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An apple is an edible
fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in particul ...
produced by an apple tree (''Malus domestica''). Apple
trees In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are u ...
are cultivated worldwide and are the most widely grown species in the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nomencla ...
''
Malus ''Malus'' ( or ) is a genus of about 30–55 species of small deciduous trees or shrubs in the family Rosaceae, including the domesticated orchard apple, crab apples, wild apples, and rainberries. The genus is native to the temperate zone of th ...
''. The
tree In botany, a tree is a perennial plant with an elongated Plant stem, stem, or trunk (botany), trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondar ...
originated in Central Asia, where its wild ancestor, ''
Malus sieversii ''Malus sieversii'' is a wild apple native to the mountains of Central Asia in southern Kazakhstan. It has recently been shown to be the primary ancestor of most cultivars of the domesticated apple (''Malus domestica''). It was first described ...
'', is still found today. Apples have been grown for thousands of years in Asia and Europe and were brought to North America by European colonists. Apples have
religious Religion is usually defined as a social-cultural system of designated behaviors and practices, morals, beliefs, worldviews, texts, sanctified places, prophecies, ethics, or organizations, that generally relates humanity to supernatural, tran ...
and
mythological Myth is a folklore genre consisting of narratives that play a fundamental role in a society, such as foundational tales or origin myths. Since "myth" is widely used to imply that a story is not objectively true, the identification of a narrati ...
significance in many cultures, including Norse,
Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...
, and European Christian tradition. Apples grown from seed tend to be very different from those of their parents, and the resultant fruit frequently lacks desired characteristics. Generally, apple cultivars are propagated by clonal
grafting Grafting or graftage is a horticultural technique whereby tissues of plants are joined so as to continue their growth together. The upper part of the combined plant is called the scion () while the lower part is called the rootstock. The suc ...
onto
rootstock A rootstock is part of a plant, often an underground part, from which new above-ground growth can be produced. It could also be described as a stem with a well developed root system, to which a bud from another plant is grafted. It can refer to a ...
s. Apple trees grown without rootstocks tend to be larger and much slower to fruit after planting. Rootstocks are used to control the speed of growth and the size of the resulting tree, allowing for easier harvesting. There are more than 7,500 known cultivars of apples. Different cultivars are bred for various tastes and uses, including
cooking Cooking, cookery, or culinary arts is the art, science and craft of using heat to prepare food for consumption. Cooking techniques and ingredients vary widely, from grilling food over an open fire to using electric stoves, to baking in vario ...
, eating raw, and cider production. Trees and fruit are prone to a number of
fungal A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, separately from t ...
, bacterial, and pest problems, which can be controlled by a number of organic and non-organic means. In 2010, the fruit's
genome In the fields of molecular biology and genetics, a genome is all the genetic information of an organism. It consists of nucleotide sequences of DNA (or RNA in RNA viruses). The nuclear genome includes protein-coding genes and non-coding gen ...
was
sequenced In genetics and biochemistry, sequencing means to determine the primary structure (sometimes incorrectly called the primary sequence) of an unbranched biopolymer. Sequencing results in a symbolic linear depiction known as a sequence which succi ...
as part of research on disease control and selective breeding in apple production. Worldwide production of apples in 2018 was 86 million
tonnes The tonne ( or ; symbol: t) is a unit of mass equal to 1000 kilograms. It is a non-SI unit accepted for use with SI. It is also referred to as a metric ton to distinguish it from the non-metric units of the short ton (United States c ...
, with China accounting for nearly half of the total.


Etymology

The word ''apple'', whose
Old English Old English (, ), or Anglo-Saxon, is the earliest recorded form of the English language, spoken in England and southern and eastern Scotland in the early Middle Ages. It was brought to Great Britain by Anglo-Saxon settlers in the mid-5th c ...
ancestor is , is descended from the
Proto-Germanic Proto-Germanic (abbreviated PGmc; also called Common Germanic) is the reconstructed proto-language of the Germanic branch of the Indo-European languages. Proto-Germanic eventually developed from pre-Proto-Germanic into three Germanic branc ...
noun , descended in turn from
Proto-Indo-European Proto-Indo-European (PIE) is the reconstructed common ancestor of the Indo-European language family. Its proposed features have been derived by linguistic reconstruction from documented Indo-European languages. No direct record of Proto-Indo- ...
. As late as the 17th century, the word also functioned as a generic term for all fruit including nuts—such as the 14th-century
Middle English Middle English (abbreviated to ME) is a form of the English language that was spoken after the Norman conquest of 1066, until the late 15th century. The English language underwent distinct variations and developments following the Old English ...
expression , meaning a
banana A banana is an elongated, edible fruit – botanically a berry – produced by several kinds of large herbaceous flowering plants in the genus ''Musa''. In some countries, bananas used for cooking may be called "plantains", distinguis ...
. This use is analogous to the
French language French ( or ) is a Romance language of the Indo-European family. It descended from the Vulgar Latin of the Roman Empire, as did all Romance languages. French evolved from Gallo-Romance, the Latin spoken in Gaul, and more specifically in No ...
use of .


Description

The apple is a
deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, afte ...
tree, generally standing tall in cultivation and up to in the wild. When cultivated, the size, shape and branch density are determined by
rootstock A rootstock is part of a plant, often an underground part, from which new above-ground growth can be produced. It could also be described as a stem with a well developed root system, to which a bud from another plant is grafted. It can refer to a ...
selection and trimming method. The leaves are alternately arranged dark green-colored simple ovals with serrated margins and slightly downy undersides.
Blossoms In botany, blossoms are the flowers of stone fruit trees (genus ''Prunus'') and of some other plants with a similar appearance that flower profusely for a period of time in spring. Colloquially, flowers of orange are referred to as such as we ...
are produced in
spring Spring(s) may refer to: Common uses * Spring (season), a season of the year * Spring (device), a mechanical device that stores energy * Spring (hydrology), a natural source of water * Spring (mathematics), a geometric surface in the shape of a h ...
simultaneously with the budding of the leaves and are produced on spurs and some long
shoot In botany, a plant shoot consists of any plant stem together with its appendages, leaves and lateral buds, flowering stems, and flower buds. The new growth from seed germination that grows upward is a shoot where leaves will develop. In the spr ...
s. The flowers are white with a pink tinge that gradually fades, five
petal Petals are modified leaves that surround the reproductive parts of flowers. They are often brightly colored or unusually shaped to attract pollinators. All of the petals of a flower are collectively known as the ''corolla''. Petals are usually ...
ed, with an
inflorescence An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are formed on ...
consisting of a cyme with 4–6 flowers. The central flower of the inflorescence is called the "king bloom"; it opens first and can develop a larger fruit. The
fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in particul ...
is a
pome In botany, a pome is a type of fruit produced by flowering plants in the subtribe Malinae of the family Rosaceae. Well-known pomes include the apple, pear, and quince. Etymology The word ''pome'' entered English in the late 14th century, and ...
that matures in late
summer Summer is the hottest of the four temperate seasons, occurring after spring and before autumn. At or centred on the summer solstice, the earliest sunrise and latest sunset occurs, daylight hours are longest and dark hours are shortest, wit ...
or
autumn Autumn, also known as fall in American English and Canadian English, is one of the four temperate seasons on Earth. Outside the tropics, autumn marks the transition from summer to winter, in September (Northern Hemisphere) or March ( South ...
, and cultivars exist in a wide range of sizes. Commercial growers aim to produce an apple that is in diameter, due to market preference. Some consumers, especially those in Japan, prefer a larger apple, while apples below are generally used for making juice and have little fresh market value. The skin of ripe apples is generally red, yellow, green, pink, or
russetted Russeting or russetting is an abnormality of fruit skin which manifests in russet-colored (brownish) patches that are rougher than healthy skin. It is a common feature in apples and pears. Russeting is typically an undesirable trait, which reduce ...
, though many bi- or tri-colored cultivars may be found. The skin may also be wholly or partly russeted i.e. rough and brown. The skin is covered in a protective layer of
epicuticular wax Epicuticular wax is a coating of wax covering the outer surface of the plant cuticle in land plants. It may form a whitish film or bloom on leaves, fruits and other plant organs. Chemically, it consists of hydrophobic organic compounds, mainly st ...
. The exocarp (flesh) is generally pale yellowish-white, though pink, yellow or green exocarps also occur.


Wild ancestors

The original wild ancestor of ''Malus domestica'' was ''Malus sieversii'', found growing wild in the
mountains of Central Asia The Mountains of Central Asia are a biodiversity hot spot designated by Conservation International which covers several montane and alpine ecoregions of Central Asia, including those of the Pamir and Tian Shan ranges, and extending across porti ...
in southern
Kazakhstan Kazakhstan, officially the Republic of Kazakhstan, is a transcontinental country located mainly in Central Asia and partly in Eastern Europe. It borders Russia to the north and west, China to the east, Kyrgyzstan to the southeast, Uzbekis ...
,
Kyrgyzstan Kyrgyzstan,, pronounced or the Kyrgyz Republic, is a landlocked country in Central Asia. Kyrgyzstan is bordered by Kazakhstan to the north, Uzbekistan to the west, Tajikistan to the south, and the People's Republic of China to the east ...
,
Tajikistan Tajikistan (, ; tg, Тоҷикистон, Tojikiston; russian: Таджикистан, Tadzhikistan), officially the Republic of Tajikistan ( tg, Ҷумҳурии Тоҷикистон, Jumhurii Tojikiston), is a landlocked country in Centr ...
, and
northwestern China Northwest China () is a statistical region of China which includes the autonomous regions of Xinjiang and Ningxia and the provinces of Shaanxi, Gansu and Qinghai. It has an area of 3,107,900 km2. The region is characterized by a (semi-)arid c ...
. Cultivation of the species, most likely beginning on the forested flanks of the
Tian Shan The Tian Shan,, , otk, 𐰴𐰣 𐱅𐰭𐰼𐰃, , tr, Tanrı Dağı, mn, Тэнгэр уул, , ug, تەڭرىتاغ, , , kk, Тәңіртауы / Алатау, , , ky, Теңир-Тоо / Ала-Тоо, , , uz, Tyan-Shan / Tangritog‘ ...
mountains, progressed over a long period of time and permitted secondary
introgression Introgression, also known as introgressive hybridization, in genetics is the transfer of genetic material from one species into the gene pool of another by the repeated backcrossing of an interspecific hybrid with one of its parent species. Intro ...
of genes from other species into the open-pollinated seeds. Significant exchange with ''
Malus sylvestris ''Malus sylvestris'', the European crab apple, is a species of the genus ''Malus'', native to Europe. Its scientific name means "forest apple" and the truly wild tree has thorns. Description Wild apple has an expanded crown and often appears mo ...
'', the crabapple, resulted in current populations of apples being more related to crabapples than to the more morphologically similar progenitor ''Malus sieversii''. In strains without recent admixture the contribution of the latter predominates.


Genome

Apples are diploid (though triploid cultivars are not uncommon), have 17 chromosomes and an estimated
genome In the fields of molecular biology and genetics, a genome is all the genetic information of an organism. It consists of nucleotide sequences of DNA (or RNA in RNA viruses). The nuclear genome includes protein-coding genes and non-coding gen ...
size of approximately 650 Mb. Several whole genome sequences have been made available, the first one in 2010 was based on the diploid cultivar '
Golden Delicious 'Golden Delicious' is a cultivar of apple. It is one of the 15 most popular apple cultivars in the United States. It is not closely related to 'Red Delicious'. History Golden Delicious arose from a chance seedling, possibly a hybrid of 'Grim ...
'. However, this first whole genome sequence turned out to contain several errors in part owing to the high degree of heterozygosity in diploid apples which, in combination with an ancient genome duplication, complicated the assembly. Recently, double- and trihaploid individuals have been sequenced, yielding whole genome sequences of higher quality. The first whole genome assembly was estimated to contain around 57,000 genes, though the more recent genome sequences support more moderate estimates between 42,000 and 44,700 protein-coding genes. Among other things, the availability of whole genome sequences has provided evidence that the wild ancestor of the cultivated apple most likely is ''Malus sieversii''. Re-sequencing of multiple accessions has supported this, while also suggesting extensive introgression from ''Malus sylvestris'' following domestication.


History

''
Malus sieversii ''Malus sieversii'' is a wild apple native to the mountains of Central Asia in southern Kazakhstan. It has recently been shown to be the primary ancestor of most cultivars of the domesticated apple (''Malus domestica''). It was first described ...
'' is recognized as a major progenitor species to the cultivated apple, and is morphologically similar. Due to the genetic variability in Central Asia, this region is generally considered the center of origin for apples. The apple is thought to have been domesticated 4000–10000 years ago in the
Tian Shan The Tian Shan,, , otk, 𐰴𐰣 𐱅𐰭𐰼𐰃, , tr, Tanrı Dağı, mn, Тэнгэр уул, , ug, تەڭرىتاغ, , , kk, Тәңіртауы / Алатау, , , ky, Теңир-Тоо / Ала-Тоо, , , uz, Tyan-Shan / Tangritog‘ ...
mountains, and then to have travelled along the
Silk Road The Silk Road () was a network of Eurasian trade routes active from the second century BCE until the mid-15th century. Spanning over 6,400 kilometers (4,000 miles), it played a central role in facilitating economic, cultural, political, and rel ...
to Europe, with hybridization and introgression of wild crabapples from Siberia (''M. baccata''), the Caucasus (''M. orientalis''), and Europe (''M. sylvestris''). Only the ''M. sieversii'' trees growing on the western side of the Tian Shan mountains contributed genetically to the domesticated apple, not the isolated population on the eastern side. Chinese soft apples, such as '' M. asiatica'' and ''
M. prunifolia ''Malus prunifolia'' is a species of crabapple tree known by the common names plumleaf crab apple, plum-leaved apple, pear-leaf crabapple, Chinese apple and Chinese crabapple. It is native to China, and is grown elsewhere for use as an ornamen ...
'', have been cultivated as dessert apples for more than 2000 years in China. These are thought to be hybrids between ''M. baccata'' and ''M. sieversii'' in Kazakhstan. Among the traits selected for by human growers are size, fruit acidity, color, firmness, and soluble sugar. Unusually for domesticated fruits, the wild ''M. sieversii'' origin is only slightly smaller than the modern domesticated apple. At the Sammardenchia-Cueis site near Udine in Northeastern Italy, seeds from some form of apples have been found in material carbon dated to around 4000 BCE. Genetic analysis has not yet been successfully used to determine whether such ancient apples were wild ''Malus sylvestris'' or ''Malus domesticus'' containing ''Malus sieversii'' ancestry. It is generally also hard to distinguish in the archeological record between foraged wild apples and apple plantations. There is indirect evidence of apple cultivation in the third millennium BCE in the
Middle East The Middle East ( ar, الشرق الأوسط, ISO 233: ) is a geopolitical region commonly encompassing Arabia (including the Arabian Peninsula and Bahrain), Asia Minor (Asian part of Turkey except Hatay Province), East Thrace (Europea ...
. There was substantial apple production in the European classical antiquity, and grafting was certainly known then. Grafting is an essential part of modern domesticated apple production, to be able to propagate the best cultivars; it is unclear when apple tree grafting was invented. Winter apples, picked in late autumn and stored just above freezing, have been an important food in Asia and Europe for millennia. Of the many Old World plants that the Spanish introduced to
Chiloé Archipelago The Chiloé Archipelago ( es, Archipiélago de Chiloé, , ) is a group of islands lying off the coast of Chile, in the Los Lagos Region. It is separated from mainland Chile by the Chacao Channel in the north, the Sea of Chiloé in the east and ...
in the 16th century, apple trees became particularly well adapted. Apples were introduced to North America by colonists in the 17th century, and the first apple orchard on the North American continent was planted in
Boston Boston (), officially the City of Boston, is the state capital and most populous city of the Commonwealth of Massachusetts, as well as the cultural and financial center of the New England region of the United States. It is the 24th- most p ...
by Reverend
William Blaxton Reverend William Blaxton (also spelled William Blackstone) (1595 – 26 May 1675) was an early English settler in New England and the first European settler of Boston and Rhode Island. Biography William Blaxton was born in Horncastle, Lincolns ...
in 1625. The only apples native to North America are crab apples, which were once called "common apples". Apple cultivars brought as seed from Europe were spread along Native American trade routes, as well as being cultivated on colonial farms. An 1845 United States apples nursery catalogue sold 350 of the "best" cultivars, showing the proliferation of new North American cultivars by the early 19th century. In the 20th century, irrigation projects in
Eastern Washington Eastern Washington is the region of the U.S. state of Washington located east of the Cascade Range. It contains the city of Spokane (the second largest city in the state), the Tri-Cities, the Columbia River and the Grand Coulee Dam, the Hanfo ...
began and allowed the development of the multibillion-dollar fruit industry, of which the apple is the leading product. Until the 20th century, farmers stored apples in frostproof cellars during the winter for their own use or for sale. Improved transportation of fresh apples by train and road replaced the necessity for storage.
Controlled atmosphere A controlled atmosphere is an agricultural storage method in which the concentrations of oxygen, carbon dioxide and nitrogen, as well as the temperature and humidity of a storage room are regulated. Both dry commodities and fresh fruit and vegetab ...
facilities are used to keep apples fresh year-round. Controlled atmosphere facilities use high humidity, low oxygen, and controlled carbon dioxide levels to maintain fruit freshness. They were first used in the United States in the 1960s.


Significance in European cultures and societies


Germanic paganism

In
Norse mythology Norse, Nordic, or Scandinavian mythology is the body of myths belonging to the North Germanic peoples, stemming from Old Norse religion and continuing after the Christianization of Scandinavia, and into the Nordic folklore of the modern period ...
, the goddess
Iðunn In Norse mythology, Iðunn is a goddess associated with apples and youth. Iðunn is attested in the ''Poetic Edda'', compiled in the 13th century from earlier traditional sources, and the ''Prose Edda'', written in the 13th century by Snorri St ...
is portrayed in the ''
Prose Edda The ''Prose Edda'', also known as the ''Younger Edda'', ''Snorri's Edda'' ( is, Snorra Edda) or, historically, simply as ''Edda'', is an Old Norse textbook written in Iceland during the early 13th century. The work is often assumed to have been ...
'' (written in the 13th century by
Snorri Sturluson Snorri Sturluson ( ; ; 1179 – 22 September 1241) was an Icelandic historian, poet, and politician. He was elected twice as lawspeaker of the Icelandic parliament, the Althing. He is commonly thought to have authored or compiled portions of the ...
) as providing apples to the
gods A deity or god is a supernatural being who is considered divine or sacred. The ''Oxford Dictionary of English'' defines deity as a god or goddess, or anything revered as divine. C. Scott Littleton defines a deity as "a being with powers greater ...
that give them
eternal youth Eternal youth is the concept of human physical immortality free of ageing. The youth referred to is usually meant to be in contrast to the depredations of aging, rather than a specific age of the human lifespan. Eternal youth is common in mytho ...
fulness. The English scholar H. R. Ellis Davidson links apples to religious practices in
Germanic paganism Germanic paganism or Germanic religion refers to the traditional, culturally significant religion of the Germanic peoples. With a chronological range of at least one thousand years in an area covering Scandinavia, the British Isles, modern Germ ...
, from which
Norse paganism Old Norse religion, also known as Norse paganism, is the most common name for a branch of Germanic religion which developed during the Proto-Norse period, when the North Germanic peoples separated into a distinct branch of the Germanic peop ...
developed. She points out that buckets of apples were found in the Oseberg ship burial site in Norway, that fruit and nuts (Iðunn having been described as being transformed into a nut in ''
Skáldskaparmál ''Skáldskaparmál'' (Old Norse: 'The Language of Poetry'; c. 50,000 words; ; ) is the second part of the ''Prose Edda''. The section consists of a dialogue between Ægir, the divine personification of the sea, and Bragi, the god of poetry, in ...
'') have been found in the early graves of the
Germanic peoples The Germanic peoples were historical groups of people that once occupied Central Europe and Scandinavia during antiquity and into the early Middle Ages. Since the 19th century, they have traditionally been defined by the use of ancient and ear ...
in England and elsewhere on the continent of Europe, which may have had a symbolic meaning, and that nuts are still a recognized symbol of fertility in southwest England. Davidson notes a connection between apples and the
Vanir In Norse mythology, the Vanir (; Old Norse: , singular Vanr ) are a group of gods associated with fertility, wisdom, and the ability to see the future. The Vanir are one of two groups of gods (the other being the Æsir) and are the namesake of the ...
, a tribe of gods associated with fertility in Norse mythology, citing an instance of eleven "golden apples" being given to woo the beautiful
Gerðr In Norse mythology, Gerðr (Old Norse: ; "fenced-in"Orchard (1997:54).) is a jötunn, goddess, and the wife of the god Freyr. Gerðr is attested in the ''Poetic Edda'', compiled in the 13th century from earlier traditional sources; the ''Prose ...
by Skírnir, who was acting as messenger for the major Vanir god
Freyr Freyr (Old Norse: 'Lord'), sometimes anglicized as Frey, is a widely attested god in Norse mythology, associated with kingship, fertility, peace, and weather. Freyr, sometimes referred to as Yngvi-Freyr, was especially associated with Sweden an ...
in stanzas 19 and 20 of ''
Skírnismál ''Skírnismál'' (Old Norse: 'The Lay of Skírnir') is one of the poems of the ''Poetic Edda''. It is preserved in the 13th-century manuscripts Codex Regius and AM 748 I 4to but may have been originally composed in the early 10th century. Many ...
''. Davidson also notes a further connection between fertility and apples in Norse mythology in chapter 2 of the ''
Völsunga saga The ''Völsunga saga'' (often referred to in English as the ''Volsunga Saga'' or ''Saga of the Völsungs'') is a legendary saga, a late 13th-century poetic rendition in Old Norse of the origin and decline of the Völsung clan (including the stor ...
'': when the major goddess
Frigg Frigg (; Old Norse: ) is a goddess, one of the Æsir, in Germanic mythology. In Norse mythology, the source of most surviving information about her, she is associated with marriage, prophecy, clairvoyance and motherhood, and dwells in the wet ...
sends King
Rerir In ''Völsunga saga'', Rerir, the son of Sigi, succeeds his murdered father and avenges his death. He rules in Hunaland and becomes a powerful ruler. Rerir's son is Völsung. Rerir and his wife were unable to have children until the goddess Frigg ...
an apple after he prays to Odin for a child, Frigg's messenger (in the guise of a crow) drops the apple in his lap as he sits atop a
mound A mound is a heaped pile of earth, gravel, sand, rocks, or debris. Most commonly, mounds are earthen formations such as hills and mountains, particularly if they appear artificial. A mound may be any rounded area of topographically higher ele ...
. Rerir's wife's consumption of the apple results in a six-year pregnancy and the birth (by
Caesarean section Caesarean section, also known as C-section or caesarean delivery, is the surgical procedure by which one or more babies are delivered through an incision in the mother's abdomen, often performed because vaginal delivery would put the baby or mo ...
) of their son—the hero
Völsung In Norse mythology, Völsung ( non, Vǫlsungr ) was the son of Rerir and the eponymous ancestor of the ill-fated Völsung clan (), which includes the well known Norse hero Sigurð. He was murdered by the Geatish king Siggeir and later avenged by ...
. Further, Davidson points out the "strange" phrase "Apples of Hel" used in an 11th-century poem by the
skald A skald, or skáld (Old Norse: , later ; , meaning "poet"), is one of the often named poets who composed skaldic poetry, one of the two kinds of Old Norse poetry, the other being Eddic poetry, which is anonymous. Skaldic poems were traditionally ...
Thorbiorn Brúnarson. She states this may imply that the apple was thought of by Brúnarson as the food of the dead. Further, Davidson notes that the potentially Germanic goddess
Nehalennia Nehalennia (spelled variously) is a goddess of unclear origin, perhaps Germanic or Celtic. She is attested on and depicted upon numerous votive altars discovered around what is now the province of Zeeland, the Netherlands, where the Schelde Rive ...
is sometimes depicted with apples and that parallels exist in early Irish stories. Davidson asserts that while cultivation of the apple in Northern Europe extends back to at least the time of the
Roman Empire The Roman Empire ( la, Imperium Romanum ; grc-gre, Βασιλεία τῶν Ῥωμαίων, Basileía tôn Rhōmaíōn) was the post-Roman Republic, Republican period of ancient Rome. As a polity, it included large territorial holdings aro ...
and came to Europe from the
Near East The ''Near East''; he, המזרח הקרוב; arc, ܕܢܚܐ ܩܪܒ; fa, خاور نزدیک, Xāvar-e nazdik; tr, Yakın Doğu is a geographical term which roughly encompasses a transcontinental region in Western Asia, that was once the hist ...
, the native varieties of apple trees growing in Northern Europe are small and bitter. Davidson concludes that in the figure of Iðunn "we must have a dim reflection of an old symbol: that of the guardian goddess of the life-giving fruit of the other world."


Greek mythology

Apples appear in many
religious traditions Religion is usually defined as a social-cultural system of designated behaviors and practices, morals, beliefs, worldviews, texts, sanctified places, prophecies, ethics, or organizations, that generally relates humanity to supernatural, tra ...
, often as a mystical or
forbidden fruit Forbidden fruit is a name given to the fruit growing in the Garden of Eden which God commands mankind not to eat. In the biblical story, Adam and Eve eat the fruit from the tree of the knowledge of good and evil and are exiled from Eden. As a ...
. One of the problems identifying apples in religion,
mythology Myth is a folklore genre consisting of narratives that play a fundamental role in a society, such as foundational tales or origin myths. Since "myth" is widely used to imply that a story is not objectively true, the identification of a narrati ...
and folktales is that the word "apple" was used as a generic term for all (foreign) fruit, other than berries, including nuts, as late as the 17th century. For instance, in
Greek mythology A major branch of classical mythology, Greek mythology is the body of myths originally told by the ancient Greeks, and a genre of Ancient Greek folklore. These stories concern the origin and nature of the world, the lives and activities of de ...
, the
Greek hero Hero cults were one of the most distinctive features of ancient Greek religion. In Homeric Greek, "hero" (, ) refers to the mortal offspring of a human and a god. By the historical period, however, the word came to mean specifically a ''dead'' ma ...
Heracles Heracles ( ; grc-gre, Ἡρακλῆς, , glory/fame of Hera), born Alcaeus (, ''Alkaios'') or Alcides (, ''Alkeidēs''), was a divine hero in Greek mythology, the son of Zeus and Alcmene, and the foster son of Amphitryon.By his adoptive ...
, as a part of his
Twelve Labours The Labours of Hercules or Labours of Heracles ( grc-gre, οἱ Ἡρακλέους ἆθλοι, ) are a series of episodes concerning a penance carried out by Heracles, the greatest of the Greek heroes, whose name was later romanised a ...
, was required to travel to the Garden of the Hesperides and pick the golden apples off the
Tree of Life The tree of life is a fundamental archetype in many of the world's mythological, religious, and philosophical traditions. It is closely related to the concept of the sacred tree.Giovino, Mariana (2007). ''The Assyrian Sacred Tree: A History ...
growing at its center. The Greek goddess of discord, Eris, became disgruntled after she was excluded from the wedding of
Peleus In Greek mythology, Peleus (; Ancient Greek: Πηλεύς ''Pēleus'') was a hero, king of Phthia, husband of Thetis and the father of their son Achilles. This myth was already known to the hearers of Homer in the late 8th century BC. Biogra ...
and
Thetis Thetis (; grc-gre, Θέτις ), is a figure from Greek mythology with varying mythological roles. She mainly appears as a sea nymph, a goddess of water, or one of the 50 Nereids, daughters of the ancient sea god Nereus. When described as ...
. In retaliation, she tossed a
golden apple The golden apple is an element that appears in various national and ethnic folk legends or fairy tales. Recurring themes depict a hero (for example Hercules or Făt-Frumos) retrieving the golden apples hidden or stolen by a monstrous antagonist. ...
inscribed Καλλίστη (''Kalliste'', sometimes transliterated ''Kallisti'', "For the most beautiful one"), into the wedding party. Three goddesses claimed the apple:
Hera In ancient Greek religion, Hera (; grc-gre, Ἥρα, Hḗrā; grc, Ἥρη, Hḗrē, label=none in Ionic and Homeric Greek) is the goddess of marriage, women and family, and the protector of women during childbirth. In Greek mythology, she ...
,
Athena Athena or Athene, often given the epithet Pallas, is an ancient Greek goddess associated with wisdom, warfare, and handicraft who was later syncretized with the Roman goddess Minerva. Athena was regarded as the patron and protectress of ...
, and
Aphrodite Aphrodite ( ; grc-gre, Ἀφροδίτη, Aphrodítē; , , ) is an ancient Greek goddess associated with love, lust, beauty, pleasure, passion, and procreation. She was syncretized with the Roman goddess . Aphrodite's major symbols include ...
.
Paris Paris () is the capital and most populous city of France, with an estimated population of 2,165,423 residents in 2019 in an area of more than 105 km² (41 sq mi), making it the 30th most densely populated city in the world in 2020. S ...
of
Troy Troy ( el, Τροία and Latin: Troia, Hittite: 𒋫𒊒𒄿𒊭 ''Truwiša'') or Ilion ( el, Ίλιον and Latin: Ilium, Hittite: 𒃾𒇻𒊭 ''Wiluša'') was an ancient city located at Hisarlik in present-day Turkey, south-west of Çan ...
was appointed to select the recipient. After being bribed by both Hera and Athena, Aphrodite tempted him with the most beautiful woman in the world, Helen of
Sparta Sparta (Doric Greek: Σπάρτα, ''Spártā''; Attic Greek: Σπάρτη, ''Spártē'') was a prominent city-state in Laconia, in ancient Greece. In antiquity, the city-state was known as Lacedaemon (, ), while the name Sparta referred t ...
. He awarded the apple to Aphrodite, thus indirectly causing the
Trojan War In Greek mythology, the Trojan War was waged against the city of Troy by the Achaeans (Greeks) after Paris of Troy took Helen from her husband Menelaus, king of Sparta. The war is one of the most important events in Greek mythology and has ...
. The apple was thus considered, in ancient Greece, sacred to Aphrodite. To throw an apple at someone was to symbolically declare one's love; and similarly, to catch it was to symbolically show one's acceptance of that love. An epigram claiming authorship by Plato states:
Atalanta Atalanta (; grc-gre, Ἀταλάντη, Atalantē) meaning "equal in weight", is a heroine in Greek mythology. There are two versions of the huntress Atalanta: one from Arcadia, whose parents were Iasus and Clymene and who is primarily know ...
, also of Greek mythology, raced all her suitors in an attempt to avoid marriage. She outran all but
Hippomenes :''The name Hippomenes may also refer to the father of Leimone.'' In Greek mythology, Hippomenes (; grc, Ἱππομένης), also known as Melanion (; Μελανίων or Μειλανίων), was a son of the Arcadian AmphidamasApollodorus, ...
(also known as
Melanion :''The name Hippomenes may also refer to the father of Leimone.'' In Greek mythology, Hippomenes (; grc, Ἱππομένης), also known as Melanion (; Μελανίων or Μειλανίων), was a son of the Arcadian AmphidamasApollodorus, ...
, a name possibly derived from ''melon'', the Greek word for both "apple" and fruit in general), who defeated her by cunning, not speed. Hippomenes knew that he could not win in a fair race, so he used three golden apples (gifts of Aphrodite, the goddess of love) to distract Atalanta. It took all three apples and all of his speed, but Hippomenes was finally successful, winning the race and Atalanta's hand.


Christian art

Though the forbidden fruit of Eden in the
Book of Genesis The Book of Genesis (from Greek ; Hebrew: בְּרֵאשִׁית ''Bəreʾšīt'', "In hebeginning") is the first book of the Hebrew Bible and the Christian Old Testament. Its Hebrew name is the same as its first word, ( "In the beginning"). ...
is not identified, popular Christian tradition has held that it was an apple that
Eve Eve (; ; ar, حَوَّاء, Ḥawwāʾ; el, Εὕα, Heúa; la, Eva, Heva; Syriac: romanized: ) is a figure in the Book of Genesis in the Hebrew Bible. According to the origin story, "Creation myths are symbolic stories describing how the ...
coaxed
Adam Adam; el, Ἀδάμ, Adám; la, Adam is the name given in Genesis 1-5 to the first human. Beyond its use as the name of the first man, ''adam'' is also used in the Bible as a pronoun, individually as "a human" and in a collective sense as ...
to share with her. The origin of the popular identification with a fruit unknown in the Middle East in biblical times is found in confusion between the
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
words ''mālum'' (an apple) and ''mălum'' (an evil), each of which is normally written ''malum''. The tree of the forbidden fruit is called "the tree of the knowledge of good and evil" in Genesis 2:17, and the Latin for "good and evil" is ''bonum et malum''.
Renaissance The Renaissance ( , ) , from , with the same meanings. is a period in European history marking the transition from the Middle Ages to modernity and covering the 15th and 16th centuries, characterized by an effort to revive and surpass ideas ...
painters may also have been influenced by the story of the
golden apple The golden apple is an element that appears in various national and ethnic folk legends or fairy tales. Recurring themes depict a hero (for example Hercules or Făt-Frumos) retrieving the golden apples hidden or stolen by a monstrous antagonist. ...
s in the
Garden of Hesperides In Greek mythology, the Hesperides (; , ) are the nymphs of evening and golden light of sunsets, who were the "Daughters of the Evening" or "Nymphs of the West". They were also called the Atlantides () from their reputed father, the Titan Atlas ...
. As a result, in the story of Adam and Eve, the apple became a symbol for knowledge, immortality, temptation, the fall of man into sin, and sin itself. The
larynx The larynx (), commonly called the voice box, is an organ in the top of the neck involved in breathing, producing sound and protecting the trachea against food aspiration. The opening of larynx into pharynx known as the laryngeal inlet is about ...
in the human throat has been called the "
Adam's apple The Adam's apple or laryngeal prominence is the protrusion in the human neck formed by the angle of the thyroid cartilage surrounding the larynx, typically visible in men, less frequently in women. Structure The topographic structure which is e ...
" because of a notion that it was caused by the forbidden fruit remaining in the throat of Adam. The apple as symbol of sexual
seduction Seduction has multiple meanings. Platonically, it can mean "to persuade to disobedience or disloyalty", or "to lead astray, usually by persuasion or false promises". Strategies of seduction include conversation and sexual scripts, paralingual ...
has been used to imply human sexuality, possibly in an ironic vein.


Proverb

The proverb, " An apple a day keeps the doctor away", addressing the supposed health benefits of the fruit, has been traced to 19th-century
Wales Wales ( cy, Cymru ) is a country that is part of the United Kingdom. It is bordered by England to the east, the Irish Sea to the north and west, the Celtic Sea to the south west and the Bristol Channel to the south. It had a population in 2 ...
, where the original phrase was "Eat an apple on going to bed, and you'll keep the doctor from earning his bread". In the 19th century and early 20th, the phrase evolved to "an apple a day, no doctor to pay" and "an apple a day sends the doctor away"; the phrasing now commonly used was first recorded in 1922.


Cultivars

There are more than 7,500 known cultivars (cultivated varieties) of apples. Cultivars vary in their yield and the ultimate size of the tree, even when grown on the same
rootstock A rootstock is part of a plant, often an underground part, from which new above-ground growth can be produced. It could also be described as a stem with a well developed root system, to which a bud from another plant is grafted. It can refer to a ...
. Different cultivars are available for
temperate In geography, the temperate climates of Earth occur in the middle latitudes (23.5° to 66.5° N/S of Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ranges throughout t ...
and
subtropical The subtropical zones or subtropics are geographical and climate zones to the north and south of the tropics. Geographically part of the temperate zones of both hemispheres, they cover the middle latitudes from to approximately 35° north and ...
climates. The UK's National Fruit Collection, which is the responsibility of the Department of Environment, Food, and Rural Affairs, includes a collection of over 2,000 cultivars of apple tree in Kent. The
University of Reading The University of Reading is a public university in Reading, Berkshire, England. It was founded in 1892 as University College, Reading, a University of Oxford extension college. The institution received the power to grant its own degrees in 192 ...
, which is responsible for developing the UK national collection database, provides access to search the national collection. The University of Reading's work is part of the European Cooperative Programme for Plant Genetic Resources of which there are 38 countries participating in the Malus/Pyrus work group. The UK's national fruit collection database contains much information on the characteristics and origin of many apples, including alternative names for what is essentially the same "genetic" apple cultivar. Most of these cultivars are bred for eating fresh (dessert apples), though some are cultivated specifically for cooking (
cooking apple A cooking apple or culinary apple is an apple that is used primarily for cooking, as opposed to a '' dessert apple'', which is eaten raw. Cooking apples are generally larger, and can be tarter than dessert varieties. Some varieties have a f ...
s) or producing cider.
Cider apple Cider apples are a group of apple cultivars grown for their use in the production of cider (referred to as "hard cider" in the United States). Cider apples are distinguished from "cookers" and "eaters", or dessert apples, by their bitterness or ...
s are typically too tart and astringent to eat fresh, but they give the beverage a rich flavor that dessert apples cannot. Commercially popular apple cultivars are soft but crisp. Other desirable qualities in modern commercial apple breeding are a colorful skin, absence of
russeting Russeting or russetting is an abnormality of fruit skin which manifests in russet-colored (brownish) patches that are rougher than healthy skin. It is a common feature in apples and pears. Russeting is typically an undesirable trait, which reduce ...
, ease of shipping, lengthy storage ability, high yields, disease resistance, common apple shape, and developed flavor. Modern apples are generally sweeter than older cultivars, as popular tastes in apples have varied over time. Most North Americans and Europeans favor sweet, subacid apples, but tart apples have a strong minority following. Extremely sweet apples with barely any acid flavor are popular in Asia, especially the
Indian subcontinent The Indian subcontinent is a physiographical region in Southern Asia. It is situated on the Indian Plate, projecting southwards into the Indian Ocean from the Himalayas. Geopolitically, it includes the countries of Bangladesh, Bhutan, India ...
. Old cultivars are often oddly shaped, russeted, and grow in a variety of textures and colors. Some find them to have better flavor than modern cultivars, but they may have other problems that make them commercially unviable—low yield, disease susceptibility, poor tolerance for storage or transport, or just being the "wrong" size. A few old cultivars are still produced on a large scale, but many have been preserved by home gardeners and farmers that sell directly to local markets. Many unusual and locally important cultivars with their own unique taste and appearance exist; apple conservation campaigns have sprung up around the world to preserve such local cultivars from extinction. In the United Kingdom, old cultivars such as '
Cox's Orange Pippin Cox's Orange Pippin, in Britain often referred to simply as Cox, is an apple cultivar first grown in 1825, at Colnbrook in Buckinghamshire, England, by the retired brewer and horticulturist Richard Cox. Though the parentage of the cultivar is ...
' and ' Egremont Russet' are still commercially important even though by modern standards they are low yielding and susceptible to disease. File:Alice (apple).jpg, 'Alice' File:Ambrosia apples 2017 A3.jpg, '
Ambrosia In the ancient Greek myths, ''ambrosia'' (, grc, ἀμβροσία 'immortality'), the food or drink of the Greek gods, is often depicted as conferring longevity or immortality upon whoever consumed it. It was brought to the gods in Olympus by ...
' File:Malus-Ananasrenette.jpg, 'Ananasrenette' File:Arkansas Black apples (cropped).jpg, 'Arkansas Black' File:Aroma (apple).jpg, 'Aroma' File:Malus-Boskoop organic.jpg, '
Belle de Boskoop Belle de Boskoop (also called Goudrenet, Goudreinet or Goudreinnette) is an apple cultivar which originated in Boskoop, Netherlands, where it began as a chance seedling in 1856. Variants include Boskoop red, yellow and green. This rustic apple is ...
' File:Bramley's Seedling Apples.jpg, 'Bramley' File:Cox orange renette2.JPG, '
Cox's Orange Pippin Cox's Orange Pippin, in Britain often referred to simply as Cox, is an apple cultivar first grown in 1825, at Colnbrook in Buckinghamshire, England, by the retired brewer and horticulturist Richard Cox. Though the parentage of the cultivar is ...
' File:Cox’s Pomona.jpg, 'Cox Pomona' File:Pink Lady Apple (4107712628).jpg, '
Cripps Pink Cripps Pink is a cultivar of apple. It is one of several cultivars sold under the trade mark name . It was originally bred by John Cripps at the Western Australia Department of Agriculture (Stoneville Research Station), by crossing the Austra ...
' File:Discovery apples.jpg, '
Discovery Discovery may refer to: * Discovery (observation), observing or finding something unknown * Discovery (fiction), a character's learning something unknown * Discovery (law), a process in courts of law relating to evidence Discovery, The Discovery ...
' File:Cross section of Egremont Russet (EMLA 1), National Fruit Collection (acc. 1979-159).jpg, ' Egremont Russet' File:Fuji apple.jpg, ' Fuji' File:Red Apple.jpg, '
Gala Gala may refer to: Music * ''Gala'' (album), a 1990 album by the English alternative rock band Lush *'' Gala – The Collection'', a 2016 album by Sarah Brightman * GALA Choruses, an association of LGBT choral groups *''Gala'', a 1986 album by ...
' File:Jabuke Gloster na stablu.2c.jpg, ' Gloster' File:Golden Delicious apple 2021 (2).jpg, '
Golden Delicious 'Golden Delicious' is a cultivar of apple. It is one of the 15 most popular apple cultivars in the United States. It is not closely related to 'Red Delicious'. History Golden Delicious arose from a chance seedling, possibly a hybrid of 'Grim ...
' File:Apfel-Berlepsch.jpg, 'Goldrenette' ( Reinette) File:Granny smith closeup.jpg, '
Granny Smith The Granny Smith, also known as a green apple or sour apple, is an apple cultivar which originated in Australia in 1868. It is named after Maria Ann Smith, who propagated the cultivar from a chance seedling. The tree is thought to be a hybrid ...
' File:Honeycrisp.jpg, '
Honeycrisp Honeycrisp (''Malus pumila'') is an apple cultivar (cultivated variety) developed at the Minnesota Agricultural Experiment Station's Horticultural Research Center at the University of Minnesota, Twin Cities. Designated in 1974 with the MN 1711 ...
' File:Malus-James-Grieve.jpg, 'James Grieve' File:Jonagold.jpg, ' Jonagold' File:Malus Lobo 4397.jpg, 'Lobo' File:McIntosh.jpg, '
McIntosh McIntosh, Macintosh, or Mackintosh ( Gaelic: ') may refer to: Products and brands * Mackintosh, a form of waterproof raincoat * Mackintosh's or John Mackintosh and Co., later Rowntree Mackintosh, former UK confectionery company now part of Nestl� ...
' File:Apple 03.jpg, '
Sciros Sciros (or Pacific Rose) is a cultivar of domesticated apple. This apple is mostly sweet with very little acidity, often compared to the 'Fuji' apple for taste, and keeps very well in storage. According to Orange Pippin it is an attractive new ...
' File:Red Delicious.jpg, '
Red Delicious Red Delicious is a type of apple with a red exterior and sweet taste that was first recognized in Madison County, Iowa, in 1872. Today, the name ''Red Delicious'' comprises more than 50 cultivars. From 1968 to 2018, it was the most produced cu ...
' File:Shampion 13.01.2013 13-17-30.jpg, ' Sampion' (Shampion) File:Mele stark.jpg, 'Stark Delicious' File:The SugarBee Apple now grown in Washington State.jpg, ' SugarBee' File:Summerred.jpg, 'Summerred' File:Tellissaare.JPG, 'Tellissaare' File:Yellow Transparent.jpg, 'Yellow Transparent'


Cultivation


Breeding

Many apples grow readily from seeds. However, more than with most perennial fruits, apples must be propagated asexually to obtain the sweetness and other desirable characteristics of the parent. This is because seedling apples are an example of " extreme heterozygotes", in that rather than inheriting genes from their parents to create a new apple with parental characteristics, they are instead significantly different from their parents, perhaps to compete with the many pests.
Triploid Polyploidy is a condition in which the cells of an organism have more than one pair of ( homologous) chromosomes. Most species whose cells have nuclei (eukaryotes) are diploid, meaning they have two sets of chromosomes, where each set contains ...
cultivars have an additional reproductive barrier in that three sets of chromosomes cannot be divided evenly during meiosis, yielding unequal segregation of the chromosomes (aneuploids). Even in the case when a triploid plant can produce a seed (apples are an example), it occurs infrequently, and seedlings rarely survive. Because apples are not true breeders when planted as seeds, although
cutting Cutting is the separation or opening of a physical object, into two or more portions, through the application of an acutely directed force. Implements commonly used for cutting are the knife and saw, or in medicine and science the scalpel and ...
s can take root and breed true, and may live for a century,
grafting Grafting or graftage is a horticultural technique whereby tissues of plants are joined so as to continue their growth together. The upper part of the combined plant is called the scion () while the lower part is called the rootstock. The suc ...
is usually used. The
rootstock A rootstock is part of a plant, often an underground part, from which new above-ground growth can be produced. It could also be described as a stem with a well developed root system, to which a bud from another plant is grafted. It can refer to a ...
used for the bottom of the graft can be selected to produce trees of a large variety of sizes, as well as changing the winter hardiness, insect and disease resistance, and soil preference of the resulting tree. Dwarf rootstocks can be used to produce very small trees (less than high at maturity), which bear fruit many years earlier in their life cycle than full size trees, and are easier to harvest. Dwarf rootstocks for apple trees can be traced as far back as 300 BCE, to the area of
Persia Iran, officially the Islamic Republic of Iran, and also called Persia, is a country located in Western Asia. It is bordered by Iraq and Turkey to the west, by Azerbaijan and Armenia to the northwest, by the Caspian Sea and Turkmeni ...
and
Asia Minor Anatolia, tr, Anadolu Yarımadası), and the Anatolian plateau, also known as Asia Minor, is a large peninsula in Western Asia and the westernmost protrusion of the Asian continent. It constitutes the major part of modern-day Turkey. The re ...
.
Alexander the Great Alexander III of Macedon ( grc, Ἀλέξανδρος, Alexandros; 20/21 July 356 BC – 10/11 June 323 BC), commonly known as Alexander the Great, was a king of the ancient Greek kingdom of Macedon. He succeeded his father Philip II to ...
sent samples of dwarf apple trees to Aristotle's
Lyceum The lyceum is a category of educational institution defined within the education system of many countries, mainly in Europe. The definition varies among countries; usually it is a type of secondary school. Generally in that type of school the ...
. Dwarf rootstocks became common by the 15th century and later went through several cycles of popularity and decline throughout the world. The majority of the rootstocks used today to control size in apples were developed in England in the early 1900s. The
East Malling Research Station NIAB EMR is a horticultural and agricultural research institute at East Malling, Kent in England, with a specialism in fruit and clonally propagated crop production. In 2016, the institute became part of the NIAB Group. History A research ...
conducted extensive research into rootstocks, and today their rootstocks are given an "M" prefix to designate their origin. Rootstocks marked with an "MM" prefix are Malling-series cultivars later crossed with trees of '
Northern Spy The Northern Spy, also called 'Spy' and 'King', is a cultivar of domesticated apple that originated on the farm of Oliver Chapin in East Bloomfield, New York, in about 1840. It is popular in upstate New York. The Northern Spy was one of four app ...
' in Merton, England. Most new apple cultivars originate as seedlings, which either arise by chance or are bred by deliberately crossing cultivars with promising characteristics. The words "seedling", "pippin", and "kernel" in the name of an apple cultivar suggest that it originated as a seedling. Apples can also form
bud sport In botany, a sport or bud sport, traditionally called ''lusus'', is a part of a plant that shows morphological differences from the rest of the plant. Sports may differ by foliage shape or color, flowers, fruit, or branch structure. The cause is ...
s (mutations on a single branch). Some bud sports turn out to be improved strains of the parent cultivar. Some differ sufficiently from the parent tree to be considered new cultivars. Since the 1930s, the Excelsior Experiment Station at the
University of Minnesota The University of Minnesota, formally the University of Minnesota, Twin Cities, (UMN Twin Cities, the U of M, or Minnesota) is a public land-grant research university in the Twin Cities of Minneapolis and Saint Paul, Minnesota, United States. T ...
has introduced a steady progression of important apples that are widely grown, both commercially and by local orchardists, throughout
Minnesota Minnesota () is a state in the upper midwestern region of the United States. It is the 12th largest U.S. state in area and the 22nd most populous, with over 5.75 million residents. Minnesota is home to western prairies, now given over to ...
and
Wisconsin Wisconsin () is a state in the upper Midwestern United States. Wisconsin is the 25th-largest state by total area and the 20th-most populous. It is bordered by Minnesota to the west, Iowa to the southwest, Illinois to the south, Lake Michi ...
. Its most important contributions have included ' Haralson' (which is the most widely cultivated apple in Minnesota), '
Wealthy Wealth is the abundance of valuable financial assets or physical possessions which can be converted into a form that can be used for transactions. This includes the core meaning as held in the originating Old English word , which is from an I ...
', 'Honeygold', and '
Honeycrisp Honeycrisp (''Malus pumila'') is an apple cultivar (cultivated variety) developed at the Minnesota Agricultural Experiment Station's Horticultural Research Center at the University of Minnesota, Twin Cities. Designated in 1974 with the MN 1711 ...
'. Apples have been acclimatized in Ecuador at very high altitudes, where they can often, with the needed factors, provide crops twice per year because of constant temperate conditions year-round.


Pollination

Apples are self-incompatible; they must
cross-pollinate Pollination is the transfer of pollen from an anther of a plant to the stigma of a plant, later enabling fertilisation and the production of seeds, most often by an animal or by wind. Pollinating agents can be animals such as insects, birds, a ...
to develop fruit. During the flowering each season, apple growers often utilize
pollinator A pollinator is an animal that moves pollen from the male anther of a flower to the female stigma of a flower. This helps to bring about fertilization of the ovules in the flower by the male gametes from the pollen grains. Insects are the maj ...
s to carry pollen.
Honey bee A honey bee (also spelled honeybee) is a eusocial flying insect within the genus ''Apis'' of the bee clade, all native to Afro-Eurasia. After bees spread naturally throughout Africa and Eurasia, humans became responsible for the current cosmop ...
s are most commonly used. Orchard mason bees are also used as supplemental pollinators in commercial orchards.
Bumblebee A bumblebee (or bumble bee, bumble-bee, or humble-bee) is any of over 250 species in the genus ''Bombus'', part of Apidae, one of the bee families. This genus is the only extant group in the tribe Bombini, though a few extinct related gener ...
queens Queens is a borough of New York City, coextensive with Queens County, in the U.S. state of New York. Located on Long Island, it is the largest New York City borough by area. It is bordered by the borough of Brooklyn at the western tip of Long ...
are sometimes present in orchards, but not usually in sufficient number to be significant pollinators.Adamson, Nancy Lee
An Assessment of Non-Apis Bees as Fruit and Vegetable Crop Pollinators in Southwest Virginia
. Diss. 2011. Web. 15 October 2015.
There are four to seven pollination groups in apples, depending on climate: * Group A – Early flowering, 1 to 3 May in England ('
Gravenstein Gravenstein (Danish: ''Gråsten'', meaning "graystone", after Gråsten Palace) is a triploid apple cultivar that originated in the 17th century or earlier. The fruit has a tart flavor, and it is heavily used as a cooking apple, especially f ...
', 'Red Astrachan') * Group B – 4 to 7 May ('
Idared Idared is a type of apple cultivar from Moscow, Idaho, United States. Variety is characterized by a non-uniform skin color. First developed at the University of Idaho Agricultural Experiment Station in 1942; it is a cross between two apple v ...
', '
McIntosh McIntosh, Macintosh, or Mackintosh ( Gaelic: ') may refer to: Products and brands * Mackintosh, a form of waterproof raincoat * Mackintosh's or John Mackintosh and Co., later Rowntree Mackintosh, former UK confectionery company now part of Nestl� ...
') * Group C – Mid-season flowering, 8 to 11 May ('
Granny Smith The Granny Smith, also known as a green apple or sour apple, is an apple cultivar which originated in Australia in 1868. It is named after Maria Ann Smith, who propagated the cultivar from a chance seedling. The tree is thought to be a hybrid ...
', '
Cox's Orange Pippin Cox's Orange Pippin, in Britain often referred to simply as Cox, is an apple cultivar first grown in 1825, at Colnbrook in Buckinghamshire, England, by the retired brewer and horticulturist Richard Cox. Though the parentage of the cultivar is ...
') * Group D – Mid/late season flowering, 12 to 15 May ('
Golden Delicious 'Golden Delicious' is a cultivar of apple. It is one of the 15 most popular apple cultivars in the United States. It is not closely related to 'Red Delicious'. History Golden Delicious arose from a chance seedling, possibly a hybrid of 'Grim ...
', 'Calville blanc d'hiver') * Group E – Late flowering, 16 to 18 May ('
Braeburn The Braeburn is a cultivar of apple that is firm to the touch with a red/orange vertical streaky appearance on a yellow/green background. Its color intensity varies with different growing conditions. It was discovered as a chance seedling in 1 ...
', 'Reinette d'Orléans') * Group F – 19 to 23 May ('Suntan') * Group H – 24 to 28 May ('Court-Pendu Gris' – also called Court-Pendu plat) One cultivar can be pollinated by a compatible cultivar from the same group or close (A with A, or A with B, but not A with C or D). Cultivars are sometimes classified by the day of peak bloom in the average 30-day blossom period, with pollenizers selected from cultivars within a 6-day overlap period.


Maturation and harvest

Cultivars vary in their yield and the ultimate size of the tree, even when grown on the same rootstock. Some cultivars, if left unpruned, grow very large—letting them bear more fruit, but making harvesting more difficult. Depending on tree density (number of trees planted per unit surface area), mature trees typically bear of apples each year, though productivity can be close to zero in poor years. Apples are harvested using three-point ladders that are designed to fit amongst the branches. Trees grafted on dwarfing rootstocks bear about of fruit per year. Farms with apple orchards open them to the public so consumers can pick their own apples. Crops ripen at different times of the year according to the cultivar. Cultivar that yield their crop in the summer include 'Gala', 'Golden Supreme', 'McIntosh', 'Transparent', 'Primate', '
Sweet Bough 'Sweet Bough' is an early ripening cultivar of domesticated apple also known by various other names including 'August Sweeting', 'Early Yellow Bough', and 'White Sugar'. It is also incorrectly known as 'Washington'. References Apple cultiva ...
', and 'Duchess'; fall producers include 'Fuji', 'Jonagold', 'Golden Delicious', 'Red Delicious', 'Chenango', 'Gravenstein', 'Wealthy', 'McIntosh', 'Snow', and 'Blenheim'; winter producers include 'Winesap', 'Granny Smith', 'King', 'Wagener', ' Swayzie', 'Greening', and 'Tolman Sweet'.


Storage

Commercially, apples can be stored for a few months in
controlled atmosphere A controlled atmosphere is an agricultural storage method in which the concentrations of oxygen, carbon dioxide and nitrogen, as well as the temperature and humidity of a storage room are regulated. Both dry commodities and fresh fruit and vegetab ...
chambers to delay
ethylene Ethylene ( IUPAC name: ethene) is a hydrocarbon which has the formula or . It is a colourless, flammable gas with a faint "sweet and musky" odour when pure. It is the simplest alkene (a hydrocarbon with carbon-carbon double bonds). Ethylene ...
-induced ripening. Apples are commonly stored in chambers with higher concentrations of carbon dioxide and high air filtration. This prevents ethylene concentrations from rising to higher amounts and preventing ripening from occurring too quickly. For home storage, most cultivars of apple can be held for approximately two weeks when kept at the coolest part of the refrigerator (i.e. below 5 °C). Some can be stored up to a year without significant degradation. Some varieties of apples (e.g. '
Granny Smith The Granny Smith, also known as a green apple or sour apple, is an apple cultivar which originated in Australia in 1868. It is named after Maria Ann Smith, who propagated the cultivar from a chance seedling. The tree is thought to be a hybrid ...
' and ' Fuji') have more than three times the storage life of others. Non-organic apples may be sprayed with a substance 1-methylcyclopropene blocking the apples' ethylene receptors, temporarily preventing them from ripening.


Pests and diseases

Apple trees are susceptible to a number of
fungal A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, separately from t ...
and
bacterial Bacteria (; singular: bacterium) are ubiquitous, mostly free-living organisms often consisting of one biological cell. They constitute a large domain of prokaryotic microorganisms. Typically a few micrometres in length, bacteria were among ...
diseases and insect pests. Many commercial orchards pursue a program of chemical sprays to maintain high fruit quality, tree health, and high yields. These prohibit the use of synthetic pesticides, though some older pesticides are allowed. Organic methods include, for instance, introducing its natural predator to reduce the population of a particular pest. A wide range of pests and diseases can affect the plant. Three of the more common diseases or pests are mildew, aphids, and apple scab. *
Mildew Mildew is a form of fungus. It is distinguished from its closely related counterpart, mould, largely by its colour: moulds appear in shades of black, blue, red, and green, whereas mildew is white. It appears as a thin, superficial growth consi ...
is characterized by light grey powdery patches appearing on the leaves, shoots and flowers, normally in spring. The flowers turn a creamy yellow color and do not develop correctly. This can be treated similarly to ''Botrytis''—eliminating the conditions that caused the disease and burning the infected plants are among recommended actions. *
Aphids Aphids are small sap-sucking insects and members of the superfamily Aphidoidea. Common names include greenfly and blackfly, although individuals within a species can vary widely in color. The group includes the fluffy white woolly aphids. A t ...
are a small insect. Five species of aphids commonly attack apples: apple grain aphid, rosy apple aphid, apple aphid, spirea aphid, and the woolly apple aphid. The aphid species can be identified by color, time of year, and by differences in the cornicles (small paired projections from their rear). Aphids feed on foliage using needle-like mouth parts to suck out plant juices. When present in high numbers, certain species reduce tree growth and vigor. *
Apple scab Apple scab is a common disease of plants in the rose family (Rosaceae) that is caused by the ascomycete fungus '' Venturia inaequalis''. While this disease affects several plant genera, including ''Sorbus, Cotoneaster,'' and '' Pyrus'', it is m ...
: Apple scab causes leaves to develop olive-brown spots with a velvety texture that later turn brown and become cork-like in texture. The disease also affects the fruit, which also develops similar brown spots with velvety or cork-like textures. Apple scab is spread through fungus growing in old apple leaves on the ground and spreads during warm spring weather to infect the new year's growth. Among the most serious disease problems is a bacterial disease called
fireblight Fire blight, also written fireblight, is a contagious disease affecting apples, pears, and some other members of the family Rosaceae. It is a serious concern to apple and pear producers. Under optimal conditions, it can destroy an entire orchard ...
, and two fungal diseases: ''
Gymnosporangium ''Gymnosporangium'' is a genus of heteroecious plant-pathogenic fungi which alternately infect members of the family Cupressaceae, primarily species in the genus ''Juniperus'' (junipers), and members of the family Rosaceae in the subfamily Amygd ...
'' rust and black spot. Other pests that affect apple trees include
Codling moth The codling moth (''Cydia pomonella'') is a member of the Lepidopteran family Tortricidae. They are major pests to agricultural crops, mainly fruits such as apples and pears. Because the larvae are not able to feed on leaves, they are highly de ...
s and
apple maggot The apple maggot (''Rhagoletis pomonella''), also known as the railroad worm (but distinct from the '' Phrixothrix'' beetle larva, also called railroad worm), is a species of fruit fly, and a pest of several types of fruits, especially apples. ...
s. Young apple trees are also prone to mammal pests like mice and deer, which feed on the soft bark of the trees, especially in winter. The larvae of the apple clearwing moth (red-belted clearwing) burrow through the bark and into the phloem of apple trees, potentially causing significant damage.


Production

World production of apples in 2020 was 86 million
tonne The tonne ( or ; symbol: t) is a unit of mass equal to 1000 kilograms. It is a non-SI unit accepted for use with SI. It is also referred to as a metric ton to distinguish it from the non-metric units of the short ton (United States c ...
s, with China producing 47% of the total (table). Secondary producers were the United States and Turkey.


Nutrition

A raw apple is 86% water and 14% carbohydrates, with negligible content of
fat In nutrition, biology, and chemistry, fat usually means any ester of fatty acids, or a mixture of such compounds, most commonly those that occur in living beings or in food. The term often refers specifically to triglycerides (triple est ...
and protein (table). A reference serving of a raw apple with skin weighing 100 grams provides 52
calorie The calorie is a unit of energy. For historical reasons, two main definitions of "calorie" are in wide use. The large calorie, food calorie, or kilogram calorie was originally defined as the amount of heat needed to raise the temperature of on ...
s and a moderate content of
dietary fiber Dietary fiber (in British English fibre) or roughage is the portion of plant-derived food that cannot be completely broken down by human digestive enzymes. Dietary fibers are diverse in chemical composition, and can be grouped generally by the ...
. Otherwise, there is low content of
micronutrient Micronutrients are essential dietary elements required by organisms in varying quantities throughout life to orchestrate a range of physiological functions to maintain health. Micronutrient requirements differ between organisms; for example, huma ...
s, with the
Daily Value The Reference Daily Intake (RDI) used in nutrition labeling on food and dietary supplement products in the U.S. and Canada is the daily intake level of a nutrient that is considered to be sufficient to meet the requirements of 97–98% of healthy ...
s of all falling below 10%.


Uses

All parts of the fruit, including the skin, except for the seeds, are suitable for human consumption. The core, from stem to bottom, containing the seeds, is usually not eaten and is discarded. Apples can be consumed in various ways:
juice Juice is a drink made from the extraction or pressing of the natural liquid contained in fruit and vegetables. It can also refer to liquids that are flavored with concentrate or other biological food sources, such as meat or seafood, such as ...
, raw in salads, baked in
pies A pie is a baked dish which is usually made of a pastry dough casing that contains a filling of various sweet or savoury ingredients. Sweet pies may be filled with fruit (as in an apple pie), nuts (pecan pie), brown sugar (sugar pie), sweeten ...
, cooked into
sauces In cooking, a sauce is a liquid, cream, or semi-solid food, served on or used in preparing other foods. Most sauces are not normally consumed by themselves; they add flavor, moisture, and visual appeal to a dish. ''Sauce'' is a French word t ...
and spreads like
apple butter Apple butter is a highly concentrated form of apple sauce produced by long, slow cooking of apples with cider or water to a point where the sugar in the apples caramelizes, turning the apple butter a deep brown. The concentration of sugar gives a ...
, and other baked dishes. Apples are sometimes used as an ingredient in savory foods, such as sausage and stuffing. Several techniques are used to preserve apples and apple products. Apples can be canned, dried or frozen. Canned or frozen apples are eventually baked into pies or other cooked dishes. Apple juice or cider is also bottled. Apple juice is often concentrated and frozen.


Popular uses

Apples are often eaten raw. Cultivars bred for raw consumption are termed dessert or
table apple Table apples or dessert apples are a group of apple cultivars grown for eating raw as opposed to cooking or cidermaking. Table apples are usually sweet and the most prized exhibit particular aroma variations that differentiate them from other ap ...
s. * In the UK, a
toffee apple Candy apples (or toffee apples in Commonwealth English) are whole apples covered in a sugar candy coating, with a stick inserted as a handle. These are a common treat at fall festivals in Western culture in the Northern Hemisphere, such as Hal ...
is a traditional confection made by coating an apple in hot
toffee Toffee is a confection made by caramelizing sugar or molasses (creating inverted sugar) along with butter, and occasionally flour. The mixture is heated until its temperature reaches the hard crack stage of . While being prepared, toffee is ...
and allowing it to cool. Similar treats in the U.S. are
candy apple Candy apples (or toffee apples in Commonwealth English) are whole apples covered in a sugar candy coating, with a stick inserted as a handle. These are a common treat at fall festivals in Western culture in the Northern Hemisphere, such as H ...
s (coated in a hard shell of crystallized sugar syrup) and
caramel apples Caramel apples or toffee apples are whole apples covered in a layer of caramel. They are created by dipping or rolling apples-on-a-stick in hot caramel, sometimes then rolling them in nuts or other small savories or confections, and allowing t ...
(coated with cooled
caramel Caramel ( or ) is an orange-brown confectionery product made by heating a range of sugars. It can be used as a flavoring in puddings and desserts, as a filling in bonbons, or as a topping for ice cream and custard. The process of caramelizati ...
). * Apples are eaten with honey at the Jewish New Year of
Rosh Hashanah Rosh HaShanah ( he, רֹאשׁ הַשָּׁנָה, , literally "head of the year") is the Jewish New Year. The biblical name for this holiday is Yom Teruah (, , lit. "day of shouting/blasting") It is the first of the Jewish High Holy Days (, , " ...
to symbolize a sweet new year. Apples are an important ingredient in many desserts, such as
apple pie An apple pie is a fruit pie in which the principal filling ingredient is apples. The earliest printed recipe is from England. Apple pie is often served with whipped cream, ice cream ("apple pie à la mode"), or cheddar cheese. It is general ...
, apple
crumble A crumble is a dish that can be made in a sweet or savoury version. Crumbles became popular in Britain during World War II, when the topping was an economical alternative to pies due to shortages of pastry ingredients as the result of rationi ...
,
apple crisp Apple crisp is a dessert made with a streusel topping. In the US and Canada, it is also called apple crumble, a word which refers to a different dessert in the UK, Australia, and New Zealand. Ingredients usually include cooked apples, butter, ...
and
apple cake Apple cakes are cakes in which apples feature as a main flavour and ingredient. Such cakes incorporate apples in a variety of forms, including diced, pureed, or stewed, and can include common additions like raisins, nuts, and 'sweet' spices suc ...
. When cooked, some apple cultivars easily form a puree known as
apple sauce Apple sauce or applesauce is a purée (not necessarily served as a true sauce) made of apples. It can be made with peeled or unpeeled apples and may be spiced or sweetened. Apple sauce is inexpensive and is widely consumed in North America and s ...
. Apples are also made into
apple butter Apple butter is a highly concentrated form of apple sauce produced by long, slow cooking of apples with cider or water to a point where the sugar in the apples caramelizes, turning the apple butter a deep brown. The concentration of sugar gives a ...
and apple jelly. They are often
baked Baking is a method of preparing food that uses dry heat, typically in an oven, but can also be done in hot ashes, or on hot stones. The most common baked item is bread but many other types of foods can be baked. Heat is gradually transferred ...
or
stewed A stew is a combination of solid food ingredients that have been cooked in liquid and served in the resultant gravy. A stew needs to have raw ingredients added to the gravy. Ingredients in a stew can include any combination of vegetables and m ...
and are also (cooked) in some meat dishes. Dried apples can be eaten or reconstituted (soaked in water, alcohol or some other liquid). Apples are milled or pressed to produce
apple juice Apple juice is a fruit juice made by the maceration and pressing of an apple. The resulting expelled juice may be further treated by enzymatic and centrifugal clarification to remove the starch and pectin, which holds fine particulate in suspe ...
, which may be drunk unfiltered (called
apple cider Apple cider (also called sweet cider, soft cider, or simply cider) is the name used in the United States and Canada for an unfiltered, unsweetened, non-alcoholic beverage made from apples. Though typically referred to simply as "cider" in the U ...
in North America), or filtered. Filtered juice is often concentrated and frozen, then reconstituted later and consumed. Apple juice can be
fermented Fermentation is a metabolic process that produces chemical changes in organic substrates through the action of enzymes. In biochemistry, it is narrowly defined as the extraction of energy from carbohydrates in the absence of oxygen. In food p ...
to make cider (called hard cider in North America),
ciderkin Ciderkin, sometimes referred to as water-cider, is a kind of weak alcoholic cider traditionally drunk by children, and made by steeping the refuse apple pomace in water. Ciderkin is currently listed alongside Cheate bread and Butter on the "Bill o ...
, and vinegar. Through
distillation Distillation, or classical distillation, is the process of separating the components or substances from a liquid mixture by using selective boiling and condensation, usually inside an apparatus known as a still. Dry distillation is the heating ...
, various alcoholic beverages can be produced, such as applejack,
Calvados Calvados (, , ) is a brandy from Normandy in France, made from apples or pears, or from apples with pears. History In France Apple orchards and brewers are mentioned as far back as the 8th century by Charlemagne. The first known record of Nor ...
, and
apfelwein ''Apfelwein'' ( German, 'apple wine'), or ''Viez'' ( Moselfranken, Saarland, Trier, ''vice'') or ''Most'' ( Austria, Switzerland, South Germany, ''must'') are German words for cider. It is made from a mix of sour tasting apples, such as "Bohna ...
.


Organic production

Organic apples are commonly produced in the United States. Due to infestations by key insects and diseases, organic production is difficult in Europe. The use of pesticides containing chemicals, such as sulfur, copper, microorganisms, viruses, clay powders, or plant extracts (
pyrethrum ''Pyrethrum'' was a genus of several Old World plants now classified as ''Chrysanthemum'' or '' Tanacetum'' which are cultivated as ornamentals for their showy flower heads. Pyrethrum continues to be used as a common name for plants formerly incl ...
,
neem ''Azadirachta indica'', commonly known as neem, nimtree or Indian lilac, is a tree in the mahogany family Meliaceae. It is one of two species in the genus ''Azadirachta'', and is native to the Indian subcontinent and most of the countries in Afri ...
) has been approved by the EU Organic Standing Committee to improve organic yield and quality. A light coating of
kaolin Kaolinite ( ) is a clay mineral, with the chemical composition Al2 Si2 O5( OH)4. It is an important industrial mineral. It is a layered silicate mineral, with one tetrahedral sheet of silica () linked through oxygen atoms to one octahedral s ...
, which forms a physical barrier to some pests, also may help prevent apple sun scalding.


Phytochemicals

Apple skins and seeds contain various
phytochemical Phytochemicals are chemical compounds produced by plants, generally to help them resist fungi, bacteria and plant virus infections, and also consumption by insects and other animals. The name comes . Some phytochemicals have been used as poisons ...
s, particularly
polyphenol Polyphenols () are a large family of naturally occurring organic compounds characterized by multiples of phenol units. They are abundant in plants and structurally diverse. Polyphenols include flavonoids, tannic acid, and ellagitannin, some of ...
s which are under preliminary research for their potential health effects.


Non-browning apples

The
enzyme Enzymes () are proteins that act as biological catalysts by accelerating chemical reactions. The molecules upon which enzymes may act are called substrates, and the enzyme converts the substrates into different molecules known as products. ...
polyphenol oxidase Polyphenol oxidase (PPO; also polyphenol oxidase i, chloroplastic), an enzyme involved in fruit browning, is a tetramer that contains four atoms of copper per molecule. PPO may accept monophenols and/or ''o''-diphenols as substrates. Th ...
causes browning in sliced or bruised apples, by catalyzing the
oxidation Redox (reduction–oxidation, , ) is a type of chemical reaction in which the oxidation states of substrate change. Oxidation is the loss of electrons or an increase in the oxidation state, while reduction is the gain of electrons or a d ...
of phenolic compounds to
o-quinone 1,2-Benzoquinone, also called ''ortho''-benzoquinone, is an organic compound with formula . It is one of the two isomers of quinone, the other being 1,4-benzoquinone. It is a red volatile solid that is soluble in water and ethyl ether. It is ...
s, a browning factor. Browning reduces apple taste, color, and food value.
Arctic apples Arctic apple is the trademark for a group of patented apples that contain a nonbrowning trait (when the apples are subjected to mechanical damage, such as slicing or bruising, the apple flesh remains as its original color) introduced through biote ...
, a non-browning group of apples introduced to the United States market in 2019, have been
genetically modified Genetic engineering, also called genetic modification or genetic manipulation, is the modification and manipulation of an organism's genes using technology. It is a set of technologies used to change the genetic makeup of cells, including ...
to silence the
expression Expression may refer to: Linguistics * Expression (linguistics), a word, phrase, or sentence * Fixed expression, a form of words with a specific meaning * Idiom, a type of fixed expression * Metaphorical expression, a particular word, phrase, ...
of polyphenol oxidase, thereby delaying a browning effect and improving apple eating quality. The US
Food and Drug Administration The United States Food and Drug Administration (FDA or US FDA) is a federal agency of the Department of Health and Human Services. The FDA is responsible for protecting and promoting public health through the control and supervision of food s ...
in 2015, and
Canadian Food Inspection Agency The Canadian Food Inspection Agency (CFIA; french: Agence canadienne d'inspection des aliments) is a regulatory agency that is dedicated to the safeguarding of food, plants, and animals (FPA) in Canada, thus enhancing the health and well-being of ...
in 2017, determined that Arctic apples are as safe and nutritious as conventional apples.


Other products

Apple seed oil Apple seed oil is a vegetable oil obtained by pressing apple seeds. It is used in manufacturing cosmetics.Krist, Sabine. (2020). ''Vegetable Fats and Oils''. Springer. pp. 61-66. Apple seed oil contains protein (about 34%). Apple seed oil may ...
is obtained by pressing apple seeds for manufacturing
cosmetics Cosmetics are constituted mixtures of chemical compounds derived from either natural sources, or synthetically created ones. Cosmetics have various purposes. Those designed for personal care and skin care can be used to cleanse or protect ...
.


Research

Preliminary research is investigating whether apple consumption may affect the risk of some types of cancer.


Allergy

One form of apple allergy, often found in northern Europe, is called birch-apple syndrome and is found in people who are also allergic to
birch A birch is a thin-leaved deciduous hardwood tree of the genus ''Betula'' (), in the family Betulaceae, which also includes alders, hazels, and hornbeams. It is closely related to the beech-oak family Fagaceae. The genus ''Betula'' contains 30 ...
pollen Pollen is a powdery substance produced by seed plants. It consists of pollen grains (highly reduced microgametophytes), which produce male gametes (sperm cells). Pollen grains have a hard coat made of sporopollenin that protects the gametophy ...
. Allergic reactions are triggered by a protein in apples that is similar to birch pollen, and people affected by this protein can also develop allergies to other fruits, nuts, and vegetables. Reactions, which entail oral allergy syndrome (OAS), generally involve itching and inflammation of the mouth and throat, but in rare cases can also include life-threatening
anaphylaxis Anaphylaxis is a serious, potentially fatal allergic reaction and medical emergency that is rapid in onset and requires immediate medical attention regardless of use of emergency medication on site. It typically causes more than one of the follow ...
. This reaction only occurs when raw fruit is consumed—the allergen is neutralized in the cooking process. The variety of apple, maturity and storage conditions can change the amount of allergen present in individual fruits. Long storage times can increase the amount of proteins that cause birch-apple syndrome. In other areas, such as the Mediterranean, some individuals have adverse reactions to apples because of their similarity to peaches. This form of apple allergy also includes OAS, but often has more severe symptoms, such as vomiting, abdominal pain and
urticaria Hives, also known as urticaria, is a kind of skin rash with red, raised, itchy bumps. Hives may burn or sting. The patches of rash may appear on different body parts, with variable duration from minutes to days, and does not leave any long-lasti ...
, and can be life-threatening. Individuals with this form of allergy can also develop reactions to other fruits and nuts. Cooking does not break down the protein causing this particular reaction, so affected individuals cannot eat raw or cooked apples. Freshly harvested, over-ripe fruits tend to have the highest levels of the protein that causes this reaction. Breeding efforts have yet to produce a
hypoallergenic Hypoallergenic, meaning "below average" or "slightly" allergenic, is a term meaning that something (usually cosmetics, pets, textiles, food, etc.) causes fewer allergic reactions. The term was first used in 1953 in an advertising campaign for cos ...
fruit suitable for either of the two forms of apple allergy.


Toxicity of seeds

Apple seeds contain small amounts of
amygdalin Amygdalin (from Ancient Greek: ' "almond") is a naturally occurring chemical compound found in many plants, most notably in the seeds (kernels) of apricots, bitter almonds, apples, peaches, cherries, and plums. Amygdalin is classified as a c ...
, a sugar and
cyanide Cyanide is a naturally occurring, rapidly acting, toxic chemical that can exist in many different forms. In chemistry, a cyanide () is a chemical compound that contains a functional group. This group, known as the cyano group, consists of a ...
compound known as a
cyanogenic glycoside In chemistry, a glycoside is a molecule in which a sugar is bound to another functional group via a glycosidic bond. Glycosides play numerous important roles in living organisms. Many plants store chemicals in the form of inactive glycosides. ...
. Ingesting small amounts of apple seeds causes no ill effects, but consumption of extremely large doses can cause
adverse reaction An adverse effect is an undesired harmful effect resulting from a medication or other intervention, such as surgery. An adverse effect may be termed a "side effect", when judged to be secondary to a main or therapeutic effect. The term complica ...
s. It may take several hours before the poison takes effect, as cyanogenic glycosides must be
hydrolyzed Hydrolysis (; ) is any chemical reaction in which a molecule of water breaks one or more chemical bonds. The term is used broadly for substitution, elimination, and solvation reactions in which water is the nucleophile. Biological hydrolysis i ...
before the cyanide ion is released. The United States
National Library of Medicine The United States National Library of Medicine (NLM), operated by the United States federal government, is the world's largest medical library. Located in Bethesda, Maryland, the NLM is an institute within the National Institutes of Health. Its ...
's
Hazardous Substances Data Bank The Hazardous Substances Data Bank (HSDB) is a toxicology database on the U.S. National Library of Medicine's (NLM) Toxicology Data Network (TOXNET). It focuses on the toxicology of potentially hazardous chemicals, and includes information on hum ...
records no cases of amygdalin poisoning from consuming apple seeds.


See also

* Apple chip *
Applecrab Applecrabs are various hybrids between crabapples and apples. They are bred for varying reasons, including disease resistance and use in cold climates because they are often hardier than apple trees and their fruit has the good eating qualities of ...
, apple–crabapple hybrids for eating *
Cooking apple A cooking apple or culinary apple is an apple that is used primarily for cooking, as opposed to a '' dessert apple'', which is eaten raw. Cooking apples are generally larger, and can be tarter than dessert varieties. Some varieties have a f ...
*
Johnny Appleseed John Chapman (September 26, 1774March 18, 1845), better known as Johnny Appleseed, was an American pioneer nurseryman who introduced apple trees to large parts of Pennsylvania, Ohio, Indiana, Illinois, and Ontario, as well as the norther ...
*
List of apple cultivars Over 7,500 cultivars of the culinary or eating apple (''Malus domestica'') are known. Some are extremely important economically as commercial products, though the vast majority are not suitable for mass production. In the following list, use for ...
*
List of apple dishes This is a list of apple dishes, that use apple as a primary ingredient. Apple beverages are also included on this list. Apple Dishes * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * ...
*
Rootstock A rootstock is part of a plant, often an underground part, from which new above-ground growth can be produced. It could also be described as a stem with a well developed root system, to which a bud from another plant is grafted. It can refer to a ...
* Welsh apples


References


Further reading

Books * * *


External links

* {{Authority control * * Fruits originating in Asia Plants described in 1768