''Archaea'' is an extinct genus of
spider
Spiders (order (biology), order Araneae) are air-breathing arthropods that have eight limbs, chelicerae with fangs generally able to inject venom, and spinnerets that extrude spider silk, silk. They are the largest order of arachnids and ran ...
s in the family
Archaeidae
Archaeidae, also known as assassin spiders and pelican spiders, is a spider family with about ninety described species in five genera. It contains small spiders, ranging from long, that prey exclusively on other spiders. They are unusual in that ...
. , four species are placed (or possibly placed) in the genus. All have been found preserved in
amber
Amber is fossilized tree resin. Examples of it have been appreciated for its color and natural beauty since the Neolithic times, and worked as a gemstone since antiquity."Amber" (2004). In Maxine N. Lurie and Marc Mappen (eds.) ''Encyclopedia ...
, either from the
Baltic
Baltic may refer to:
Peoples and languages
*Baltic languages, a subfamily of Indo-European languages, including Lithuanian, Latvian and extinct Old Prussian
*Balts (or Baltic peoples), ethnic groups speaking the Baltic languages and/or originatin ...
or
Bitterfeld
Bitterfeld () is a town in the district of Anhalt-Bitterfeld, Saxony-Anhalt, Germany. Since 1 July 2007 it has been part of the town of Bitterfeld-Wolfen. It is situated approximately 25 km south of Dessau, and 30 km northeast of Hall ...
, Germany.
[ First described in 1854, ''Archaea'' species have a distinctive "neck" separating the head from the thorax, and very long ]chelicerae
The chelicerae () are the arthropod mouthparts, mouthparts of the subphylum Chelicerata, an arthropod group that includes arachnids, horseshoe crabs, and sea spiders. Commonly referred to as "jaws", chelicerae may be shaped as either articulated ...
("jaws").[
]
Description
In the genus ''Archaea'', the head is separated from the thorax by a deep fold, forming a "neck", so that the globular head appears to sit on the thorax. There are eight eyes, four on each side of the head, arranged in the shape of a rhombus
In plane Euclidean geometry, a rhombus (: rhombi or rhombuses) is a quadrilateral whose four sides all have the same length. Another name is equilateral quadrilateral, since equilateral means that all of its sides are equal in length. The rhom ...
, with the front eyes being the largest. The curved chelicerae
The chelicerae () are the arthropod mouthparts, mouthparts of the subphylum Chelicerata, an arthropod group that includes arachnids, horseshoe crabs, and sea spiders. Commonly referred to as "jaws", chelicerae may be shaped as either articulated ...
are very long, longer than the head, with long, strong fangs. The pedipalp
Pedipalps (commonly shortened to palps or palpi) are the secondary pair of forward appendages among Chelicerata, chelicerates – a group of arthropods including spiders, scorpions, horseshoe crabs, and sea spiders. The pedipalps are lateral to ...
s of the female are very small and thin, shorter than the chelicerae, the third and last segments being short, the latter needle-shaped. Those of the male are sturdier, the third segment being almost as long as the second, with the end segment bearing the palpal bulb
The two palpal bulbs – also known as palpal organs and genital bulbs – are the copulatory organs of a male spider. They are borne on the last segment of the pedipalps (the front "limbs" of a spider), giving the spider an appearance often descr ...
s. The thorax is relatively flat, narrower at the back than the front, and without thoracic grooves. The legs are long and thin. The first pair are the longest, the second somewhat longer than the fourth, the third being the shortest. The patellae and tarsi are short, without spines or bristles.[
Living members of the family Archaeidae are predators on other spiders (araneophageous); it is assumed that extinct members of the family had the same habits.][
]
Taxonomy
The genus ''Archaea'' was erected in 1854 by C. L. Koch and G. C. Berendt,[ initially with three extinct species, all found in ]amber
Amber is fossilized tree resin. Examples of it have been appreciated for its color and natural beauty since the Neolithic times, and worked as a gemstone since antiquity."Amber" (2004). In Maxine N. Lurie and Marc Mappen (eds.) ''Encyclopedia ...
from the Baltic Sea
The Baltic Sea is an arm of the Atlantic Ocean that is enclosed by the countries of Denmark, Estonia, Finland, Germany, Latvia, Lithuania, Poland, Russia, Sweden, and the North European Plain, North and Central European Plain regions. It is the ...
or Bitterfeld
Bitterfeld () is a town in the district of Anhalt-Bitterfeld, Saxony-Anhalt, Germany. Since 1 July 2007 it has been part of the town of Bitterfeld-Wolfen. It is situated approximately 25 km south of Dessau, and 30 km northeast of Hall ...
in Saxony-Anhalt
Saxony-Anhalt ( ; ) is a States of Germany, state of Germany, bordering the states of Brandenburg, Saxony, Thuringia and Lower Saxony. It covers an area of
and has a population of 2.17 million inhabitants, making it the List of German states ...
, Germany. It was then the only genus in the family Archaeidae.[ The first living species in the family was described in 1881 by O. Pickard-Cambridge. He placed it in the genus '' Eriauchenius'' as '' E. workmani''. In 1895, ]Eugène Simon
Eugène Louis Simon (; 30 April 1848 – 17 November 1924) was a French naturalist who worked particularly on insects and spiders, but also on birds and plants. He is by far the most prolific spider Taxonomy (biology), taxonomist in history, ...
transferred this species to ''Archaea'' as ''Archaea workmani''.[ Other living species were placed in the genus later, but all are now considered to belong to different genera.][
]
Species
*†?''Archaea bitterfeldensis'' Wunderlich, 2004 – Bitterfeld amber
*†''Archaea compacta'' Wunderlich, 2004 – Baltic amber
*†''Archaea paradoxa'' C.L. Koch & Berendt, 1854 (syn. ''Archaea laevigata'' C. L. Koch & Berendt, 1854, ''Archaea incompta'' Menge, 1854) (type species
In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
) – Baltic amber
*†''Archaea pougneti'' Simon, 1884 – Baltic amber
Fossil species that have been placed in ''Archaea'' but are now placed elsewhere include:[
*†''Eroarchaea hyperoptica'' (Menge, 1854) = ''Archaea hyperoptica''
*†''Baltarchaea conica'' (Koch & Berendt 1854) = ''Archaea conica''
]
References
{{Taxonbar, from=Q2859968
Archaeidae
Araneomorphae genera
Extinct arachnids
Fossil taxa described in 1854