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archaeogenetics Archaeogenetics is the study of ancient DNA using various molecular genetic methods and DNA resources. This form of genetic analysis can be applied to human, animal, and plant specimens. Ancient DNA can be extracted from various fossilized specime ...
, the term Ancient North Eurasian (generally abbreviated as ANE) is the name given to an ancestral component that represents a lineage ancestral to the people of the
Mal'ta–Buret' culture The Mal'ta–Buret' culture is an archaeological culture of 24,000 to 15,000 BP / 22'050 to 13'050 BC in the Upper Paleolithic on the upper Angara River in the area west of Lake Baikal in the Irkutsk Oblast, Siberia, Russian Federation. Th ...
and populations closely related to them, such as from Afontova Gora and the
Yana Rhinoceros Horn Site The Yana Rhinoceros Horn Site (Yana RHS) is an Upper Palaeolithic archaeological site located near the lower Yana river in northeastern Siberia, Russia, north of the Arctic Circle in the far west of Beringia. It was discovered in 2001, after thawi ...
, and populations descended from them. ANE ancestry developed from a sister lineage of West-Eurasians with significant admixture from early
East Asians East Asian people (East Asians) are the people from East Asia, which consists of China, Taiwan, Japan, Mongolia, North Korea, and South Korea. The total population of all countries within this region is estimated to be 1.677 billion and 21% of the ...
. ANE ancestry has spread throughout Eurasia and the Americas in various migrations since the
Upper Paleolithic The Upper Paleolithic (or Upper Palaeolithic) is the third and last subdivision of the Paleolithic or Old Stone Age. Very broadly, it dates to between 50,000 and 12,000 years ago (the beginning of the Holocene), according to some theories coi ...
, and more than half of the world's population today derives between 5 to 40% of their genomes from the Ancient North Eurasians. Significant ANE ancestry can be found in the
indigenous peoples of the Americas The Indigenous peoples of the Americas are the inhabitants of the Americas before the arrival of the European settlers in the 15th century, and the ethnic groups who now identify themselves with those peoples. Many Indigenous peoples of the Am ...
, as well as in regions of Europe,
South Asia South Asia is the southern subregion of Asia, which is defined in both geographical and ethno-cultural terms. The region consists of the countries of Afghanistan, Bangladesh, Bhutan, India, Maldives, Nepal, Pakistan, and Sri Lanka.;;;;;;; ...
, Central Asia, and
Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part of ...
. It has been suggested that their mythology may have included a narrative, found in both Indo-European and some Native American fables, in which a dog guards the path to the afterlife.


Genetic studies

The ANE lineage is defined by association with the MA-1, or " Mal'ta boy", the remains of an individual who lived during the
Last Glacial Maximum The Last Glacial Maximum (LGM), also referred to as the Late Glacial Maximum, was the most recent time during the Last Glacial Period that ice sheets were at their greatest extent. Ice sheets covered much of Northern North America, Northern Eur ...
, 24,000 years ago in central
Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part of ...
, discovered in the 1920s. Ancient North Eurasians are described as a lineage "which is deeply related to Paleolithic/Mesolithic hunter-gatherers in Europe", meaning that they diverged from Paleolithic Europeans a long time ago. Ancient North Eurasians originated either in Europe or in the Middle East. Ancient North Eurasians also had varying degrees of admixture from an ancient East Asian-related population, such as the Tianyuan Man. Sikora et al. (2019) found that the oldest ANE-associated Yana RHS sample (31,600 BP) in Northeastern Siberia "can be modelled as early West Eurasian with an approximately 22% contribution from early East Asians" suggesting early contact in Northeastern Siberia. Populations genetically similar to MA-1 were an important genetic contributor to Native Americans,
Europeans Europeans are the focus of European ethnology, the field of anthropology related to the various ethnic groups that reside in the states of Europe. Groups may be defined by common genetic ancestry, common language, or both. Pan and Pfeil (2004) ...
, Ancient Central Asians,
South Asians South Asian ethnic groups are an ethnolinguistic grouping of the diverse populations of South Asia, including the nations of India, Pakistan, Bangladesh, Nepal, Bhutan, the Maldives, and Sri Lanka. While Afghanistan is variously considered to b ...
, and some East Asian groups, in order of significance. Lazaridis et al. (2016:10) note "a cline of ANE ancestry across the east-west extent of Eurasia." The ancient Bronze-age-steppe
Yamnaya The Yamnaya culture or the Yamna culture (russian: Ямная культура, ua, Ямна культура literal translation, lit. 'culture of pits'), also known as the Pit Grave culture or Ochre Grave culture, was a late Copper Age to ea ...
and Afanasevo cultures were found to have a noteworthy ANE component at ~25% (some studies estimated a higher percentage at nearly ~50%). According to Moreno-Mayar et al. 2018 between 14% and 38% of Native American ancestry may originate from gene flow from the Mal'ta–Buret' people (ANE). This difference is caused by the penetration of posterior Siberian migrations into the Americas, with the lowest percentages of ANE ancestry found in Eskimos and Alaskan Natives, as these groups are the result of migrations into the Americas roughly 5,000 years ago. Estimates for ANE ancestry among first wave Native Americans show higher percentages, such as 42% for those belonging to the
Andean The Andes, Andes Mountains or Andean Mountains (; ) are the longest continental mountain range in the world, forming a continuous highland along the western edge of South America. The range is long, wide (widest between 18°S – 20°S ...
region in South America. The other gene flow in Native Americans (the remainder of their ancestry) was of East Asian origin. Gene sequencing of another south-central Siberian people (Afontova Gora-2) dating to approximately 17,000 years ago, revealed similar autosomal genetic signatures to that of Mal'ta boy-1, suggesting that the region was continuously occupied by humans throughout the Last Glacial Maximum. Genomic studies also indicate that the ANE component was introduced to Western Europe by people related to the
Yamnaya culture The Yamnaya culture or the Yamna culture (russian: Ямная культура, ua, Ямна культура lit. 'culture of pits'), also known as the Pit Grave culture or Ochre Grave culture, was a late Copper Age to early Bronze Age arch ...
, long after the Paleolithic. It is reported in modern-day Europeans (10%–20%). Additional ANE ancestry is found in European populations through Paleolithic interactions with
Eastern European Hunter-Gatherers In archaeogenetics, the term Eastern Hunter-Gatherer (EHG), sometimes East European Hunter-Gatherer, or Eastern European Hunter-Gatherer is the name given to a distinct ancestral component that represents descent from Mesolithic hunter-gatherers of ...
, which resulted in populations such as Scandinavian Hunter-Gatherers.


Groups partially derived from the Ancient North Eurasians

Eastern Hunter-Gatherer In archaeogenetics, the term Eastern Hunter-Gatherer (EHG), sometimes East European Hunter-Gatherer, or Eastern European Hunter-Gatherer is the name given to a distinct ancestral component that represents descent from Mesolithic hunter-gatherers ...
(EHG) is a lineage which derived significant ancestry from ANE, ranging between 9% to up to 75%, with the remaining ancestry from a group more closely related to, but distinct from,
Western Hunter-Gatherers In archaeogenetics, the term Western Hunter-Gatherer (WHG), West European Hunter-Gatherer or Western European Hunter-Gatherer names a distinct ancestral component of modern Europeans, representing descent from a population of Mesolithic hunter-gat ...
(WHGs). It is represented by two individuals from
Karelia Karelia ( Karelian and fi, Karjala, ; rus, Каре́лия, links=y, r=Karélija, p=kɐˈrʲelʲɪjə, historically ''Korjela''; sv, Karelen), the land of the Karelian people, is an area in Northern Europe of historical significance fo ...
, one of Y-haplogroup R1a-M417, dated c. 8.4 kya, the other of Y-haplogroup J, dated c. 7.2 kya; and one individual from
Samara Samara ( rus, Сама́ра, p=sɐˈmarə), known from 1935 to 1991 as Kuybyshev (; ), is the largest city and administrative centre of Samara Oblast. The city is located at the confluence of the Volga and the Samara rivers, with a population of ...
, of Y-haplogroup R1b-P297, dated c. 7.6 kya. After the end of the Last Glacial Maximum, the
Western Hunter-Gatherer In archaeogenetics, the term Western Hunter-Gatherer (WHG), West European Hunter-Gatherer or Western European Hunter-Gatherer names a distinct ancestral component of modern Europeans, representing descent from a population of Mesolithic hunter-g ...
s (WHG) and EHG lineages merged in Eastern Europe, accounting for early presence of ANE-derived ancestry in Mesolithic Europe. Evidence suggests that as Ancient North Eurasians migrated West from Eastern Siberia, they absorbed Western Hunter-Gatherers and other West Eurasian populations as well.
Scandinavian Hunter-Gatherer In archaeogenetics, the term Scandinavian Hunter-Gatherer (SHG) is the name given to a distinct ancestral component that represents descent from Mesolithic hunter-gatherers of Scandinavia. Genetic studies suggest that the SHGs were a mix of W ...
(SHG) is represented by several individuals buried at
Motala Motala () is a locality and the seat of Motala Municipality, Östergötland County, Sweden with 29,823 inhabitants (41,956 in the entire municipality) in 2010. It is the third largest city of Östergötland, following Linköping and Norrköping. ...
, Sweden ca. 6000 BC. They were descended from
Western Hunter-Gatherer In archaeogenetics, the term Western Hunter-Gatherer (WHG), West European Hunter-Gatherer or Western European Hunter-Gatherer names a distinct ancestral component of modern Europeans, representing descent from a population of Mesolithic hunter-g ...
s who initially settled
Scandinavia Scandinavia; Sámi languages: /. ( ) is a subregion in Northern Europe, with strong historical, cultural, and linguistic ties between its constituent peoples. In English usage, ''Scandinavia'' most commonly refers to Denmark, Norway, and Swed ...
from the south, and later populations of EHG who entered Scandinavia from the north through the coast of
Norway Norway, officially the Kingdom of Norway, is a Nordic country in Northern Europe, the mainland territory of which comprises the western and northernmost portion of the Scandinavian Peninsula. The remote Arctic island of Jan Mayen and th ...
. Ancient Beringian/Ancestral Native American are specific archaeogenetic lineages, based on the genome of an infant found at the Upward Sun River site (dubbed USR1), dated to 11,500 years ago. The AB lineage diverged from the Ancestral Native American (ANA) lineage about 20,000 years ago. West Siberian Hunter-Gatherer (WSHG) are a specific archaeogenetic lineage, first reported in a genetic study published in
Science Science is a systematic endeavor that builds and organizes knowledge in the form of testable explanations and predictions about the universe. Science may be as old as the human species, and some of the earliest archeological evidence for ...
in September 2019. WSGs were found to be of about 30% EHG ancestry, 50% ANE ancestry, and 20% to 38% East Asian ancestry.
Western Steppe Herders In archaeogenetics, the term Western Steppe Herders (WSH), or Western Steppe Pastoralists, is the name given to a distinct ancestral component first identified in individuals from the Eneolithic steppe around the turn of the 5th millennium BCE, ...
(WSH) is the name given to a distinct ancestral component that represents descent closely related to the
Yamnaya culture The Yamnaya culture or the Yamna culture (russian: Ямная культура, ua, Ямна культура lit. 'culture of pits'), also known as the Pit Grave culture or Ochre Grave culture, was a late Copper Age to early Bronze Age arch ...
of the
Pontic–Caspian steppe The Pontic–Caspian steppe, formed by the Caspian steppe and the Pontic steppe, is the steppeland stretching from the northern shores of the Black Sea (the Pontus Euxinus of antiquity) to the northern area around the Caspian Sea. It extend ...
. This ancestry is often referred to as Yamnaya ancestry or Steppe ancestry. Late Upper Paeolithic Lake Baikal - Ust'Kyakhta-3 (UKY) 14,050-13,770 BP were mixture of 30% ANE ancestry and 70% East Asian ancestry. Lake Baikal Holocene - Baikal Eneolithic (Baikal_EN) and Baikal Early Bronze Age (Baikal_EBA) derived 6.4% to 20.1% ancestry from ANE, while rest of their ancestry was derived from East Asians. Fofonovo_EN near by Lake Baikal were mixture of 12-17% ANE ancestry and 83-87% East Asian ancestry.
Jōmon people is the generic name of several peoples who lived in the Japanese archipelago during the Jōmon period (). The Jōmon people may have consisted of multiple groups, which arrived and merged at different times in the Japanese archipelago, using mult ...
, the pre-
Neolithic The Neolithic period, or New Stone Age, is an Old World archaeological period and the final division of the Stone Age. It saw the Neolithic Revolution, a wide-ranging set of developments that appear to have arisen independently in several part ...
population of Japan, mainly derived their ancestry from East Asian lineages, but also received geneflow from the ANE-related "Ancient North Siberians" (represented by samples from the
Yana Rhinoceros Horn Site The Yana Rhinoceros Horn Site (Yana RHS) is an Upper Palaeolithic archaeological site located near the lower Yana river in northeastern Siberia, Russia, north of the Arctic Circle in the far west of Beringia. It was discovered in 2001, after thawi ...
) prior to the migration from the Asian mainland to the Japanese archipelago. Jōmon ancestry is still found among the inhabitants of present-day Japan: most markedly among the
Ainu people The Ainu are the indigenous people of the lands surrounding the Sea of Okhotsk, including Hokkaido Island, Northeast Honshu Island, Sakhalin Island, the Kuril Islands, the Kamchatka Peninsula and Khabarovsk Krai, before the arrival of the Yam ...
, who are considered the direct descendants of the Jōmon people, and to a small, but significant degree among the majority of the Japanese population.


Evolution of blond hair

Single nucleotide polymorphisms In genetics, a single-nucleotide polymorphism (SNP ; plural SNPs ) is a germline substitution of a single nucleotide at a specific position in the genome. Although certain definitions require the substitution to be present in a sufficiently larg ...
of the KITLG gene are associated with blonde hair color in various human populations. One of these polymorphisms is associated with blond hair. The earliest known individual with this allele is a female south-central Siberian ANE individual from Afontova Gora 3 site, which is dated to 16130-15749 BCE., "Japanese research in 2006 found that the genetic mutation that prompted the evolution of blond hair dates to the ice age that happened around 11,000 years ago. Since then, the 17,000-year-old remains of a blond- haired North Eurasian hunter-gatherer have been found in eastern Siberia, suggesting an earlier origin." Mathieson, et al (2018) thus argued that this gene originated in the Ancient North Eurasian population. "Supplementary Information page 52: "The derived allele of the KITLG SNP rs12821256 that is associated with – and likely causal  for blond hair in Europeans7,8 is present in one hunter-gatherer from each of Samara, Motala  and Ukraine (I0124, I0014 and I1763), as well as several later individuals with Steppe ancestry. Since the allele is found in populations with EHG but not WHG ancestry, it suggests  that its origin is in the Ancient North Eurasian (ANE) population. Consistent with this, we observe that the earliest known individual with the derived allele (supported by two reads) is the ANE individual Afontova Gora 3,6 which is directly dated to 16130-15749 cal BCE (14710±60 BP, MAMS-27186: a previously unpublished date that we newly report here). We  cannot determine the status of rs12821256 in Afontova Gora 2 and MA-1 due to lack of  sequence coverage at this SNP.9" Geneticist David Reich said that the KITLG gene for blond hair probably entered continental Europe in a population migration wave from the Eurasian steppe, by a population carrying substantial Ancient North Eurasian ancestry. "The earliest known example of the classic European blond hair mutation is in an Ancient North Eurasian from the Lake Baikal region of eastern Siberia from seventeen thousand years ago. The hundreds of millions of copies of this mutation in central and western Europe today likely derive from a massive migration of people bearing Ancient North Eurasian ancestry, an event that is related in the next chapter." Hanel and Carlberg (2020) likewise report that populations bearing Ancient North Eurasian ancestry were responsible for contributing this gene to Europeans.


Comparative mythology

Since the term 'Ancient North Eurasian' refers to a genetic bridge of connected mating networks, scholars of
comparative mythology Comparative mythology is the comparison of myths from different cultures in an attempt to identify shared themes and characteristics.Littleton, p. 32 Comparative mythology has served a variety of academic purposes. For example, scholars have used ...
have argued that they probably shared myths and beliefs that could be reconstructed via the comparison of stories attested within cultures that were not in contact for millennia and stretched from the
Pontic–Caspian steppe The Pontic–Caspian steppe, formed by the Caspian steppe and the Pontic steppe, is the steppeland stretching from the northern shores of the Black Sea (the Pontus Euxinus of antiquity) to the northern area around the Caspian Sea. It extend ...
to the
American continent The Americas, which are sometimes collectively called America, are a landmass comprising the totality of North and South America. The Americas make up most of the land in Earth's Western Hemisphere and comprise the New World. Along with t ...
. For instance, the
mytheme In structuralism-influenced studies of mythology, a mytheme is a fundamental generic unit of narrative structure (typically involving a relationship between a character, an event, and a theme) from which myths are thought to be constructed—a mi ...
of the dog guarding the
Otherworld The concept of an otherworld in historical Indo-European religion is reconstructed in comparative mythology. Its name is a calque of ''orbis alius'' (Latin for "other Earth/world"), a term used by Lucan in his description of the Celtic Otherworld ...
possibly stems from an older Ancient North Eurasian belief, as suggested by similar motifs found in
Indo-European The Indo-European languages are a language family native to the overwhelming majority of Europe, the Iranian plateau, and the northern Indian subcontinent. Some European languages of this family, English, French, Portuguese, Russian, Dutc ...
, Native American and Siberian mythology. In
Siouan Siouan or Siouan–Catawban is a language family of North America that is located primarily in the Great Plains, Ohio and Mississippi valleys and southeastern North America with a few other languages in the east. Name Authors who call the entire ...
, Algonquian,
Iroquoian The Iroquoian languages are a language family of indigenous peoples of North America. They are known for their general lack of labial consonants. The Iroquoian languages are polysynthetic and head-marking. As of 2020, all surviving Iroquoian la ...
, and in Central and South American beliefs, a fierce guard dog was located in the
Milky Way The Milky Way is the galaxy that includes our Solar System, with the name describing the galaxy's appearance from Earth: a hazy band of light seen in the night sky formed from stars that cannot be individually distinguished by the naked eye. ...
, perceived as the path of souls in the afterlife, and getting past it was a test. The Siberian Chukchi and Tungus believed in a guardian-of-the-afterlife dog and a spirit dog that would absorb the dead man's soul and act as a guide in the afterlife. In Indo-European myths, the figure of the dog is embodied by
Cerberus In Greek mythology, Cerberus (; grc-gre, Κέρβερος ''Kérberos'' ), often referred to as the hound of Hades, is a multi-headed dog that guards the gates of the Underworld to prevent the dead from leaving. He was the offspring of the mo ...
, Sarvarā, and
Garmr In Norse mythology, Garmr or Garm (Old Norse: ; "rag") is a wolf or dog associated with both Hel and Ragnarök, and described as a blood-stained guardian of Hel's gate. Name The Old Norse name ''Garmr'' has been interpreted as meaning "rag." ...
. In Zoroastrianism, two four-eyed dogs guard the bridge to the afterlife called
Chinvat Bridge The Chinvat Bridge ( Avestan: 𐬗𐬌𐬥𐬬𐬀𐬙𐬋 𐬞𐬈𐬭𐬈𐬙𐬏𐬨 ''Cinvatô Peretûm'', "bridge of judgement" or "beam-shaped bridge") or the Bridge of the Requiter in Zoroastrianism is the sifting bridge, which separates t ...
.
Anthony Anthony or Antony is a masculine given name, derived from the '' Antonii'', a ''gens'' ( Roman family name) to which Mark Antony (''Marcus Antonius'') belonged. According to Plutarch, the Antonii gens were Heracleidae, being descendants of Anton, ...
and Brown note that it might be one of the oldest mythemes recoverable through
comparative mythology Comparative mythology is the comparison of myths from different cultures in an attempt to identify shared themes and characteristics.Littleton, p. 32 Comparative mythology has served a variety of academic purposes. For example, scholars have used ...
. A second canid-related series of beliefs, myths and rituals connected dogs with healing rather than death. For instance, Ancient Near Eastern and Turkic- Kipchaq myths are prone to associate dogs with healing and generally categorised dogs as impure. A similar myth-pattern is assumed for the Eneolithic site of Botai in Kazakhstan, dated to 3500 BC, which might represent the dog as absorber of illness and guardian of the household against disease and evil. In Mesopotamia, the goddess Nintinugga, associated with healing, was accompanied or symbolized by dogs. Similar absorbent-puppy healing and sacrifice rituals were practiced in Greece and Italy, among the
Hittites The Hittites () were an Anatolian people who played an important role in establishing first a kingdom in Kussara (before 1750 BC), then the Kanesh or Nesha kingdom (c. 1750–1650 BC), and next an empire centered on Hattusa in north-centra ...
, again possibly influenced by Near Eastern traditions.


Steam bath ritual

Sven Kurbel has suggested that, as a result of the high levels of dust in the atmosphere during the Last Glacial Maximum, the Ancient North Eurasians may have developed steam baths to cleanse the skin and respiratory organs, as well as perhaps serving some sort of spiritual function. Kurbel speculates that this may explain why sauna-type steam baths are typical of Northwest European and Native American populations, who perhaps inherited this practice from their Ancient North Eurasian ancestors.


See also

*
Western Hunter-Gatherer In archaeogenetics, the term Western Hunter-Gatherer (WHG), West European Hunter-Gatherer or Western European Hunter-Gatherer names a distinct ancestral component of modern Europeans, representing descent from a population of Mesolithic hunter-g ...
*
Prehistory of Siberia The Prehistory of Siberia is marked by several archaeologically distinct cultures. In the Chalcolithic, the cultures of western and southern Siberia were pastoralists, while the eastern taiga and the tundra were dominated by hunter-gatherers unt ...


References


Notes


Bibliography

* * * * * * * * * * * * * * * * * * * * * * * * * * * {{Human genetics
ANE Ane or ane may refer to: * Āne, a village in Latvia * Ane, Netherlands, a village in Overijssel, Netherlands, also ** Battle of Ane (1227), a battle fought near the village * -ane, a suffix in organic chemistry, or specifically ** Alkanes, whi ...
Genetic history of Europe Peopling of the world Peopling of the Americas Indo-European genetics Last Glacial Maximum Mesolithic