''Annularia'' is a
form taxon
Form classification is the classification of organisms based on their morphology, which does not necessarily reflect their biological relationships. Form classification, generally restricted to palaeontology, reflects uncertainty; the goal of s ...
, applied to
fossil
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
foliage belonging to extinct plants of the genus ''
Calamites
''Calamites'' is a genus of extinct arborescent (tree-like) horsetails to which the modern horsetails (genus '' Equisetum'') are closely related. Unlike their herbaceous modern cousins, these plants were medium-sized trees, growing to heights o ...
'' in the order
Equisetales
Equisetales is an order of subclass Equisetidae with only one living family, Equisetaceae, containing the genus '' Equisetum'' (horsetails).
Classification
In the molecular phylogenetic classification of Smith et al. in 2006, Equisetales, in i ...
.
Description
''Annularia'' is a
form taxon
Form classification is the classification of organisms based on their morphology, which does not necessarily reflect their biological relationships. Form classification, generally restricted to palaeontology, reflects uncertainty; the goal of s ...
name given to leaves of ''
Calamites
''Calamites'' is a genus of extinct arborescent (tree-like) horsetails to which the modern horsetails (genus '' Equisetum'') are closely related. Unlike their herbaceous modern cousins, these plants were medium-sized trees, growing to heights o ...
''. In that species, the leaves formed radiating leaf whorls at each stem node, in a similar way to the branches of ''
Equisetum
''Equisetum'' (; horsetail, snake grass, puzzlegrass) is the only living genus in Equisetaceae, a family of ferns, which reproduce by spores rather than seeds.
''Equisetum'' is a "living fossil", the only living genus of the entire subclass ...
'', an extant genus of
horsetail
''Equisetum'' (; horsetail, snake grass, puzzlegrass) is the only living genus in Equisetaceae, a family of ferns, which reproduce by spores rather than seeds.
''Equisetum'' is a "living fossil", the only living genus of the entire subclass ...
s. ''Annularia'' leaves are arranged in whorls of between 8-13 leaves. The leaf shape is quite variable, being oval in ''Annularia sphenophylloides'' and linear to lanceolate in ''Annularia radiata'', but they are always flat and of varying lengths.
[MSN]
/ref>
''Calamites'' were arborescent and grew to a height of 32 feet (10 meters).
Fossil records
Fossils
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
of this genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
have been discovered in the Permian
The Permian ( ) is a geologic period and stratigraphic system which spans 47 million years from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.9 Mya. It is the last period of the Paleo ...
strata of Russia
Russia (, , ), or the Russian Federation, is a transcontinental country spanning Eastern Europe and Northern Asia. It is the largest country in the world, with its internationally recognised territory covering , and encompassing one-ei ...
and in the Carboniferous
The Carboniferous ( ) is a geologic period and system of the Paleozoic that spans 60 million years from the end of the Devonian Period million years ago ( Mya), to the beginning of the Permian Period, million years ago. The name ''Carboniferou ...
(around ) strata of the United States
The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country Continental United States, primarily located in North America. It consists of 50 U.S. state, states, a Washington, D.C., ...
, Canada
Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by to ...
, China
China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's List of countries and dependencies by population, most populous country, with a Population of China, population exceeding 1.4 billion, slig ...
and Europe
Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
.[Paleobiology Database]
/ref>
Calamitaceae - Annularia stellata.JPG, Specimen of ''Annularia stellata'' from Italy on display at the Museo Civico di Storia Naturale di Milano
The Museo Civico di Storia Naturale di Milano (Milan Natural History Museum) is a museum in Milan, Italy. It was founded in 1838 when
naturalist Giuseppe de Cristoforis donated his collections to the city. Its first director was Giorgio Jan. ...
Calamitaceae foliage.jpg, The foliage
A leaf ( : leaves) is any of the principal appendages of a vascular plant stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, ...
of ''Calamites''
Calamites reconstrucción.png, Calamites reconstruction
See also
*Coal forest
Coal forests were the vast swathes of wetlands that covered much of the Earth's tropical land areas during the late Carboniferous ( Pennsylvanian) and Permian times.Cleal, C. J. & Thomas, B. A. (2005). "Palaeozoic tropical rainforests and their e ...
References
External links
Fossil Museum
{{Taxonbar, from=Q2065549
Horsetails
Carboniferous plants
Prehistoric plant genera
Carboniferous first appearances
Carboniferous extinctions
Fossil taxa described in 1821
Fossils of Georgia (U.S. state)
Paleozoic life of New Brunswick
Paleozoic life of Nova Scotia
Prehistoric plants of North America