegal Senegal,; Wolof: ''Senegaal''; Pulaar: 𞤅𞤫𞤲𞤫𞤺𞤢𞥄𞤤𞤭 (Senegaali); Arabic: السنغال ''As-Sinighal'') officially the Republic of Senegal,; Wolof: ''Réewum Senegaal''; Pulaar : 𞤈𞤫𞤲𞤣𞤢𞥄𞤲𞤣� ...
and
Gambia The Gambia,, ff, Gammbi, ar, غامبيا officially the Republic of The Gambia, is a country in West Africa. It is the smallest country within mainland AfricaHoare, Ben. (2002) ''The Kingfisher A-Z Encyclopedia'', Kingfisher Publicatio ...
, albeit unconfirmed. It tends to occur around woodlands of ''
Brachystegia ''Brachystegia'' is a genus of tree of the subfamily Detarioideae that is native to tropical Africa. Trees of the genus are commonly known as miombo, and are dominant in the miombo woodlands of central and southern tropical Africa. The Zambez ...
'' and '' Baikiaea'' in the core south-central African part of their distribution. It is not typically a forest dwelling species and prefers mature woodland among often irregular,
rocky ''Rocky'' is a 1976 American sports drama film directed by John G. Avildsen and written by and starring Sylvester Stallone. It is the first installment in the ''Rocky'' franchise and stars Talia Shire, Burt Young, Carl Weathers, and Burges ...
terrain. Occasionally but not regularly they may habituate to ''
Acacia ''Acacia'', commonly known as the wattles or acacias, is a large genus of shrubs and trees in the subfamily Mimosoideae of the pea family Fabaceae. Initially, it comprised a group of plant species native to Africa and Australasia. The genus na ...
''
plantation A plantation is an agricultural estate, generally centered on a plantation house, meant for farming that specializes in cash crops, usually mainly planted with a single crop, with perhaps ancillary areas for vegetables for eating and so on. The ...
s and
riparian zone A riparian zone or riparian area is the interface between land and a river or stream. Riparian is also the proper nomenclature for one of the terrestrial biomes of the Earth. Plant habitats and communities along the river margins and banks a ...
s. Ayres's hawk-eagles tend to be rare in hyper
arid A region is arid when it severely lacks available water, to the extent of hindering or preventing the growth and development of plant and animal life. Regions with arid climates tend to lack vegetation and are called xeric or desertic. Most ...
and, of course, treeless areas.


Biology

Ayres's hawk-eagle is a bird hunter, almost to the exclusion of any other type of prey, especially
dove Columbidae () is a bird family consisting of doves and pigeons. It is the only family in the order Columbiformes. These are stout-bodied birds with short necks and short slender bills that in some species feature fleshy ceres. They primarily ...
s and
pigeon Columbidae () is a bird family consisting of doves and pigeons. It is the only family in the order Columbiformes. These are stout-bodied birds with short necks and short slender bills that in some species feature fleshy ceres. They primarily ...
s, it soars high above the ground to search for prey. It is unique for a
Aquilinae The Aquilinae are a subfamily of eagles of the family Accipitridae. The general common name used for members of this subfamily is "booted eagle", although this is also the common name of a member of the subfamily.Lerner, H., Christidis, L., Gamau ...
eagle in its
falcon Falcons () are birds of prey in the genus ''Falco'', which includes about 40 species. Falcons are widely distributed on all continents of the world except Antarctica, though closely related raptors did occur there in the Eocene. Adult falcons ...
-like highly aerial method for hunting birds on the wing, an aptitude it shares with the Asian rufous-bellied eagle. Favored in diet in some areas are laughing doves and red-eyed doves. Once a bird has been singled out, the eagle stoops to intercept it in mid air. Other than birds it has been recorded as catching a few mammals including bush squirrels, and
fruit bat Megabats constitute the family Pteropodidae of the order Chiroptera ( bats). They are also called fruit bats, Old World fruit bats, or—especially the genera ''Acerodon'' and ''Pteropus''—flying foxes. They are the only member of the ...
s. They are an exceptional agile and fast eagle and can overtake many birds in flight. The Ayres's hawk-eagle predominantly hunt quite small birds, typically weighing from , and can sometimes include nestling birds in their diet. However, it quite commonly takes heavier birds than in the aforementioned weight range, including
rock pigeon The rock dove, rock pigeon, or common pigeon ( also ; ''Columba livia'') is a member of the bird family Columbidae (doves and pigeons). In common usage, it is often simply referred to as the "pigeon". The domestic pigeon (''Columba livia dom ...
s and African grey hornbills. Sometimes they will overtake birds up to over twice their size including white-faced whistling ducks, assorted
francolin Francolins are birds in the tribe Gallini that traditionally have been placed in the genus ''Francolinus'', but now commonly are divided into multiple genera. As previously defined, they were paraphyletic as the genus '' Pternistis'', which wa ...
s and
guineafowl Guineafowl (; sometimes called "pet speckled hens" or "original fowl") are birds of the family Numididae in the order Galliformes. They are endemic to Africa and rank among the oldest of the gallinaceous birds. Phylogenetically, they branched ...
s and even other birds of prey including gabar goshawks and shikras. Ayres's hawk-eagle is a monogamous, territorial solitary nester. The nest consists of a large platform of a twigs and sticks, lined with green leaves and typically concealed in the fork within a well-leafed, large tree, usually from above the ground. The nest can be from across. Eggs are laid from April to September, peaking in April and May. A single egg is laid, measuring on average . The female is almost solely responsible for incubation which takes about 43 days, the male brings her food every two to three days. The chick is fed almost daily, fledging at about 73 days old and becoming fully independent from its parents approximately three months after fledging.Dewhurst, C. E., & van Someren, C. (1988). Observation on the breeding ecology of Ayres' hawk eagle Hieraaetus ayresii at Karen, Nairobi, Kenya. GABAR., 3(2), 85-93.


Conservation status

Although Ayres's hawk-eagle is an uncommon bird throughout its range, it is classified as
Least Concern A least-concern species is a species that has been categorized by the International Union for Conservation of Nature (IUCN) as evaluated as not being a focus of species conservation because the specific species is still plentiful in the wild. T ...
(LC) by the IUCN, due to its large range and its numbers, while small, appearing stable at the present time. The estimated global population is only 1000 to 10,000 individuals. The main threat faced by the species is
cutting Cutting is the separation or opening of a physical object, into two or more portions, through the application of an acutely directed force. Implements commonly used for cutting are the knife and saw, or in medicine and science the scalpel and ...
of woodland. Furthermore, it is persecuted at times for hunting down
domestic pigeon The domestic pigeon (''Columba livia domestica'' or ''Columba livia'' ''forma'' ''domestica'') is a pigeon subspecies that was derived from the rock dove (also called the rock pigeon). The rock pigeon is the world's oldest domesticated bird. Me ...
s.


References


External links

* (Ayres's hawk-eagle = ) Ayres' eagle
Species text in The Atlas of Southern African Birds
{{Taxonbar, from=Q1260400 Ayres's hawk-eagle Birds of prey of Sub-Saharan Africa Ayres's hawk-eagle Ayres's hawk-eagle Taxonomy articles created by Polbot