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''Aralia'' , or spikenard, is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
of the
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
Araliaceae The Araliaceae are a family of flowering plants composed of about 43 genera and around 1500 species consisting of primarily woody plants and some herbaceous plants. The morphology of Araliaceae varies widely, but it is predominantly distinguish ...
, consisting of 68 accepted
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of
deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, ...
or
evergreen In botany, an evergreen is a plant which has foliage that remains green and functional through more than one growing season. This also pertains to plants that retain their foliage only in warm climates, and contrasts with deciduous plants, whic ...
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...
s, shrubs, and
rhizomatous In botany and dendrology, a rhizome (; , ) is a modified subterranean plant stem that sends out roots and shoots from its nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from axillary buds and grow ho ...
herbaceous
perennial A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also wid ...
s. The genus is native to Asia and the Americas, with most species occurring in mountain woodlands. ''Aralia'' plants vary in size, with some herbaceous species only reaching tall, while some are trees growing to tall. ''Aralia'' plants have large bipinnate (doubly compound) leaves clustered at the ends of their stems or branches; in some species the leaves are covered with bristles. The stems of some woody species are quite prickly, as in ''
Aralia spinosa ''Aralia spinosa'', commonly known as devil's walking stick, is a woody species of plant in the genus ''Aralia'', family Araliaceae, native to eastern North America. The various names refer to the viciously sharp, spiny stems, petioles, and even ...
''. The
flower A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechani ...
s are whitish or greenish occurring in terminal panicles, and the spherical dark purple berry-like
fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in particu ...
s are popular with
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweig ...
s. ''Aralia'' species are used as food plants by the
larva A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. ...
e of some Lepidoptera species, including the
common emerald The common emerald (''Hemithea aestivaria'') is a moth of the family Geometridae. The species is found throughout the Nearctic and Palearctic regions and the Near East. It is mostly commonly found in the southern half of the British Isles. It was ...
(''Hemithea aestivaria''). There are many colours of aralia flowers. The main flower is whitish aralia. Some species, notably ''Aralia cordata'', are edible and are cultivated for human consumption.


Taxonomy

The taxonomic circumscription of the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
''Aralia'' has varied greatly. Species formerly included in wider views of the genus are now included in such separate genera as ''
Fatsia ''Fatsia'' is a small genus of three species of evergreen shrubs in the family Araliaceae native to southern Japan and Taiwan. They typically have stout, sparsely branched stems bearing spirally-arranged, large leathery, palmately lobed leav ...
'', '' Macropanax'', '' Oreopanax'', ''
Panax The ''Panax'' (ginseng) genus belongs to the ''Araliaceae'' (ivy) family. ''Panax'' species are characterized by the presence of ginsenosides and gintonin. ''Panax'' is one of approximately 60 plant genera with a classical disjunct east Asian and ...
'', ''
Polyscias ''Polyscias'' is a genus of flowering plants in the Family (biology), family Araliaceae. They bear pinnately compound leaves. In 2003, a checklist and nomenclator (nomenclature), nomenclator was published for Araliaceae.David G. Frodin and Rafa� ...
'', ''
Pseudopanax ''Pseudopanax'' (Latin for "false ginseng") is a small genus of 12–20 species of evergreen plants, the majority of which are endemic to New Zealand. Flowers of the genus occur in terminal umbels. Taxonomy A 2000 molecular study established th ...
'', ''
Schefflera ''Schefflera'' is a genus of flowering plants in the family Araliaceae. With an estimated 600–900 species, the genus represents about half of its family. The plants are trees, shrubs or lianas, growing tall, with woody stems, the absence of a ...
'', and '' Tetrapanax''. The genus ''Dimorphanthus'', formerly considered distinct by some, is now included within ''Aralia'' as a section within that genus.


Species

, Plants of the World Online accepts the following species: *'' Aralia apioides'' Hand.-Mazz. *'' Aralia armata'' (Wall. ex G.Don) Seem. *'' Aralia atropurpurea'' Franch. *'' Aralia bahiana'' J.Wen *'' Aralia bicrenata'' Wooton & Standl. *'' Aralia bipinnata'' Blanco *'' Aralia cachemirica'' Decne. *'' Aralia caesia'' Hand.-Mazz. *'' Aralia californica'' S.Watson *'' Aralia castanopsiscola'' (Hayata) J.Wen *'' Aralia chinensis'' L. *''
Aralia continentalis ''Aralia'' , or spikenard, is a genus of the family (biology), family Araliaceae, consisting of 68 accepted species of deciduous or evergreen trees, shrubs, and rhizome, rhizomatous herbaceous plant, herbaceous perennial plant, perennials. The ge ...
'' Kitag. *''
Aralia cordata ''Aralia cordata'' is an upright herbaceous perennial plant growing up to in height, native to Japan, Korea and eastern China. Its common names include spikenard, herbal aralia, ''udo'' (from ja, ウド), Japanese spikenard, and mountain as ...
'' Thunb. *'' Aralia dasyphylla'' Miq. *'' Aralia dasyphylloides'' (Hand.-Mazz.) J.Wen *'' Aralia debilis'' J.Wen *''
Aralia decaisneana ''Aralia'' , or spikenard, is a genus of the family (biology), family Araliaceae, consisting of 68 accepted species of deciduous or evergreen trees, shrubs, and rhizome, rhizomatous herbaceous plant, herbaceous perennial plant, perennials. The ge ...
'' Hance *'' Aralia delavayi'' J.Wen *'' Aralia devendrae'' Pusalkar *'' Aralia duplex'' R.Chaves *'' Aralia echinocaulis'' Hand.-Mazz. *''
Aralia elata ''Aralia elata'', the Japanese angelica tree, Chinese angelica-tree, or Korean angelica-tree, is a woody plant belonging to the family Araliaceae. It is known as ''tara-no-ki'' (; ) in Japanese, and ''dureup-namu'' () in Korean. Description It i ...
'' (Miq.) Seem. *'' Aralia excelsa'' (Griseb.) J.Wen *'' Aralia fargesii'' Franch. *'' Aralia ferox'' Miq. *'' Aralia finlaysoniana'' (Wall. ex G.Don) Seem. *'' Aralia foliolosa'' Seem. ex C.B.Clarke *'' Aralia frodiniana'' J.Wen *'' Aralia gigantea'' J.Wen *'' Aralia gintungensis'' C.Y.Wu ex K.M.Feng *'' Aralia glabra'' Matsum. *'' Aralia glabrifoliolata'' (C.B.Shang) J.Wen *'' Aralia henryi'' Harms *'' Aralia hiepiana'' J.Wen & Lowry *'' Aralia hispida'' Vent. *'' Aralia humilis'' Cav. *'' Aralia hypoglauca'' (C.J.Qi & T.R.Cao) J.Wen & Y.F.Deng *'' Aralia indonesica'' Doweld *'' Aralia kansuensis'' G.Hoo *'' Aralia kingdon-wardii'' J.Wen, Lowry & Esser *''
Aralia leschenaultii ''Aralia leschenaultii'' is a species of plant in the family Araliaceae. It is found in China, India, Myanmar, and Sri Lanka Sri Lanka (, ; si, ශ්‍රී ලංකා, Śrī Laṅkā, translit-std=ISO (); ta, இலங்கை, ...
'' (DC.) J.Wen *'' Aralia malabarica'' Bedd. *'' Aralia melanocarpa'' (H.Lév.) Lauener *'' Aralia merrillii'' C.B.Shang *'' Aralia mexicana'' (C.B.Shang & X.P.Li) Frodin *'' Aralia montana'' Blume *'' Aralia nudicaulis'' L. *'' Aralia officinalis'' Z.Z.Wang *''
Aralia parasitica ''Aralia'' , or spikenard, is a genus of the family Araliaceae, consisting of 68 accepted species of deciduous or evergreen trees, shrubs, and rhizomatous herbaceous perennials. The genus is native to Asia and the Americas, with most species occu ...
'' (D.Don) Buch.-Ham. ex Otto *'' Aralia plumosa'' H.L.Li *'' Aralia racemosa'' L. *'' Aralia regeliana'' Marchal *'' Aralia rex'' (Ekman) J.Wen *'' Aralia scaberula'' G.Hoo *'' Aralia scopulorum'' Brandegee *'' Aralia searelliana'' Dunn *'' Aralia soratensis'' Marchal *'' Aralia spinifolia'' Merr. *''
Aralia spinosa ''Aralia spinosa'', commonly known as devil's walking stick, is a woody species of plant in the genus ''Aralia'', family Araliaceae, native to eastern North America. The various names refer to the viciously sharp, spiny stems, petioles, and even ...
'' L. *'' Aralia stellata'' (King) J.Wen *'' Aralia stipulata'' Franch. *''
Aralia subcordata ''Aralia'' , or spikenard, is a genus of the family (biology), family Araliaceae, consisting of 68 accepted species of deciduous or evergreen trees, shrubs, and rhizome, rhizomatous herbaceous plant, herbaceous perennial plant, perennials. The ge ...
'' (Wall. ex G.Don) J.Wen *'' Aralia thomsonii'' Seem. ex C.B.Clarke *'' Aralia tibetana'' G.Hoo *'' Aralia tomentella'' Franch. *'' Aralia undulata'' Hand.-Mazz. *'' Aralia urticifolia'' Blume ex Miq. *'' Aralia verticillata'' (Dunn) J.Wen *'' Aralia vietnamensis'' Ha *'' Aralia wangshanensis'' (W.C.Cheng) Y.F.Deng *'' Aralia warmingiana'' (Marchal) J.Wen *'' Aralia wilsonii'' Harms *''
Aralia yunnanensis ''Aralia'' , or spikenard, is a genus of the family Araliaceae, consisting of 68 accepted species of deciduous or evergreen trees, shrubs, and rhizomatous herbaceous perennials. The genus is native to Asia and the Americas, with most species occu ...
'' Franch.


Fossil record

One
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
endocarp Fruit anatomy is the plant anatomy of the internal structure of fruit. Fruits are the mature ovary or ovaries of one or more flowers. They are found in three main anatomical categories: aggregate fruits, multiple fruits, and simple fruits. Aggr ...
of †''Aralia pusilla'' has been described from a middle Miocene stratum of the Fasterholt area near
Silkeborg Silkeborg () is a Danish town with a population of 49,747 (1 January 2022).Jutland Jutland ( da, Jylland ; german: Jütland ; ang, Ēota land ), known anciently as the Cimbric or Cimbrian Peninsula ( la, Cimbricus Chersonesus; da, den Kimbriske Halvø, links=no or ; german: Kimbrische Halbinsel, links=no), is a peninsula of ...
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. Several
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
fruits of ''Aralia rugosa'' and †''Aralia tertiaria'' have been extracted from bore hole samples of the Middle Miocene fresh water deposits in Nowy Sacz Basin,
West Carpathians The Western Carpathians are a mountain range and geomorphological province that forms the western part of the Carpathian Mountains. The mountain belt stretches from the Low Beskids range of the Eastern Carpathians along the border of Poland wit ...
,
Poland Poland, officially the Republic of Poland, is a country in Central Europe. It is divided into 16 administrative provinces called voivodeships, covering an area of . Poland has a population of over 38 million and is the fifth-most populou ...
. Several ''Aralia'' macrofossils have been recovered from the late
Zanclean The Zanclean is the lowest stage or earliest age on the geologic time scale of the Pliocene. It spans the time between 5.332 ± 0.005 Ma (million years ago) and 3.6 ± 0.005 Ma. It is preceded by the Messinian Age of the Miocene Epoch, and ...
stage of
Pliocene The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58Pocapaglia,
Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical ...
.Messian to Zanclean vegetation and climate of Northern and Central Italy by Adele Bertini & Edoardo Martinetto, Bollettino della Societa Paleontologica Italiana, 47 (2), 2008, 105-121. Modena, 11 July 2008.


Notes


References

*Frodin, D. G. and R. Govaerts. 2003. World Checklist and Bibliography of Araliaceae. Kew, UK: The Royal Botanic Gardens, Kew. *Wen, J. 2004. Systematics and biogeography of ''Aralia'' L. sect. ''Dimorphanthus'' (Miq.) Miq. (Araliaceae). ''Cathaya'' 15-16: 1–187.


External links


''Aralia''
{{Taxonbar, from=Q132636 Apiales genera Flora of North America Flora of Asia