''Asiaceratops'' (meaning "Asian horned face") is a

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...

of herbivorous

ceratopsian Ceratopsia or Ceratopia ( or ; Ancient Greek, Greek: "horned faces") is a group of herbivore, herbivorous, beaked dinosaurs that thrived in what are now North America, Europe, and Asia, during the Cretaceous Period (geology), Period, although anc ...

dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...

. It lived during the

Early Early may refer to: History * The beginning or oldest part of a defined historical period, as opposed to middle or late periods, e.g.: ** Early Christianity ** Early modern Europe Places in the United States * Early, Iowa * Early, Texas * Early ...

Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', ...

. The type species, ''A. salsopaludalis'' is known from

Uzbekistan Uzbekistan (, ; uz, Ozbekiston, italic=yes / , ; russian: Узбекистан), officially the Republic of Uzbekistan ( uz, Ozbekiston Respublikasi, italic=yes / ; russian: Республика Узбекистан), is a doubly landlocked co ...

, while ''A. sulcidens'' is known from

China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's List of countries and dependencies by population, most populous country, with a Population of China, population exceeding 1.4 billion, slig ...

and

Mongolia Mongolia; Mongolian script: , , ; lit. "Mongol Nation" or "State of Mongolia" () is a landlocked country in East Asia, bordered by Russia to the north and China to the south. It covers an area of , with a population of just 3.3 million ...

Discovery and naming

The

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specim ...

, ''Asiaceratops salsopaludalis'', was formally described by Lev Nesov, L.F. Kaznyshkina and Gennadiy Olegovich Cherepanov in 1989. The generic name combines a reference to Asia with ''~ceratops'', "horned face". The specific name means "of the salt marsh" in

Latin Latin (, or , ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through ...

. In the same publication '' Microceratops sulcidens'' Bohlin 1953 was renamed into a second species of ''Asiaceratops'': ''Asiaceratops sulcidens''.L.A. Nessov, L F. Kaznyshkina, and G.O. Cherepanov. (1989). esozoic ceratopsian dinosaurs and crocodiles of central Asia In: Bogdanova and Khozatskii (eds.), ''Theoretical and Applied Aspects of Modern Palaeontology'' pp 144-154 The

holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of seve ...

of ''Asiaceratops salsopaludalis'', CCMGE 9/12457, was found in Uzbekistan in a layer of the

Khodzhakul Formation The Khodzhakul Formation is a Cenomanian aged geologic formation in Uzbekistan. Dinosaur remains are among the fossils that have been recovered from the formation.Weishampel, et al., 2004, "Dinosaur distribution." pp.517-607 As well as those of ...

dating from the early

Cenomanian The Cenomanian is, in the ICS' geological timescale, the oldest or earliest age of the Late Cretaceous Epoch or the lowest stage of the Upper Cretaceous Series. An age is a unit of geochronology; it is a unit of time; the stage is a unit in ...

, about ninety-nine million years old. It consists of a part of a left

maxilla The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. T ...

. Some other fragments were in 1989 referred to the species, among them teeth and a phalanx. In 1995 Nesov referred more material, from three Uzbek sites, mostly skull elements and a partial humerus, of individuals of different ages.L.A. Nessov, (1995), ''Dinozavri severnoi Yevrazii: Novye dannye o sostave kompleksov, ekologii i paleobiogeografii'', Institute for Scientific Research on the Earth's Crust, St. Petersburg State University, St. Petersburg 156 pp A second species, ''A. sulcidens'', was created to house ''

Microceratus ''Microceratus'' (meaning "small-horned") is a genus of small ceratopsian dinosaur that lived in the Cretaceous period in Asia. It walked on two legs, had short front arms, a characteristic ceratopsian frill and beak-like mouth, and was aroun ...

sulcidens''. Bohlin, B. (1953). ''Fossil reptiles from Mongolia and Kansu. Reports from the Scientific Expedition to the North-western Provinces of China under Leadership of Dr. Sven Hedin. VI. Vertebrate Palaeontology 6.'' The Sino-Swedish Expedition Publications 37, 113 pp It is known from remains discovered in

Early Cretaceous The Early Cretaceous (geochronology, geochronological name) or the Lower Cretaceous (chronostratigraphy, chronostratigraphic name), is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 145& ...

(

Aptian The Aptian is an age in the geologic timescale or a stage in the stratigraphic column. It is a subdivision of the Early or Lower Cretaceous Epoch or Series and encompasses the time from 121.4 ± 1.0 Ma to 113.0 ± 1.0 Ma (million years ag ...

Albian The Albian is both an age of the geologic timescale and a stage in the stratigraphic column. It is the youngest or uppermost subdivision of the Early/Lower Cretaceous Epoch/ Series. Its approximate time range is 113.0 ± 1.0 Ma to 100.5 ± 0 ...

)-aged rocks located in

(

Xinminpu Group The Xīnmínbǎo Group () is a group of geological formations in north central China. They occur across a large depression between the Altai mountains of Mongolia to the north and the Qilian mountains of the Qinghai Plateau to the south, in the ...

) and

and the holotype of ''A. sulcidens'', no inventory number given, consists of two teeth, centra, an incomplete tibia, tarsals and left pes. ''Asiaceratops'' has often been considered a ''

nomen dubium In binomial nomenclature, a ''nomen dubium'' (Latin for "doubtful name", plural ''nomina dubia'') is a scientific name that is of unknown or doubtful application. Zoology In case of a ''nomen dubium'' it may be impossible to determine whether a s ...

'', in view of the limited holotype material. A basicranium tentatively referred to ''Asiaceratops'' may show diagnostic characters of the taxon.

Classification

''Asiaceratops'' belonged to the

Ceratopsia Ceratopsia or Ceratopia ( or ; Greek: "horned faces") is a group of herbivorous, beaked dinosaurs that thrived in what are now North America, Europe, and Asia, during the Cretaceous Period, although ancestral forms lived earlier, in the Jurass ...

(the name is

Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...

for "horned faces"), a group of herbivorous dinosaurs with

parrot Parrots, also known as psittacines (), are birds of the roughly 398 species in 92 genera comprising the order Psittaciformes (), found mostly in tropical and subtropical regions. The order is subdivided into three superfamilies: the Psittacoide ...

-like beaks which thrived in

North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and th ...

and

Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an are ...

during the Cretaceous Period, which ended roughly 66 million years ago. All ceratopsians became extinct at the end of this era. In 1995 Nesov assigned ''Asiaceratops'' to an Asiaceratopsinae of its own. Recent

cladistic Cladistics (; ) is an approach to biological classification in which organisms are categorized in groups ("clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is typically shared derived char ...

analyses indicated, despite the presumed status as a ''nomen dubium'', a basal position in the

Leptoceratopsidae Leptoceratopsidae is an extinct family of neoceratopsian dinosaurs from Asia, North America and Europe. Leptoceratopsids resembled, and were closely related to, other neoceratopsians, such as the families Protoceratopsidae and Ceratopsidae, b ...

.M.J. Ryan, D.C. Evans, P.J. Currie, C.M. Brown, and D. Brinkman, (2012), "New leptoceratopsids from the Upper Cretaceous of Alberta, Canada", ''Cretaceous Research'' 35(1): 69-80

Diet

''Asiaceratops'', like all ceratopsians, was a

herbivore A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthpar ...

. During the Cretaceous, flowering plants were "geographically limited on the landscape", and so it is likely that this dinosaur fed on the predominant plants of the era: ferns, cycads, and conifers. It would have used its sharp ceratopsian beak to bite off the leaves or needles.

References

{{Taxonbar, from=Q134381 Leptoceratopsids Late Cretaceous dinosaurs of Asia Fossil taxa described in 1989 Ornithischian genera

Discovery and naming

Classification

Diet

See also

References